Original

in (McVaugh & Koelz
Brandt 2242); coastal
and primordia of new
tquila (Aquila, Hinton
Hinton 12479)- "11 mi
2-6 (-10) m high on
:hinly pilose, the whole
eath, the inflorescencend stipels, like those of
nflorescences like those
the peduncles 1-10 cm
;xceptionally -100!) cm
y attenuate to the tip;
m of the axis; flowers 2
ose of vars. crassa and
imbricated even in the
ite or sub acute to shortced below the point of
sdicel, and corolla-color,
i often partly united into
tened, 2.5-2.7 cm wide,
obovoid, 1.8-2 cm long,
rare, crassa and strobiloJ-shaped, again abruptly
:h inflexed tip, the whole
d strobilophora; fruiting
species, as was done by
& Steyermark (1946). On
:ms possible to distinguish
inflorescence, smaller and
als, and more uniformly
er hand, in aspect and in
Jrom Vigna strobilophora.
'ubescence, are seemingly
irescence, used in the key
, making identification of
arietal level.
se, the holotype of which
ri. Rose (1903) said of R.
>ut more glabrous throughh long apiculations to the
a moderately slender lax
tie bracts seem to resemble
e-axis suggests that of var.
LEGUMINOSAE - PAPILIONOIDEAE
753
Vigna strobilophora var. crassa McVaugh, var. nov., a var. strobilophora
bracteis primariis obtusis vel subacutis, bracteolis late ovatis vel suborbicularibus
1 mm longis, inflorescentiae axe in statu fructifero quam pedunculo saepe crassiore, usque ad 4 mm crasso, differt; a var. buseri (ut etiam similiter var. strobilophora) inflorescentiae immaturae bracteis primariis late imbricatis capitulum
strobiliforme formantibus, inflorescentiae axe breviore, nodis congestis, differt.
Wooded slopes and ravines, stream-valleys, thickets and roadsides, openings
in oak forest or subdeciduous forest with Hura, Brosimum, Ficus, sometimes
persistent in secondary forest with Cordia, 200-1200 m in the mountains and
foothills of the Pacific slope, flowering Aug-Sep, fruiting Oct-Dec.
Nay. ("9 mi" N of Compostela, fl 29 Aug, McVaugh 16529, MICH!, the
holotype; fr 12 Nov, McVaugh & Koelz 514, MICH), Jal. Endemic as far as
known.
NE of Ixcatan, SW of Santa Cruz del Guaibel (Bauml et al. 1159, HNT,
MICH); W of Jalcocotan, road from Tetitata (Feddema 1026); between Las Varas
and Compostela (Gentry 22327, Dieterle 3979, both MICH); "10 mi" SE of
Ahuacatlan, road to Barranca del Oro (McVaugh & Koelz 816); Santa Cruz de
Vallarta (Mexia 1268, MICH); seaward-facing slopes SW of Autlan (Wilbur 2276,
McVaugh 19944, both MICH); "12 mi" SW of La Huerta (Anderson & Laskowski 3771, MICH); S of Cuautitlan, road to Casimiro Castillo (Iltis et al. 382,
MICH).
Woody vine up to 6-10 m long, often high-climbing on trees, the tips of the
stems herbaceous, the base up to 2.5-3 cm thick; plants copiously or thinly pilose
almost throughout (except for the flowers) with soft pale short hairs, the inflorescence-axis usually glabrous or nearly so, the ovary glabrous or essentially so, the
fruit glabrous; leaves long-petiolate, the rachis mostly 1.5-4 cm long, the leaflets
rather broadly ovate, acuminate, the lateral ones oblique; leaflets of major leaves
10-14 cm long; stipules thick, ovate to deltoid, blunt or acute, subcoriaceous, up to
6 mm wide at the persistent corky base, often reflexed; stipels linear-lanceolate,
several-striate, coriaceous, 2-3.5 (-5) mm long; inflorescences axillary, the naked
peduncles (2-) 6-10 (-14) cm long, 1.5-2.7 mm thick, terminating in the flowering
axis 3-6 (-13) cm long, commonly thicker than the peduncle, up to 4 mm thick in
fruit; flowering nodes conspicuous (enlarged), crowded, 5-7 (-10) to 1 cm of the
axis; flowers 2 at a node; pedicels 9-12 (-14) mm long; floral (primary) bracts in
the young inflorescence broadly imbricated over the buds in a conelike aggregation; mature bracts white, fleshy, multistriate, 18-20 mm long, 12-16 mm wide,
broadly ovate, obtuse to subacute with a very short blunt point, deciduous before
full anthesis; bracteoles broadly ovate to suborbicular, produced below the point of
insertion, 1 mm long, deciduous at anthesis; calyx ca 7 mm long, campanulate,
essentially glabrous except the ciliate margins, the teeth subequal or the adaxial
pair united into an emarginate lobe, rounded to ovate and bluntly acuminate, erect
or the abaxial one recurved, acute; pedicels and calyx whitish or reddish to violet at
anthesis; corolla white to lavender or violet, the standard white or pale or with
yellow spot at base, the keel usually pale, the wings deep lavender, violet, or
purple; blade of the standard orbicular-reniform above a broad claw 2 mm long, ca
2 cm long, 2.5 cm wide with 2 lamellae at the conduplicate base, these 5 mm long, 1
mm high at base, produced proximally into a pair of short rounded auricles, nar-
754
FLORA NOVO-GALICIANA
rowed and convergent distally; wings adherent to the keel, with obliquely rhombic
blade 14-15 mm long, 9-12 mm wide, the claw 6-8 mm long; keel-blades ca 20-26
mm long, more or less united on the inner edges except at base and apex, gradually
tapered above the middle, strongly twisted laterally and curved inward (toward the
base) near the narrow rounded free tips; vexillar stamen abruptly bent near base,
strongly callose-thickened at the bend on the outer side; ovary sometimes with a
few fine hairs; style 28-30 mm long, not twisted with the keel but bent near the
middle and again below the apex, forming a squarish hook; fruit hard, green,
nearly terete, 1.5-2 cm thick, 12-17 cm long including the straight beak 1-1.5 cm
long; pedicels in fruit (6 collections) 1.5-3.5 mm thick; valves separating along both
sutures, curling when dry; seeds 8-10, biconvex, subquadrate or broadly reniform,
ca 8-10 mm long, 7-8 mm wide.
This is a fairly common plant on the Pacific slope and in the lower part of the
Santiago basin; throughout this range it seems morphologically uniform and constant in appropriate habitats. Less well known is the following, which has been
collected a few times at widely scattered localities in the upper basin of the Rio
Santiago:
Vigna strobilophora var. strobilophora. Vigna strobilophora B. L. Rob., as to
type. Ramirezella pubescens Rose, Contr. U.S. Nat. Herb. 8: 45. 1903. Phaseolus
pubescens (Rose) Morton, Contr. U.S. Nat. Herb. 29: 85. 1944. Ramirezella
strobilophora var. pubescens (Rose) Marechal et al. Taxon 27: 199. 1978.
Rocky slopes and ravines, in dry tropical deciduous forest with Bursera,
Ceiba, and arborescent Ipomoea, 1300-1800 m, in Nueva Galicia mostly in the
higher parts of the Santiago basin, flowering Aug-Sep.
Reported by Piper (1926) from Chihuahua; ?s Zac., Ags., Jal. (near Guadalajara, Pringle 5163, GH!, the holotype; US!, isotype; Pringle 4503).
Between Huejuquilla and Mezquitic (Rose 2562, US, cited by Rose as Ramirezella glabrata); near km 40, road to Calvillo W of Aguascalientes (McVaugh
18311); barranca of Rio Verde between Tepatitlan and Yahualica (McVaugh
17339); summit "6 mi" E of Atotonilco el Alto (McVaugh 17239); Mpio. Zapopan, Barranca de Huentitan (Diaz Luna 347).
The above collections are alike in having the primary floral bracts prominently acuminate with a sharp point, the body of the bracts 13-17 mm long, 1016 mm wide, and the bracteoles lance-ovate, 1.5-2 mm long. In most respects the
plants are indistinguishable from those of var. crassa, but some characters vary in
ways not yet fully understood:
Inflorescence-axis often densely pilose; ovary pilose on the valves; leaf-rachis
0.8-2 cm long; flowering axis not conspicuously thicker than the peduncle, the
flowering nodes 7-10 in 1 cm (in specimens from near Guadalajara) or 3-4 in 1
cm; pedicels (5-) 7-12 mm long; fruit thinly pilose or glabrescent; pedicels in fruit
(one collection) 1.5 mm thick.
The holotype of Ramirezella pubescens was from Guerrero (Nelson 2046,
US!). Marechal et al., after extensive numerical analysis of this and other taxa,
retained pubescens at the rank of variety, noting only that (in contrast to the
situation in var. strobilophora) the calyces and fruits were pubescent.
A specimen from summits SW of Autlan, 1300-1600 m, fl Nov (McVaugh
L
14180) was cited by M
men of this number at
species if one accepts
Leaves short, the r
lets silky-pilose beneat
a node; pedicels 4-8 r
mm wide, obtuse; flow
broadly ovate to subor
Vigna vexillata (L.
10: 191. 1845. Phaseolu
Roadsides, thicket
nearby foothills, sea-le
type presumably from
and variable, many var
Miramar, 15 km S
(Norris 13855); Valle d
Vallarta (Dieterle 3104c
Trailing herbaceous
with brownish hairs; p
to ovate, acute or acu
triangular, cordate-aun
1.5-3 mm thick at ba
often retrorsely setose;
a node, with a thick
bracteoles subulate, 32.5 mm long (-4 mm
panulate, 4.5-7 mm lo
8 mm long; flowers va
purple, or with purple
to 3 cm wide, with ret
longed into the auricles
keel broad, strongly i
laterally with a hollow
recurved toward the b
stamen not thickened a
7-11 cm long, 15-20-s
reniform or oblong, 4-
Vigna lonchophylla
Known only from
Feb 1901, by G. M. Er
Stems "sparsely n
"loose, 10 to 15 flower*