Trichopilia aenigma Garay, sp. nov. TYPE:

GARAY ET AL.-INQUlLINA ORCHIDACEA II
Trichopilia aenigma Garay, sp. nov. TYPE:
ANDES, without precise locality. Type specimen
is cultivated by Dr. Wojciech Klikunas s.n. (Holotype: AMES). Fig. 4.
Planta epiphytic a, ca. 38 cm alta; radicibus
crassiusculis, glabris; pseudobulbis approximatis,
lateraliter valde compressis, in ambitu oblique
ovatis vel ovato-oblongis, monophyllis, usque ad
12 cm longis, 3.5 cm latis; foliis subcoriaceis, oblongo-ellipticis, acutis, sessilibus, usque ad 26 cm
longis, 6 cm latis, vulgo minoribus; inflorescentia
arcuata vel arcuatim pendula, pauciflora; bracteis
ovatis, acutis, ovariis adpressis; floribus plus minusve camosis, apertis; sepalis petalisque pallide
233
GARAY ET AL.-INQUILlNA ORCHIDACEA II
CO\u,..,.. ....
Fig. 4.
Trichopilia aenigma. Reproduced from Dunsterville & Garay, Venezuelan Orchids Illustrated 3: 317, 1965.
nandrio dorsaliter lacerato, usque ad 20 mm 10nga; ovario teretiusculo, usque ad 25 mm longo.
PARATYPES: VENEZUELA. Depto. Bolivar,
above Guayaraca, 3,500 ft. alt. G. C. K. Dunsterville 800 (AMES). COLOMBIA. Depto. huila,
Suza, between 500 and 1,000 malt. G. Escobar
No. 948 (AMES). Without precise locality. C.
Loddiges 1680 (K).
This is indeed an enigmatic species. It has been
the source of confusion ever since Loddiges introduced it in 1843 from Colombia. Of that intro-
viridibus, labello albido in funde luteo; sepalis inter se simillimis, liberis, linearibus, margine paululo undulatis, acuminatis, dorsal iter obscure carinatis, usque ad 40 mm longis, 7 mm latis; petalis
simillimis, margine interdum crispato-undulatis,
usque ad 37 mm longis, 7 mm latis; labello e
cuneata basi elliptico, obscure lobulato, margine
anteriori undulato-erosulo, apice bilobulo, disco a
basi ad medium usque distincte carinato, carina
obscura in fundo utrinque donato, usque ad 30
mm longo, 20 mm lato; columna cylindrica, cli234
GARAY ET AL.-INQUIUNA ORCHIDACEA II
Guide pt. 3: 1023, 1979 as T. albida Wendl. Orquideologia 8: 97, 1973 as T. wageneri (Rchb.f.)
Rchb .f.
The following key is offered to facilitate the
identification of species of the Trichopilia fragrans complex.
duction there is an inflorescence with two flowers
in the Lindley Herbarium at Kew, and also an
excellent drawing of a flower, column, and pollinia by Lindley's hand. On the herbarium sheet in
Lindley's hand is added in pencil "P[ilumna] candida m[ihi]. "Pilumna candida or rather "Trichopilia candida" appeared the first time under Pilumna fragrans Lindl. in Lindley, Orchidaceae
Lindenianae, in 1846, in association with a collection by Linden No. 649 (K) from Venezuela,
Merida. Since Dunsterville's No. 800 was identical with the Loddiges material we reported and
illustrated it in 1965 as T. fragrans (Lindl.)
Rchb.f. In 1973, when I received photographs and
dried flowers from Escobar No. 948, it became
apparent that both Loddiges No. 1680 and Dunsterville No. 800 are not only identical with Escobar No. 948, but that they are also different
from T. fragrans which up to that time was considered to include T. wageneri Rchb.f. In the intervening time the true T. wagneri has been rediscovered by Dunsterville (No. 990). This new
rediscovery left the ColombianNenezuelan materials without a name. While preparing the Orchids of Venezuela, Field Guide in 1979, Dunsterville and I assigned these collections to T. albida Wendl. with much hesitation. In retrospect
this choice again becomes untenable. With the arrival of new fresh material from the Trichopilia
collection of Dr. Klikunas I wish to put to rest
this ongoing and mounting confusion by naming
it appropriately an enigmatic Trichopilia.
ETYMOLOGY: From the Greek aenigma
meaning a riddle, something obscure, in reference
to its history.
ILLUSTRATIONS: Dunsterville & Garay,
Venezuelan Orchids Illustrated 3: 316-317, 1965
as T. fragrans (Lindl.) Rchb.f. Dunsterville &
Garay, Orchids of Venezuela, An Illustrated Field
1. Dorsal and lateral sepals 5 to 6 cm long . . .. ... . 2
la. Dorsal and lateral sepals 2.5 to 4 cm long . . . . . . . 3
2. Sepals, petals and lip without undulate margins; lip
obscurely pandurate, slightly emarginate in front .. ..
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . T. wageneri
2a. Sepals commonly, but not always with straight edges,
petals with undulate margins; lip distinctly lobed in
middle, deeply emarginate to bilobed in front (hence
quasi 4-lobed) . . . . . . . . . . . . . . . . . . .. T. fragrans
3. Sepals and petals white; lip from a cuneate-unguiculate base subquadrate, more precisely obtrapezoid; clinandrium triangular, entire ...... . .. . . . T. nobilis
3a. Sepals and petals greenish; lip with a cuneate, but not
unguiculate base; clinandrium lacerate-fimbriate ... 4
4. Sepals 4 cm long; the apical half of lip with eroseundulate margin, disc shallowly 3-carinate in throat
... . .. ..... .. .. . ..... . ... . .... T. aenigma
4a. Sepals 2.5 to 3 cm long; the apical half of lip lobulate
with entire margin, disc with an abruptly elevated, lamella-like keel in throat ..... . . .... .. .. T. grata
LITERATURE CITED
Ames, O. 1907. Orchidaceae Ha\conenses. Phil. 1. Sci. 2(4):
311-337.
- -. 1924. Orchidaceae in E. D. Merrill, An enumeration of
Philippine flowering plants. (Bur. Sci. Publ. Manila 18)
1(3): 333-334.
Argus [only name provided in original publication] . 1895.
Dies Orchidianae. Orch. Rev. 3: 135-137.
Dunsterville, G. C. K., and L. A. Garay. 1965. Venezuelan
Orchids Illustrated 3. Andre Deutsch, London.
- -, and L. A. Garay. 1979. Orchids of Venezuela, An Illustrated Field Guide. Botanical Museum, Harvard University, Cambridge, Massachusetts.
Lindley, J. 1846. Orchidaceae Lindenianae. Bradbury &
Evans, London.
Loddiges, c., and Sons. 1844. Orchideae. Wilson & Ogilvy,
London.
Rumphius, G. E. 1750. Herbarium Amboinense pI. 6: 115116, T. 52, fig . 2. Amsterdam, Den Haag.
Willdenow, C. L. 1805. Species Plantarum 4: 122. G. C.
Nank Imp., Berlin.
Willemet, P. R. 1796. Herbarium Mauritianum in P. Usteri,
Neue Ann. Bot. 6(18): 52-53.
235