PANČIĆ`S SPECIMENS IN THE HERBARIUM COLLECTION OF

Transcription

Bulletin of the Natural History Museum, 2012, 5: 37-71.
Received 29 Nov 2012; Accepted 05 Dec 2012.
UDC: 582.3(497.113)
PANČIĆ’S SPECIMENS IN THE HERBARIUM COLLECTION OF
ANDREAS RAFAEL WOLNY
RANKO PERIĆ1, KSENIJA JAKOVLJEVIĆ2, VIDA STOJŠIĆ1, SNEŽANA VUKOJIČIĆ2
1
Institute for Nature Conservation of Vojvodina Province, Radnička 20a, 21000
Novi Sad, Serbia, e-mails: [email protected], [email protected]
2
Faculty of Biology, Institute of Botany and Botanical Garden “Jevremovac”, 11000
Belgrade, Serbia, e-mails: [email protected], [email protected]
The herbarium collection of Andreas R. Wolny in the Gymnasium of Sremski
Karlovci contains the specimens of well-known Serbian botanist Dr. Josif Pančić,
who donated them during and after his visit to the Gymnasium. Also, one small
part of the specimens from this collection is deposited in the herbarium of the
Institute for Nature Conservation of Vojvodina Province in Novi Sad. A list of 281
herbarium sheets with 295 taxa is presented for the first time to the botanical
community, with detailed information on where and when the plants were
collected, and notes on the specimens and revision data.
Key words: herbarium collection, Josif Pančić, Andreas R. Wolny, Gymnasium of Sremski Karlovci
INTRODUCTION
During his long botanical career Dr. Josif Pančić (1814-1888) investigated the plant life in the territories of present-day Serbia, Montenegro,
Romania, Bulgaria, and, to a lesser extent, Croatia, Hungary and Austria,
38
PERIĆ, R. ET AL.: PANČIĆ’S SPECIMENS IN HERBARIUM OF A. WOLNY
which resulted in the first national floras of the Principalities of Serbia
(Pančić 1874, 1884), Montenegro (Pančić 1875), Bulgaria (Pančić 1883,
1886), as well as many other botanical papers (Janković 1970). He gave
special attention to gathering a lot of plant material, which he later included
in his herbarium collection. The majority of these specimens are deposited
in the Herbarium of the Institute of Botany and Botanical Garden
“Jevremovac” - BEOU (Thiers 2012), within the special collection Herbarium Pancicianum (Vukojičić et al. 2011). A small part of his specimens is
deposited in the Herbarium of the Natural History Museum in Belgrade
(BEO, Thiers 2012) and in the herbarium collection of Andreas Rafael
Wolny in the Gymnasium of Sremski Karlovci. Also, an unknown number
of specimens are scattered through other Herbaria, mainly in central
Europe.
The herbarium collection of Andreas Rafael Wolny (1757-1827) in the
Gymnasium of Sremski Karlovci is the oldest known collection in Serbia,
with approximately 7300 sheets. Within this collection are stored specimens of vascular plants, mosses, lichens, fungi and algae, principally
originating from the territory of the former Habsburg Monarchy, including
the present-day province of Vojvodina (Serbia). It can be divided into three
different collections: authentic material of Wolny (197 sheets), general
school collection (about 6800 sheets) and material of Josif Pančić (270
sheets). After an official decision of the Institute for Protection and Study
of cultural monuments (nr. 240/50, March 13th, 1950), the herbarium
collection of Andreas Wolny in the Gymnasium of Sremski Karlovci was
inventoried and protected by the state (Marčetić 1950, 1952).
Wolny was highly respected both by his contemporaries (Waldstein &
Kitaibel 1802, Sadler 1820, 1830, Rochel 1828, Schuster 1829, Haberle
1830) and by subsequent botanists like Kanitz (1865), Schulzer et al.
(1866), Janka, Borbás and Jávorka (Bunke 1996) as well. Because of its
great historical and scientific importance many botanists (A. Rochel, A.
Kanitz, L. Adamović, A. Forenbacher, N. Košanin) visited Wolny’s collection (Marčetić 1950). During his botanical excursion to Srem in 1857 Pančić also visited and studied Wolny’s collection (Hirc 1919, Mitranović
1964). As was usual for that time he donated herbarium material during his
visit (he also sent a few specimens afterwards). This part of Pančić’s botanical legacy from the earliest known period of his work is presented for the
first time to the botanical community.
MATERIAL AND METHODS
The list of herbarium exsiccates of Josif Pančić in Andreas R. Wolny’s
collection is given according to their number in the Herbarium Inventory
BULLETIN OF THE NATURAL HISTORY MUSEUM, 2012, 5: 37-71.
39
(198-467) with a few additional sheets without number. All information
from labels is stored in the database in original form. Classification and
proper determination of each taxon were checked. Processed information
includes revised or properly and fully written names of taxa and authors, as
well as various abbreviations with updated toponymy and notes concerning
the sheets and specimens characteristics. If the identification was either
wrong, or still valid but now considered a synonym (Tutin et al. 1968,
1972, 1976, 1980, 1993), the appropriate taxon (specific or subspecific)
name was entered after the revision of the material. During the review of
the herbarium collection all specimens were photographed in the original.
Regarding the List of herbarium exsiccates in the following chapter, the
valid names were marked with bold letters. If names were misapplied or
representing synonyms, standard symbols were used (“–” misapplication of
name; “≡” isotypic synonyms; “=” heterotypic synonyms). In cases where
some taxa are considered to be particularly interesting from the chorological, historical or nature protection point of view their significance is
discussed in a special section (Obs.).
RESULTS AND DISCUSSION
List of herbarium exsiccates with revised nomenclature1
Achillea ptarmica L.
198 [Serbia] grows in Danube marshes [in Serbian].
Obs. With last known records from Knić and Gruža (Veljović 1967) this
species is supposed to be extinct in Serbia. All previous records in
Serbia are from areas near the Danube and Sava Rivers (Stevanović &
Vukojičić 1999).
Achillea umbellata DC. = A. umbellata Sibth. & Sm.
199 [Greece] Graecia. in M. Parnasso [Mount Parnassos], Aug 855
[1855].
Obs. Endemic to south-central and south Greece (Richardson 1976).
Aethionema saxatile RBr. [(L.) R. Br.]
200 [Serbia] rupestrib. [rupestribus] serpentinaceis ad Brđane [Brđani,
near Gornji Milanovac], Serb. Merid [meridionalis], Jun, l. Pančić.
Aira caespitosa L. [cespitosa] var. ochroleuca Rhb. ≡ Deschampsia
cespitosa (L.) Beauv. subsp. cespitosa
201 No data.
Note: There is one more label which is unreadable.
1
commutationem (com.) = acquired by exchange
40
Ajuga reptans L.
202 No data.
Allium moschatum L.
203 [Serbia] in saxosis ad Brđane [Brđani, near Gornji Milanovac],
Serb. Centr., Jul 858 [1858], l. Pančić; declivib. [declivibus] asperis ad
Ljuljaci [near Kragujevac], Serb. Centr. [centralis], Aug., l. Pančić.
Note: Plants from different localities are in the same sheet.
Alsine Parnassica Boiss. & Spr. = Minuartia stellata (E. D. Clarke) Maire
& Petitmengin
204 [Greece] Graecia. In cacumine M. Parnassi [Mt Parnassos], Aug.
855 [1855].
Obs. Endemic to mountains of Greece and south Albania (Halliday 1993).
Alyssum montanum L.
205 No data.
Alyssum rostratum Stev. [Steven] – A. tortuosum Willd.
206 [Serbia] arenosis Bjelobrdo [Deliblato Sand] in Banatus [Banat], Jul
1857, l. Pančić.
Anchusa Barrelieri [barrelieri] Boss. [(All.) Vitman]
207 [Serbia] collinis Serbidin prope Jagodina, Jun, l. Pančić.
Anchusa italica Retz. = A. azurea Miller
208 No data.
Anchusa officinalis L.
209 No data.
Androsace septentrionalis L.
210 Through the middle and eastern Europe [in Serbian].
Anemone sylvestris L.
211 [Serbia] Ad latera sylvarum ad Topčider prope Belgrad [Belgrade],
Maj 854 [1854], l. Pančić.
Note: It is considered that this species disappeared from Belgrade,
because in the last 50 years it has not been recorded in this area
(Jakovljević 2006).
Anthemis austriaca Jacq [Jacq.]
212 [Serbia] In arvis ad Deligrad [near Aleksinac], Serb. austral.
[australis], Jul 847 [1847], l. Pančić.
Anthriscus cerefolium L. [(L.) Hoffm.]
213 [Romania] in saxosis ad Ruskberg [Rusca Montană, Caraş-Severin
County] in Banatus [Banat], Jul 844 [1844], l. Pančić.
41
Apera spica-venti PB. [(L.) Beauv.]
214 No data.
“I guess that inflorescence branches are accidentally elongated and if
they are constantly long like this, obviously it is new!” [J. Pančić, in
Serbian].
Arceuthobium oxycedri MB. [(DC.) Bieb.]
215 [Serbia] super oxycedrum [Juniperus oxycedrus L.] in rupestrib.
[rupestribus] serpentinaceis M. Stolovi, Serb. merid. [meridionalis],
Aug., l. Pančić.
Artemisia scoparia WK. [Waldst. & Kit.]
216 [Serbia] through the tower of Slankamen [J. Pančić, in Serbian].
Asperula arvensis L.
217 [Hungary] M. Schwabenberg [Svábhegy] ad Budam [Buda: western
part of Budapest], Jul 837 [1837], l. Pančić.
Asperula hexaphylla All. – A. capitata Kit. ex Schult.
218 [Romania] Rupestrib. [rupestribus] M. Domugled [Domogled]
prope Mehadijam [Mehadia, Caraş-Severin County], Jun 844 [1844], l.
Pančić.
Obs. Endemic to south and east Carpathians (Ehrendorfer & Krendl
1976: 12).
Asplenium adianthum-nigrum [adiantum-nigrum L.] var. serpentini sensu
Pančić = A. cuneifolium Viv.
219 [Serbia] in rupestrib. [rupestribus] Serpentinaceis ad Brđane
[Brdjani, near Gornji Milanovac], Serb. Centr. [centralis], Jul 858
[1858], l. Pančić.
Asplenium trichomanes L.
220 [Serbia] in rupib. [rupibus] ad Topčider [Belgrade], Jul 854 [1854],
l. Pančić.
Astragalus contortuplicatus L.
221 [Serbia] Arenosis ad Danubium [Danube] in procul a Belgrado
[Belgrade], Aug. 1857, l. Pančić.
Astragalus dasyanthus Pall.
222 [Serbia] (florens) Arenosis Bjelobrdo in Banatus [Deliblato Sand],
Jul 1857; (fructifer) ad ripas arenosis fluvii Ibar [Valley of river Ibar],
Serb. merid. [meridionalis], Jul 856 [1856], l. Pančić.
Astragalus hamosus L. var. multiflorus Griseb.
223 [Serbia] pratis siccis ad Kajaburun [Karaburma] prope Belgrad
[Belgrade], Jun, l. Pančić.
42
Astrantia minor L.
224 [Italy] Grenier in Pedemontio [near Oyace, Valle d’Aosta], com.
Pančić.
Athamantha mathioli Wulf. = A. turbith (L.) Brot. subsp. hungarica
(Borbás) Tutin
225 [Romania] rupib. [rupibus] calcareis ad Mehadia [Mehadia, Caraş-Severin County], Jul 1845, l. Pančić.
Avena neglecta Savi = Trisetum paniceum (Lam.) Pers.
226 [Serbia?] rare... [unreadable] [J. Pančić, in Serbian] [Srem?]
Note: Besides two fragments of T. paniceum, there are also two
fragments of Festuca sp.
Avena tenuis Mnch. = Ventenata dubia (Leers) Coss.
227 No data.
Azalea procumbens L. ≡ Loiseleuria procumbens (L.) Desv.
228 [Romania] In cacumine M. Gugu [Gugu peak, Godeanu mountain
in Caraş-Severin County] in Banatus [Banat], Aug. 844 [1844], l.
Pančić.
Barkhausia setosa Rchb. [(Haller f.) DC.] ≡ Crepis setosa Haller f.
229 No data.
Barkhausia rhoe[a]difolia MB. [(M. Bieb.) M. Bieb.] ≡ Crepis foetida L.
subsp. rhoeadifolia (Bieb.) Čelak.
230 No data.
Berula angustifolia [Koch in Mert. & Koch] = B. erecta (Huds.) Coville
231 No data.
Bifora radians MB. [M. Bieb.]
232 [Serbia] in arvis ad Banja [Sokobanja near Aleksinac], Serb. austral.
Blitum virgatum L. = Chenopodium foliosum (Moench) Asch.
233 No data.
Bromus asper Murr. = B. ramosus Hudson
234 [Serbia] ad rivum prope Banja [Sokobanja near Aleksinac], Serb.
austral. [australis], Jun 847 [1847], l. Pančić.
Bromus vernalis Panč. [Pančić] – B. erectus Hudson
235 [Serbia] Rupestrib. [rupestribus] serpentinaceis ad Brđane [Brdjani,
near Gornji Milanovac], Serb. Central. [centralis], Apr, l. Pančić.
Bruckenthalia spiculiflora Rchb [spiculifolia (Salisb.) Reichenb.]
236 [Serbia] M. Crni vr’ [Crni vrh] ultra confinium C. Gurgusovac
[Knjaževac], Jul, l. Pančić.
43
Bupleurum apiculatum Gm. [Friv.]
237 [Serbia] In arvis sterilibus ad Deligrad [near Aleksinac], Serb.
austral. [australis], Jul, l. Pančić.
Bupleurum falcatum L.
238 No data.
Bupleurum gerardi sensu Pančić = B. praealtum L.
239 [Serbia] In graminosum sub M. Avala [Belgrade], Serb. Sept.
[septentrionalis], Jul 857 [1857], l. Pančić.
Bupleurum glumaceum Sibth. [Sibth. & Sm.]
240 [Greece] Attica [Attikí], In M. Lycabettus [in Athens], Apr. 855
[1855].
Bupleurum junceum sensu Pančić = B. praealtum L.
241 No data.
Bupleurum odontites L. – B. baldense Turra subsp. gussonei (Arcangeli)
Tutin.
242 [Serbia] Declivib. [declivibus] Asperis ad Raška, Serb. merid.
[meridionalis], Jul 1856, l. Pančić.
Bupleurum semicompositum L.
243 [Hungary] In graminosis salin [salinis] ad Libiscum [Tibiscum =
Tisa] non procul Szegedino [Szeged], Aug. 840 [1840], l. Pančić.
Obs. Corresponds well to the specimen from herbarium of C. L.
Willdenow (B-W-05659-02 0) (Röpert 2000). Nevertheless, in Europe
this species is confined to the Mediterranean and Atlantic coastal areas
(Snogerup & Snogerup 2001: 276) and the record is likely to be
erroneous.
Calamintha patavina Host. [(Jacq.) Host] var. rotundifolia Benth. = Acinos
alpinus (L.) Moench subsp. majoranifolius (Mill) P. W. Ball
244 [Serbia] In saxosis serpentinaceis ad Brđane [Brdjani, near Gornji
Milanovac], Serb. Centr. [centralis] Jul 854 [1854], l. Pančić.
Note: With one fragment of Acinos arvensis (Lam.) Dandy.
Campanula carpathica [carpatica] L. [L. fil.] = C. carpatica Jacq.
245 [Romania] In M. Magura Carpathosum [Măgura, Braşov], Septentr.
[septentrionalis], Aug 1840, l. Pančić.
Campanula crassipes Heuff.
246 [Serbia] Ad rupis veteris arvis Golubac, Serb. borealis, Sept. 850
[1850], l. Pančić.
Campanula linifolia – Campanula sp. [C. rotundifolia agg.]
247 No data.
“Probab. [probabiliter] Campanula linifolia!” [J. Pančić]
44
Campanula rapunculoides L.
248 No data.
Cardamine glauca Spr. [Sprengel] – C. pancicii Hayek
249 [Serbia] Cacumine M. Kopaonik, Serb. merid. [meridionalis], Jul, l.
Pančić.
Carex acuta L. var. chlorostachya Rchb.
250 No data.
Carex hirta L.
251 No data.
Carex muricata L. var. virens L. [(Steud.) Lam. ex Rchb.] – C. spicata
Hudson
252 No data.
Carex riparia Curtis
253 No data.
Carlina acanthifolia All.
254 [Serbia] In pratis montanis ad Topčider prope Belgrad [Belgrade],
Aug. 858 [1858], l. Pančić.
Chenopodium polyspermum L. var. acutifolium (W. K) [(Sm.) Kostel.]
255 No data.
Chrysanthemum alpinum L. ≡ Leucanthemopsis alpina (L.) Heywood
256 [Romania] In herbidis alpis Branu [Bran, Braşov] in Banatus
[Banat], Aug, 845 [1845], l. Pančić.
Centaurea axillaris W. = C. triumfetti subsp. aligera (Gugler) Dostál
257 No data.
Centaurea scabiosa L.
258 No data.
“Teratologica modificatio/ capitulus” [J. Pančić]
Cerastium glomeratum Thuill. – C. fontanum Baumg. subsp. vulgare
(Hartm.) Greuter & Burdet
259 No data.
Note: With one specimen of Silene vulgaris (Moench) Garcke.
Cerastium grandiflorum WK. [Waldst. & Kit.] – C. banaticum (Rochel)
Heuff. subsp. banaticum
260 [Serbia] Rupestrib. [rupestribus] M. Starica prope Majdanpek, Serb.
Boreal [borealis], Maj, l. Pančić.
Cerinthe aspera Roth. – Cerinthe major L.
261 No data.
45
Cicer arietinum L. [arietum L.]
262 No data.
Obs. Probably introduced by Turks and cultivated in Serbia during the
Ottoman period. Seeds are rich in protein content and they were milled
and used as sourdough admixture (Pančić 1856). In recent times roasted
seeds are still popular in some parts of Serbia (“leblebija”).
Cirsium pannonicum Gaud. = C. pannonicum (L. f.) Link
263 [Hungary] Pratis sylvestrib. [sylvestribus] M. Schwabenberg [Svábhegy] ad Budam [Buda: western part of Budapest], Jul 1838, l. Pančić.
Convolvulus sylvaticus WK [Kit.] ≡ Calystegia silvatica (Kit.) Griseb.
264 [Romania] Ad margines sylvarum rupestribus ad Mehadiam
[Mehadia, Caraş-Severin County], Jun. 1845, l. Pančić.
Coronilla minima L.
265 [Italy] St. Germani in Pedomonttia [Piedmont], com. Pančić.
Cotoneaster vulgaris Lindl [Lindl.] = C. integerrimus Medik.
266 [Serbia] M. Kopaonik, Serb. merid. [meridionalis], Jul 856 [1856],
l. Pančić.
Crataegus melanocarpa MB. [M. Bieb.] = C. pentagyna Waldst. & Kit. ex
Willd.
267 [Serbia] In vepretis ad Trnava [near Raška], Serb. Centr. [centralis],
Jun. Aug., l. Pančić.
Crataegus nigra WK. [Waldst. & Kit.]
268 [Serbia] Insularis ad Savum, Jun Aug., l. Pančić.
Crepis biennis L.
269 No data.
Crepis pulchra L.
270 [Serbia] In arvis ad Banja [Sokobanja near Aleksinac], Serb. austr.
Crocus banaticus Heuff [Heuffel] = C. vernus (L.) Hill subsp. vernus
271 [Romania] Sakul [Sacul near Rusca Montană] in Banatus [Banat],
Apr. 844 [1844], l. Pančić.
Cytisus biflorus ĽHer [Ľ Hér.] = Chamaecytisus ratisbonensis (Schaeffer)
Rothm.
272 No data.
“Syn. C. ratisbonensis Schaef. var. minor Koch. Is it from Srem? It was
mixed with nigricans” [Lembotropis nigricans (L.) Griseb.] [J. Pančić,
in Serbian].
46
Daucus Carota L. [carota]
273 No data.
Danthonia provincialis DC. = D. alpina Vest
274 [Serbia] Pratis sylvestrib. [sylvestribus], ad Trmbas [near Kragujevac], Serb. centralis, Jun, l. Pančić.
Dianthus leptopetalus W. [Willd.]
275 [Serbia] Collinis arvidis ad Deligrad [near Aleksinac], Serb. austral.
[australis], Jul, l. Pančić.
Dianthus superbus L.
276 No data.
Digitalis ferruginea L.
277 [Serbia] In collib. [collibus] siccis ad Jagodina, Serb. centr.
[centralis], Jun 847 [1847], l. Pančić.
Epilobium Dodonei Vill. [dodonaei Vill.]
278 [Romania] Ad ripas Ren mare [probably river Râul Mare (Retezat),
Hunedoara County], in Transylvania, Jul 844 [1844], l. Pančić.
Epilobium palustre L.
279 No data.
Equisetum pratense Ehrh.
280 [Serbia] Ad ripas Danubii [Danube] prope Belgrad [Belgrade], Jun
854 [1854], l. Pančić.
Note: Only label without specimen in the same sheet with next
specimen.
Obs. The presence of this species in Serbia is doubtful (Jalas &
Suominen 1972).
Equisetum telmateja L. [telmateia Ehrh.]
280 [Serbia] Ad fossas prope Rakovica [Belgrade], Serb. bor. [borealis],
aug 855 [1855], l. Pančić.
Note: In the same sheet with previous specimen.
Erigeron alpinus L.
281 [Romania] M. Branu in Banatus [Bran, Braşov], Aug. 1843, l.
Pančić.
Eryngium serbicum Pančić
282 [Serbia] Collib. [collibus] herbidis ad Dobrača [near Kragujevac],
Serb. centr. [centralis], Aug., l. Pančić.
Obs. From locus classicus (Pančić 1856). Endemic to Serbia (Chater
1968: 323)
47
Eryngium tricuspidatum L. sensu Pančić = E. palmatum Pančić & Vis.
283 [Serbia] In glareosis M. Zlatibor [near Užice], Serb. merid.
[meridionalis], Jul 856 [1856], l. Pančić; Sylvestrib. [sylvestribus]
saxosis ad Ravanica [near Ćuprija], Serb. centr. [centralis], Jul, l.
Pančić.
Note: both labels are within the same sheet
Obs. Ravanica is locus classicus for this species (Visiani & Pančić
1870). Endemic to the central part of the Balkan peninsula (Chater
1968).
Erysimum angustifolium Ehrh. = Syrenia cana (Piller & Mitterp.) Neilr.
284 [Serbia] Arenosis ad Radujevac, Serb. austral. [australis], Jun, l.
Pančić.
Erysimum cuspidatum DC. [(M. Bieb.) DC.]
285 [Serbia] Ruderalis ad Kučajna, Serb. boreal. [borealis], Jun, l.
Pančić.
Erysimum helveticum DC. [(Jacq.) DC.] – E. linariifolium Tausch [rev. M.
Niketić]
286 [Serbia] Rupestrib. [rupestribus] serpentinaceis ad Brdjane [Brđani,
near Gornji Milanovac], Serb. Centr. [centralis], Maj, l. Pančić.
Erythraea emarginata Rchb. [Waldst. & Kit.] = Centaurium pulchellum
(Swartz) Druce
287 [Serbia] in uliginosis ad Mokri lug prope Belgrad [Belgrade], Aug.
858 [1858], l. Pančić.
Erythraea pulchella Fries [(Swartz) Fries] ≡ Centaurium pulchellum
(Swartz) Druce
288 [Serbia] In arenosis Morava [near Ćuprija], Serb. Centr. [centralis],
Jun 847 [1847], l. Pančić.
Ervum ervilia L. ≡ Vicia ervilia (L.) Willd.
289 [Serbia] In arvis circa Belgrad [Belgrade], Jul 857 [1857], l. Pančić.
Obs. This species has been cultivated in present day Turkey since the
7th millenium BC (Zohary & Hopf 2000) . It was probably introduced
into Serbia during the Ottoman period.
Ervum tetraspermum L. ≡ Vicia tetrasperma (L.) Schreber
290 [Serbia] In graminosis ad Kragujevac, Serb. Centr. [centralis], Jun
849 [1849], l. Pančić.
Eudianthe coeli-rosa Rchb [Rchb.] = Silene coeli-rosa (L.) Godron
291 [Serbia] Inter segetes ad Žarkovo in procul a Belgrado [Belgrade],
Jun / 1857, l. Pančić.
Obs. In Serbia cultivated for ornament.
48
Euphorbia lucida WK [Waldst. & Kit.]
292 No data.
Euphorbia stricta L. = E. serrulata Thuill.
293 No data.
Euphrasia lanceolata Gaud [Gaudin] ≡ Odontites lanceolata (Gaudin)
Rchb.
294 [Italy] Apiemont [?] in Pedemontio [Piedmont], com Pančić.
Evax perpusilla Boiss & Heldr. [Boiss. & Heldr.]
295 [Greece] Graecia. In M. Parnass [Mount Parnassos], Jul 855 [1855].
Obs. Endemic to mountains of south and central Greece (Holub 1976).
Foeniculum officinale L. = F. vulgare Miller
296 No data.
Fragaria vesca L.
297 [Serbia] Rakovac am Berge (Schan 2 / Fr. parviflora).
Fritillaria tenella MB. [M. Bieb.] – F. montana Hoppe
298 [Serbia] M. Medvednik, Serb. merid. [meridionalis], Jul 856 [1856],
l. Pančić.
Obs. New record for Serbia. Pančić did not include this locality in his
“Flora principatus Serbiae” (Pančić 1874) nor was it mentioned by
Tomović et al. (2009).
Fraxinus ornus L.
299 [Serbia] M. Avala prope Belgrad [Belgrade], Maj 854 [1854], l
Pančić.
Gagea arvensis Schult. [(Pers.) Schult.] ≡ G. arvensis (Pers.) Dumort.
300 [Serbia] In arvis ad Belgrad [Belgrade], Apr 857 [1857], l Pančić.
Galium boreale L.
301 [Hungary] In pratis humidis prope Pesthinum [Pest: eastern part of
Budapest], Jun 842 [1842], l. Pančić.
Galium aristatum sensu Pančić = Galium sp. (G. ser. Nemoralia M.
Popova ex Ehrend.)
302 [Serbia] Topčider [Belgrade].
Obs. Incomplete specimen, probably G. pseudaristatum Schur.
Galium pedemontanum All. [(Bellardi) All.] ≡ Cruciata pedemontana
(Bellardi) Ehrend.
303 [Serbia] Jagodina, on Đurđevo hill, Maj 847 [1847], Pančić [in
Serbian].
49
Galium tricorne WK [auct.] = G. tricornutum Dandy
304 [Serbia] In arvis ad Beloševac [near Kragujevac], Serb. Centr. [centralis], Jun 852 [1852], l. Pančić.
Galium vernum Scop. = Cruciata glabra (L.) Ehrend.
305 [Serbia] Ad margines sylvarum prope Drača [near Kragujevac],
Serb. Centr. [centralis], Maj. 850 [1850], l. Pančić.
Gentiana ciliata L. ≡ Gentianella ciliata (L.) Borkh.
306 [Austria] In graminosis ad Baden prope Viennam [Wien], Aug 844
[1844], l. Pančić.
Gentiana obtusifolia Willd. ≡ Gentianella germanica (Willd.) E. F. Warb.
307 [Romania] M. Ruszka [Ruszka Mountains, Caraş-Severin County]
in Banatus [Banat], Jul 844 [1844], l. Pančić.
Gentiana punctata L.
308 [Romania] M. Branu [Bran, Braşov] in Banatus [Banat], Aug 844
[1844], l. Pančić.
Geranium fasciculatum Panč. [Pančić] = G. asphodeloides Burm. fil.
309 [Serbia] Collinis herbidis prope Kragujevac, Serb. Centr. [centralis],
Maj, l. Pančić.
Geranium palustre L.
310 No data.
Geranium Robertianum L. [robertianum L.]
311 No data.
Geranium umbrosum WK. [Waldst. & Kit.] = G. pyrenaicum Burm. f.
312 No data.
Glyceria aquatica Presl. [(L.) J. & C. Presl.] ≡ Catabrosa aquatica (L.) P.
Beauv.
313 [Hungary] Ad ripas Danubii prope Pesthinum [Pest: eastern part of
Glycyrrhiza glabra L.
314 No data.
Gypsophila illyrica Sm. [Sibth. & Sm.] = Petrorhagia illyrica (Ard.) P. W.
Ball & Heywood
315 [Serbia] Declivib [-us] arvidis ad Banja Još. [Jošanička banja, near
Kopaonik], Serb. merid. [meridionalis], Jul 1856, l. Pančić.
Herniaria glabra L.
316 [Serbia] In arenosis Morava ad Cuprija [Ćuprija], Serb. Centr.
[centralis], Aug. 847 [1847], l. Pančić.
50
Herniaria hirsuta L.
317 [Serbia] In collinis siccis ad Deligrad [near Aleksinac], Serb.
austral. [australis], Jul 847 [1847], l. Pančić.
Hieracium prealtum Koch [praealtum Vill. ex Gochnat] var. bauhini
[(Schultes) Koch] = H. praealtum Vill. ex Gochnat subsp. bauhinii
(Besser) Petunin.
318 No data.
Hieracium prealtum Koch. [praealtum Vill. ex Gochnat] var. Decipiens
[decipiens W. D. J. Koch]
319 No data.
Hieracium prealtum Koch. [praealtum Vill. ex Gochnat] var. florentinum
Koch. [Koch] = H. praealtum Vill. ex Gochnat
320 No data.
Himanthoglossum [Himantoglossum] hircinum MB [(L.) Sprengel]
321 [Serbia] In sylvestrib [sylvestribus] ad Topčider prope Belgrad
[Belgrade], Jul, l. Pančić.
Hippuris vulgaris L.
322 [Serbia] In uliginosis ad Danubium [Danube] prope Belgrad
[Belgrade], Maj 854 [1854], l. Pančić.
Obs. Critically endangered species in Serbia (Vučković & Panjković
1999).
Hottonia palustris L.
323 No data.
Hypericum nummulariefolium L. [nummularium L.]
324 [France] Chambéry [Chambéry, Savoie] in Pedemontio, Jul, com
Pančić.
I. [Inula] ensifolia L.
325 No data.
Inula vaillantii Vill. [(All.) Vill.] = I. helvetica Weber
326 [Italy] Chambéry [Chambéry, Savoie] in Pedemontio, Aug, com
Pančić.
Iris graminea L.
327 [Serbia] In graminosis sylvestrib [sylvestribus] ad Topčider [Belgrade], Jun 854 [1854], l. Pančić.
Iris lutescens Lam. – I. reichenbachii Heuff.
328 [Serbia] Rupestrib. [rupestribus]s..iticis [sylvaticis?] ad Borač [near
Kragujevac], Serb. Centr. [centralis], Maj, l. Pančić.
51
Jasione supina Sieb. = J. laevis subsp. orbiculata (Griseb. ex Velen.) Tutin
329 [Serbia] M. Kopaonik, Serb. merid [meridionalis], Jul, l. Pančić.
Folia primordialia / speciei Aceri!
330 No data.
“If am not mistaken!” [J. Pančić, in Serbian].
Juncus obtusiflorum [obtusiflorus] Ehrh. = J. subnodulosus Schrank
331 [Hungary] In turfosis Rakos prope Pesth. [Pesthinum] [Pest: eastern
Knautia hybrida Coult. [(All.) Coult.] var. bidens Sm. = K. integrifolia (L.)
Bertol.
332 [Serbia] Pratis arenosis ad Negotin, Serb. austral. [australis], Jun, l.
Pančić.
Lathraea squamaria L.
333 No data.
Lathyrus Nissolia L. [nissolia]
334 [Serbia] In collinis ad Topčider prope Belgrad [Belgrade], Jun, l.
Pančić.
Lathyrus palustris L.
335 No data.
Leontodon villarsii DC. [(Willd.) Loisel.] = L. hirtus L.
336 [Italy] Anex Rochelles [?] in Pedemontio [Piedmont], Jul, com
Pančić.
Obs. Endemic to south France and Italy (Finch & Sell 1976).
Limodorum abortivum Sm. [(L.) Sw.]
337 [Serbia] In sylvaticis umbrosi [umbrosis] ad Topčider [Belgrade],
Jun 857 [1857], l. Pančić.
Linum hirsutum L.
338 [Serbia] In pratis circa Belgrad [Belgrade], Jun 847 [1847], Pančić.
Lloidia graeca Schrb. [Lloydia graeca (L.) Endl. ex Kunth] ≡ Gagea
graeca (L.) A. Terracc.
339 [Greece] Attica [Attikí], in collinis, Apr. 855 [1855].
Obs. Endemic to south Greece and Aegean region (Richardson 1980).
Lonicera Caprifolium L. [caprifolium]
340 [Serbia] Ad margines sylvar [“sylvarum”] ad Topčider prope
Belgrad [Belgrade], Maj 854 [1854], l. Pančić.
Luzula forsteri DC. [(Sm.) DC].
341 [Serbia] Sylvaticis M. Crni vr [Crni Vrh near Jagodina] Serb. Centr.
[centralis], Maj, l. Pančić.
52
Luzula pilosa Gaud. [(Sm.) DC].
342 [Serbia] M. Crni vr [Crni Vrh near Jagodina] Serb. Centr.
[centralis], Apr. 850 [1850], l. Pančić.
Lychnis dioica L. ≡ Silene dioica (L.) Clairv.
343 No data.
Lysimachia nemorum L.
344 [Serbia] Sylvaticis M. Murtenica [near Nova Varoš], Serb. Merid.
[meridionalis], Jul 856 [1856], l. Pančić.
Obs. Most likely new record for Serbia. This species is rare with records
from the vicinity of Bajina Bašta (Derventa, Zaovine, Mitrovac) and
near Povlen mountain (“Mađarvode”) (Pančić 1874, Čolić 1953, Nikolić
1973).
Lysimachia punctata L.
345 No data.
Lythrum graefferi Ten. = L. junceum Banks & Sol.
346 [France] Bayonne in Gallia, com. Pančić.
Malcolmia graeca Boiss. [Boiss. & Spruner]
347 [Greece] Graecia. In M. Lycabettus [near Athens], Jun 855 [1855].
Obs. Endemic to central, south and northwestern Greece with one
station in south Albania (Ball & Akeroyd 1993).
Medicago carstiensis Jacq.
348 [Serbia] Ad latera sylvarum M. Ozren [near Sokobanja], Serb.
austral. [australis], Jul, l. Pančić.
Medicago falcata L. var. major Koch ≡ M. sativa L. subsp. falcata (L.)
Arcang. f. major Koch
349 No data.
Medicago lupulina L.
350 No data.
Medicago prostrata Jacq.
351 [Serbia] Collib. [collibus] asperis ad Dobrača [near Kragujevac],
Serb. Centr. [centralis], Jun. l. Pančić.
Moehringia trinervia Clair. [(L.) Clairv.]
352 No data.
Melampyrum nemorosum L.
353 No data.
Meum Mutel[l]ina Gartn. [(L.) Gaertn.] – Carum graecum Boiss. & Heldr.
354 [Serbia] In cacumine M. Rtanj, Serb. Austral. [australis], Jul., l.
Pančić.
53
Myosotis versicolor Pers [(Pers.) Sm.] – M. ramosissima Rochel
355 [Serbia] In pratis siccis circa Čačak, Serb. Centr. [centralis], Apr., l.
Pančić.
Nepeta lanceolata Lam. = N. nepetella L.
356 [France] In Labandia [?], com Pančić.
Notochlaena Marantae RBr. [Notholaena marantae (L.) Desv.] ≡
Cheilanthes marantae (L.) Domin
357 [Serbia] In rupestrib. [rupestribus] Serpentinaceis ad Brđane
[Brđani, near Gornji Milanovac], Serb. austr. [australis], Jul 858 [1858],
l. Pančić.
Oenanthe peucedanifolia Poll.
358 No data.
Ononis repens L.
359 No data.
“Keep in mind this plant, seems that it shows some peculiarities, better
exemplars are needed” [J. Pančić, in Serbian].
Onosma stellulatum WK [stellulata Waldst. & Kit.] – O. heterophylla
Griseb. [rev. M. Niketić]
360 [Serbia] Montosis asperis ad Vraćevšnica [near Kragujevac], Serb.
central. [centralis], Jun, l. Pančić.
Orlaya grandiflora Hoffm. [(L.) Hoffm.]
361 No data.
Ornithogalum chloranthum Saut. = O. boucheanum Asch.
362 [Serbia] In uno prato in procul in a Belgrado [Belgrade], Maj, l.
Pančić.
Ornithogalum exscapum Ten. – O. orthophyllum Ten. subsp. kochii (Parl.)
Zahar.
363 [Serbia] In arvis circa Belgrad [Belgrade], Apr., l. Pančić.
Ornithogalum nutans L.
364 No data.
Paliurus australis G. [Gaertn.] = P. spina-christi Miller
365 [Serbia] (čalija, drač), Topčider close to Belgrade [in Serbian], Sep.
846 [1846], Pančić.
“Sand in Banat”, [added to label by anonymous].
Pedicularis Barrelieri Rchb. = P. ascendens Schleich. ex Gaudin
366 [Italy] Grenier in Pedemontio [near Oyace, Valle d’Aosta], com
Pančić.
Obs. Endemic to Alps (Mayer 1972).
54
Pedicularis cenisia Gaud.
367 [France] M. Cenis [Mont Cenis, Savoie] in Pedemontio, com
Pančić.
Obs. Endemic to France and Italy (Mayer 1972).
Pedicularis comosa L.
368 [France] Chamberg [Chambéry, Savoie] in Pedemontio, com
Pančić.
Pedicularis foliosa L. – P. exaltata Besser
369 [Romania] In pratis montanis ad Štje [?] in Transylvania, Jul 844
[1844], l. Pančić.
Pedicularis gyroflexa Vill [Vill.]
370 [Italy] Grenier in Pedemontio [near Oyace, Valle d’Aosta], com.
Pančić.
Peucedanum austriacum Koch [(Jacq.) W. D. J. Koch]
371 No data.
Phyteuma fistulosum Host = P. orbiculare L.
372 [Hungary] Lindenberg prope Budam [Buda: western part of
Phyteuma hemisphaericum L. – P. confusum A. Kerner
373 [Romania] M. Branu [Bran, Braşov] in Banatus [Banat], Aug. 845
[1845], l. Pančić.
Phyteuma limonifolium Sm. [Sibth. & Sm.] = Asyneuma limonifolium (L.)
Janch.
374 [Serbia] Asperis M. Stol, Serb. merid. [meridionalis], Jul, l. Pančić.
Pimpinella magna L. ≡ P. major (L.) Hudson
375 [Austria] Leopoldiberg [Leopoldberg] prope Viennam [Wien], Aug.
845 [1845], l. Pančić.
Plantago montana L. = P. atrata Hoppe
376 [Romania] In pascuis uliginosis M. Sidjimir [?] in Banatus [Banat],
Aug. 844, l. Pančić; Alpinis Branu [Bran, Braşov] in Transylvania,
Aug., l. Pančić.
Plantago carinata Schrad [Schrad. ex Mert & W. D. J. Koch, non Moench]
= P. holosteum Scop.
377 [Serbia] Asperis serpentinaceis M. Zlatibor [near Užice], Serb.
merid. [meridionalis], Jul, l. Pančić.
Plantago Cynops L. = P. sempervirens Crantz
378 [Austria] M. Kalvarienberg prope Baden in Austria, aug 1845, l.
Pančić.
55
Podospermum Jacquinianum Koch = Scorzonera cana (C. A. Mey.) O.
Hoffm.
379 No data.
Polygala alpestris Rchb [Rchb.]
380 [Serbia] M. Kopaonik Serbiae meridionalis, Aug., l. Pančić.
Polygala amara Jacq [L.]
381 No data.
Potentilla petiolulata Gaud. = P. caulescens L. [subsp. petiolulata sensu
Nyman]
382 [France] Chambery [Chambéry, Savoie] in Pedemontio, com.
Pančić.
Polygonum bistorta L.
383 [Serbia] –vig [?] in Syrmio [Srem].
Obs. This record is uncertain, because Srem is ecologically an unusual
habitat for this montane species.
Primula pedemontana Thom. [E. Thomas ex Gaudin]
384 [France] M. Cerni [Mont Cenis, Savoie] in Labandia [?], com
Pančić.
Obs. Endemic to southwestern Alps and Cordillera Cantábrica (Valentine & Kress 1972).
Pyrola secunda L. ≡ Orthilia secunda (L.) House
385 [Serbia] In fagetis M. Rasovati kamen [near Knjaževac], Serb. aust.
[australis], Aug 853 [1853], l. Pančić.
Pyrola media Sw.
386 [Romania] In sylvaticis ad Ruszkberg [Rusca Montană, Caraş-Severin County], Banatus [Banat], Jun 844 [1844], l. Pančić.
Pyrola uniflora L. ≡ Moneses uniflora (L.) A. Gray
387 [Romania] Convalle alpina K’m [?] in Banatus [Banat], Aug 844
[1844], l. Pančić.
Pyrus amygdaliformis Vill.
388 [Serbia] Collinis ad Deligrad [near Aleksinac], Serb. austral
[australis], Maj Jul, l. Pančić.
Peucedanum Chabraei Rchb [(Jacq.) Rchb.] = P. carvifolia Vill.
389 No data.
Quercus brutia Ten. = Q. robur L. subsp. brutia (Ten.) O. Schwarz
390 No data.
56
“Could be a Kitaibel Q. pendulina” [J. Pančić, in Serbian] [Q. ×
pendulina Kit. (Q. robur subsp. robur × Q. virgiliana / pubescens sensu
Soó)].
Quercus sessiliflora Sm. [Salisb.] = Q. petraea (Matt.) Liebl.
391 No data.
“All appearances that oak species are not possible to determine without
fruits” [J. Pančić, in Serbian].
Queria hispanica Loefl. [L.] = Minuartia hamata (Hausskn.) Mattf.
392 [Serbia] asperis ad Topčider prope Belgrad [Belgrade], maj, l.
Pančić.
Rhamnus alpina L. sensu Pančić = R. alpinus L. subsp. fallax (Boiss.)
Maire & Petitm.
393 [Serbia] in sylvis saxosis ad Beli potok [near Belgrade], serb. merid.
[meridionalis], Jul, 846 [1846], l. Pančić.
Rhamnus rupestris Scop [Scop.] ≡ Frangula rupestris (Scop.) Schur
394 [Serbia] in rupib. [rupibus] ad Ravanica C. [coenobium], cuprianensis [near Ćuprija], Apr. 847 [1847], l. Pančić.
Rhamnus tinctoria WK [tinctorius Waldst. & Kit.] ≡ R. saxatilis Jacq.
subsp. tinctorius (Waldst. & Kit.) Nyman
395 [Serbia] In collib. [collibus] supra Topčider [Belgrade], Maj. Aug, l.
Pančić.
Ranunculus auricomus L. var. grandiflorus Rchb [Rchb.]
396 [Serbia] Bukovački branik prope Jagodina, Maj, l. Pančić.
Ranunculus millefoliatus Vahl
397 [Serbia] infra collem Golobrdo prope Knić, Serb. central. [centralis], Maj, l. Pančić.
Ranunculus pedatus WK [Waldst. & Kit.]
398 [Hungary] In prato ad Stadtmeyerhof [part of/near Budapest], Maj
1837, l. Pančić.
Rosa lutea Mill. – Rosa sp. (R. canina group)
399 [Serbia] collinis ad Kragujevac Serb. Centr. [centralis], Maj 850
[1850], l. Pančić.
Obs. Incomplete plant material!
Ruscus aculeatus L.
400 [Serbia] In fagetis ad Ramaća [near Kragujevac] Serb. Centr.
[centralis], Maj 850 [1850], l. Pančić.
Salvia horminoides Pour. = S. verbenaca L.
401 [France] Toulonne [Toulon] in Gallia, com Pančić.
57
Saponaria glutinosa MB. [M. Bieb.]
402 [Serbia] Declivib. [Declivibus] saxosis ad Banja [Sokobanja near
Aleksinac (Pančić 1874: 172], Serb. austral. [australis], Jul, l. Pančić.
Sarothamnus vulgaris Willd. [Wimmer] = Cytisus scoparius (L.) Link
subsp. scoparius
403 No data.
“cult?” [J. Pančić].
Saxifraga aizoides L.
404 [Romania] M. Brustur [Brusturi peak, Bihor County] in Banatus
[Banat], Aug 844 [1844], l. Pančić.
Saxifraga Aizoon Jacq. = S. paniculata Mill.
405 [Romania] M. Domugled [Domogled] ad Mehadiam [Mehadia,
Caraş-Severin County], In Banatus [Banat], Jun 844 [1844], l. Pančić.
Saxifraga biflora All. subsp. biflora
406 [Austria] M. Grossglockner in Carinthia, Aug 844 [1844], l. Pančić.
Obs. Endemic to Alps (Webb 1993).
Saxifraga bryoides L.
407 [Romania] M. Brustur [Brusturi peak, Bihor County] in Banatus
Saxifraga controversa Strb. [Sternb.] = S. adscendens L.
408 [Romania] M. Nedjea [Nedeia peak, Munţii Căpăţânii in Parâng
mountains group] in Banatus [Banat], Aug 844 [1844], l. Pančić.
Saxifraga stellaris L.
409 [Romania] M. Branu [Bran, Braşov] in Banatus [Banat], Aug. 844
[1844], l. Pančić.
Scabiosa banatica WK [Waldst. & Kit.] = S. columbaria L.
410 [Romania] In lapidosis ad Kazan [Đerdap Gorge] in Banatus
Scabiosa ucranica L. = S. argentea L.
411 [Serbia] pratis arvidis ad Umka [near Belgrade], Serb. boreal
[borealis], Jul, l. Pančić.
Scleropoa maritima Parlat [(L.) Parl.] ≡ Cutandia maritima (L.) Barbey
412 [Greece] Attica [Attikí], in arenosis Phaleri [Phalerum: Phaleron or
Faliron Bay, near Athens], Apr. 855 [1855].
Scandix Pecten L. – S. pecten-veneris L. subsp. pecten-veneris
413 [Serbia] In agri M. Zlatibor Serb. merid., Jul 856 [1856], l. Pančić.
58
Scirpus radicans Schk [Schkuhr] ≡ S. sylvaticus L. subsp. radicans
(Schkuhr) Rouy
414 No data.
Scleranthus perennis L.
415 [Serbia] In rupestrib. [rupestribus] ad Borač [near Kragujevac]
Serb. Centr. [centralis], Jun 849 [1849], l. Pančić.
Scorzonera montana Mut [Mutel] = S. hispanica L. subsp. hispanica
416 [France] Chambéry [Chambéry, Savoie] in Pedemontio, Jun, com.
Pančić.
Scrophularia hopii Koch [W. D. J. Koch] ≡ S. canina L. subsp. hopii (W.
D. J. Koch) P. Fourn.
417 [Serbia] Arvidis serpentinaceis ad Brđane [Brđani, near Gornji
Milanovac] Serb. Centr. [centralis], Maj, l. Pančić.
Scrophularia laciniata WK [Waldst. & Kit.] ≡ S. heterophylla Willd.
subsp. laciniata (Waldst. & Kit.) Maire & Petitm.
418 [Romania] Ad rupem Solyma Ruskberg [Rusca Montană, Caraş-Severin County] in Banatus [Banat], Jun 844 [1844], l. Pančić.
Scutellaria alpina L.
419 [France] M. Cenis [Mont Cenis, Savoie] in Labandia [?], com.
Pančić.
Scutellaria columnae All.
420 [Serbia] Ad latera sylvar [sylvarum] prope Ražanj Serbia austral.
[australis], Jul., l. Pančić.
Senecio aquaticus Huds. = S. aquaticus Hill subsp. barbareifolius (Wimmer & Grab.) Walters
421 [Serbia] In herbidis M. Jablanik [near Valjevo] Serb. merid.
[meridionalis], Jul 856 [1856], l. Pančić.
Senecio umbrosus WK [Waldst. & Kit.] ≡ S. doria L. subsp. umbrosus
(Waldst. & Kit.) Soó
422 [Serbia?]
“Is it from Srem? It grows on high mountains!” [J. Pančić, in Serbian].
Seseli bienne Crtz. [Crantz] = S. annuum L.
423 No data.
Seseli hippomarathrum L.
424 [Hungary] In collib. [collibus] siccis circa Budam [Buda: western
59
Seseli leucospermum WK [Waldst. & Kit.]
425 [Hungary] Collinis Wolpithal [Wolfsthal?] prope Budam [Buda:
western part of Budapest], Jul 1836, l. Pančić.
Obs. Endemic to western Hungary (Ball 1968).
Seseli rigidum WK [Waldst. & Kit.]
426 [Serbia] rupib. [rupibus] calcareis ad Ravanica [near Ćuprija] Serb.
Central [centralis], Jul, l. Pančić.
Seseli tortuosum L.
427 [Serbia] Collinis arvidis ad Ražanj Serb. austral [australis], Jul, l.
Pančić.
Silene paradoxa L.
428 [Serbia] Asperis serpentinaceis ad Jelakci [near Aleksandrovac]
Serb. merid. [meridionalis], Jul, l. Pančić.
Silene dichotoma Ehrh.
429 [Serbia] In arvis ad vineas Topčider prope Belgrad [Belgrade], Jul,
l. Pančić.
Silene echinata Ehrh. [Otth]
430 No data.
“In Petrović’s label, which has been accidentally lost, it was stated that
this is Silene dichotoma. Is it from Srem?” [J. Pančić, in Serbian].
Silene nemoralis WK [Waldst. & Kit.] ≡ S. italica (L.) Pers. subsp.
nemoralis (Waldst. & Kit.) Nyman
431 [Serbia] declivilate meridionalis M. Rtanj Serb. austral. [australis],
Jul, l. Pančić.
Smyrnium perfoliatum Mill. [L.]
432 [Serbia] In saxosis ad Manasija [near Ćuprija] Serb. Centr.
[centralis], Apr. Jun 847 [1847], l. Pančić.
Sorbus hybrida L.
433 [France] Prope Chambery [Chambéry, Savoie] in Pedemontio, com.
Pančić.
Obs. This north european (Scandinavian) species is not native to France
(Warburg & Kárpáti, 1968)
Stachys palustris L.
434 No data.
Statice tatarica L. ≡ Goniolimon tataricum (L.) Boiss.
435 [Serbia] Rupestrib. [rupestribus] serpentinaceis ad Raška, Serb.
merid. [meridionalis], Jul, l. Pančić.
Obs. Besides this record, Pančić never included this species in his “Flora
principatus Serbiae”, nor “Additamenta” (Pančić 1874, 1884). This
60
species was officially recorded for the first time for Serbian flora in the
vicinity of Preševo and Prokuplje (Ranđelović 1984). More recently, it
has been recorded again near Raška (Krivošej et al. 2008).
Teucrium montanum L. var. supinum Jacq. = T. montanum L. f. supinum
(L.) Lam. & DC.
436 [Serbia] In collib. [collibus] asperi [asperis] ad Kutlovo [Kragujevac] Serb. Centr. [centralis], Jul 850 [1850], l. Pančić.
Torilis helvetica Gmel. [C. C. Gmel.] = T. arvensis (Huds.) Link subsp.
arvensis
437 No data.
Tragopogon floccosus WK. [Waldst. & Kit.] subsp. floccosus
438 [Serbia] Arenosis Bjelobrdo [Deliblato Sand] in Banatus [Banat],
Aug 857 [1857], l. Pančić.
Tragopogon major L. [Jacq.] = T. dubius Scop. [“subsp.” major (Jacq.)
Vollmann]
439 No data.
Trapa natans L.
440 [Serbia] In palude Kriva bara prope Jagodina, Jul 847 [1847], l.
Pančić.
Trifolium angustifolium L.
441 [Serbia] Pratis sterilib. [sterilibus] ad Aleksinac Serb. austral
[australis], Jul, l. Pančić.
Trifolium hirtum All.
442 [Serbia] Collinis siccis ad Dušica [?] Serb. austral [australis], Jul, l.
Pančić.
Trifolium pannonicum L. = T. pannonicum Jacq.
443 [Serbia] Pratis montanis Ozren [near Sokobanja] Serb. aus[t]ralis,
Jul, l. Pančić.
Trifolium medium L.
444 No data.
Note: In the same sheet with next specimen.
Trifolium reclinatum WK. [Waldst. & Kit.] = T. echinatum M. Bieb.
444 [Serbia] Sub vineis ad Pancsova [Pančevo] in Banatus [Banat], Jul
857 [1857], l. Pančić.
“Loco classico kitaibeliano !” [J. Pančić].
Note: In the same sheet with previous specimen.
61
Trifolium resupinatum L.
445 [Serbia] Pratis pinguib. [pinguibus] ad Danubium [Danube] prope
Belgrad [Belgrade], Jul, l. Pančić.
Trifolium speciosum W. [Willd.]
446 [Greece] Attica. [Attikí] In M. Hymetto [Hymettus or Hymettos],
Maj 855 [1855].
Triodia decumbens PB. [P. Beauv.] = Danthonia decumbens (L.) DC.
447 [Serbia] In pratis montanis ad Trešnjevica [near Aranđelovac] Serb.
Centr. [centralis], Jun, l. Pančić.
Trinia [sp.]
448 No data.
“Umbellifera prob. [probabile] Trinia! I found it in splice of Arenaria
serpyllifolia. Pay attention to this herb” [J. Pančić, in serbian].
Vaccinium vitis [-] idaea L.
449 [Serbia] In sylvis supra Majdanpek Serb. bor. [borealis], Jul 853
[1853], l. Pančić.
Valeriana echinata L. ≡ Valerianella echinata (L.) DC.
450 [Greece] Graecia. In M. Pentelico [Penteliko Mountain, in Attikí],
Apr. 855 [1855].
Valerianella carinata Lam. = V. carinata Loisel.
[451] [Serbia] In saxosis ad Rudnik Serb. Centr. [centralis], Maj 848
[1848], l. Pančić.
Verbena supina L.
452 [Greece] Graecia. In M. Parnasso [Mount Parnassus], Jul 855
[1855].
Veronica alpina L. var. serratifolia Roch.
453 [Romania] M. Gugu [Gugu peak, Godeanu mountain in Caraş-Severin County] in Banatus [Banat], Aug 845 [1845], l. Pančić.
Veronica agrestis L.
454 [Serbia] M. Ovčar [near Čačak] Serb. Central [centralis], Apr. 851
[1851], l. Pančić.
Veronica bellidifolia L. [V. bellidioides L.]
455 [Romania] M. Hribina Skenlni [?] ad Banatus [Banat], Aug 845, l.
Pančić.
Veronica buxbaumi [buxbaumii] Ten. = V. persica Poir.
456 No data.
62
Veronica hederifolia L.
457 [Serbia] M. Crni Vr’ [near Jagodina?] Serb. Centr. [centralis], Apr.
850 [1850], l. Pančić.
Vicia cassubica L.
458 [Serbia] In graminosis sylvaticis ad Knić Serb. Centr. [centralis],
Jun 850 [1850], l. Pančić.
Vicia lathyroides L.
459 [Serbia] In glareosis M. Georgis [Đurđevo brdo] ad Jagodina Serb.
Centr. [centralis], Apr. 847 [1847], l. Pančić.
Vicia lutea L.
460 [Serbia] Pratis pinguib. [pinguibus] ad Danubium [Danube] prope
Belgrad [Belgrade], Jul 1847, l. Pančić.
Vicia pannonica Jacq. var. purpurescens Koch = V. pannonica Crantz
subsp. striata (M. Bieb.) Nyman
461 [Romania] In pratis ad Mehadiam [Mehadia, Caraş-Severin County], Maj 845 [1845], l. Pančić.
Vicia pannonica Jacq. = V. pannonica Crantz
462 No data.
Vicia pisiformis L.
463 No data.
Vicia sativa L.
464 [Hungary] In pratis sylvestribus prope Budam [Buda: western part
of Budapest], Maj 840 [1840], l. Pančić.
Vicia serratifolia Jacq. = V. narbonensis L.
465 No data.
“(syn. V. narbonensis β Koch.)” [J. Pančić].
Vinca herbacea WK. [Waldst. & Kit.]
466 [Austria] In lapidosis Bisamberg prope Viennam [Wien], Maj 847
[1847], l. Pančić.
Waldsteinia geoides Willd. var. obtusiloba Panč. [Pančić]
467 [Serbia] Ad cacumen M. Ovčar [near Čačak] Serb. Centr. [centralis], Maj, l. Pančić.
It should be mentioned that an additional 11 sheets with specimens
collected by Pančić are deposited in the Herbarium of the Institute for
Nature Conservation of Vojvodina Province. This herbarium material was
accidentally found, along with a few hundred other sheets, in the
Petrovaradin fortress powder stockroom about 35-40 years ago (V. Stojšić
pers. comm.). According to the characteristics (size, shape and colour) and
label information of the sheets, it is clear that they belong to the Pančić
63
collection in the Gymnasium herbarium. Because they lack the inventory
mark given by Marčetić and Babić who made the first full collection
inventory in 1950 (Marčetić 1950), it is evident that this material was
separated from the main collection before December 1950. These sheets
were probably lost during the many relocations of the whole collection in
the past (at least after the construction of the new Gymnasium building in
1891 or during the World Wars).
Alkanna tinctoria Tausch. [(L.) Tausch]
[Hungary] In arenosis Stadtwäldchen [Városliget] prope Pesth. [Pesthinum: Pest-eastern part of Budapest], Jul 841 [1841], l. Pančić.
Aubrietia graeca Gris. ≡ A. deltoidea (L.) DC. var. graeca (Griseb.) Regel
[Greece] Graecia. In M. Hymetto [Hymettus or Hymettos], Maj 854
[1854].
Avena tenuis Mch. ≡ Ventenata dubia (Leers) Coss.
[Serbia] Collinis herbidis prope Jagodina, Serb. Centr. [centralis], Jun, l.
Pančić.
Note: In the same sheet are also 3 specimens of Achnatherum
calamagrostis (L.) P. Beauv. and one unknown Gramineae (only
fragment of spike), all without data.
Crocus iridiflorus Heuf. = C. banaticus J. Gay
[Romania] Ad latera sylvar. [sylvarum] Ruszkberg [Rusca Montană,
Caraş-Severin County] in Banatus [Banat], Sept. 844 [1844], l. Pančić.
Cynosurus echinatus L.
[Serbia] Herbidi supra Borač [near Kragujevac], Serb. Centr. [centralis],
Maj, l. Pančić.
Elymus crinitus Schreb [Schreb.] = Taeniatherum caput-medusae (L.)
Nevski
[Serbia] [...] inter Banja [Sokobanja near Aleksinac] et M. Rtanj, Serb.
Austr. [australis], Jul, l. Pančić.
Leontopodium alpinum DC. [Cass.]
[Austria?] In alpe Rax Austria [?], Jul 845 [1845], l. Pančić.
Orchis papilionacea L.
[Serbia] In graminosis supra Topčider prope Belgrad [Belgrade], Maj
855 [1855], l. Pančić.
Paronychia capitata Lam. – P. kapela (Hacq.) A. Kerner subsp. kapela
[Serbia] M. Stol, Serb. Austral. [australis], Jun 850 [1850], l. Pančić.
Saponaria lutea L.
[France] M. Cenis [Mont Cenis, Savoie] in Pedemon. [Pedemontio], Jun
Jul, com Pančić.
64
Obs. Endemic to Alps (Chater 1993).
Trigonella monspeliaca L.
[Serbia] In arvi[s] ad Belgrad [Belgrade], Jul 857 [1857], l. Pančić.
The herbarium collection of Josif Pančić at the Gymnasium of Sremski
Karlovci, with additional sheets in the Institute for Nature Conservation of
Vojvodina Province comprises 281 sheets placed in 3 boxes (two in the
Gymnasium and one in the Institute). Within this material there are 295
taxa on species (278) and subspecies level (30) classified into 169 genera
Fig. 1 Herbarium specimen of Minuartia stellata (E. D. Clarke)
Maire & Petitmengin, Mt Parnassos (Greece).
and 49 families. Almost one third of the herbarium specimens in this
collection lacks information about the date and the locality of collecting (76
sheets). Only 126 sheets include information on the year of collecting the
plant material.
Pančić’s collection from Sremski Karlovci contains herbarium specimens of plants collected in the territories of Serbia (129), Romania (27),
Hungary (12) and Austria (6). In addition, this collection includes the herbarium material acquired through exchange from France (12), Greece (11)
65
and Italy (7). It is interesting to note that in this collection there are more
specimens from France and Greece than in the Herbarium Pancicianum of
the Institute of Botany and the Botanical Garden “Jevremovac”, University
of Belgrade, that contains only 9 exsiccates from France and 7 from Greece
(Vukojičić et al. 2011). However, a significant number (76) of sheets lack
information about the locality. The most important part of this collection
are herbarium specimens of 15 endemic species (Achillea umbellata,
Asperula capitata, Eryngium serbicum, Eryngium palmatum, Evax perpusilla, Gagea graeca, Leontodon hirtus, Malcolmia graeca, Minuartia stellata (Fig. 1), Pedicularis ascendens, Pedicularis cenisia, Primula pedemontana, Saponaria lutea, Saxifraga biflora and Seseli leucospermum). In
addition, there are specimens of Eryngium serbicum (Fig. 2) and E.
palmatum originating from the locus classicus, and probably representing
type material of these two species. The collection also contains exsiccates
of Achillea ptarmica, species that disappeared from the territory of Serbia
(Stevanović & Vukojičić 1999: 98), Hippuris vulgaris, critically endangered species in Serbia (Vučković & Panjković 1999), as well as Fritillaria
montana from Mt Medvednik and Lysimachia nemorum from Mt Murtenica, likely to be a new localites for these species in Serbia.
Fig. 2. - Herbarium specimen of Eryngium serbicum Pančić, Dobrača (Serbia).
Collection analysis showed that Pančić’s specimens in Sremski Karlovci dated from the very beginning of his botanical work. This period
includes three subsequent periods: student days at University of Pest (1832-1843), “Ruskberg period” (1844-1845), and early years in Serbia (1846-1858). The oldest material originated from Hungary and dates from 1836 to
66
1839 (Asperula arvensis, Cirsium pannonicum, Juncus subnodulosus, Ranunculus pedatus, Phyteuma orbiculare, Seseli leucospermum). During the
“Ruskberg period”, Pančić explored flora of the Carpathian Mountains
(especially the Transylvanian part) and the Carinthian Alps. In this collection there are 27 specimens from this period (Fig. 3). The third period
Fig. 3. - Herbarium specimen of A. capitata Kit. ex Schult.,
Domogled (Romania).
coincides with Pančić’s practice of medicine in Jagodina and Kragujevac
(1846-1853) and first years in Belgrade Lyceum (from 1853 onwards).
Although the majority of Pančić’s specimens are deposited in the
Herbarium of the Institute of Botany and the Botanical Garden “Jevremovac”, also of great significance is the minor part of his specimens in the
67
special collection Herbarium Pancicianum. Besides its historical and
cultural value, this herbarium material is of great importance for present
and future botanical explorations especially throughout the Balkan Peninsula and needs to be fully digitized which would prevent damage to the
material after its use and properly presented to a wider audience.
Acknowledgements
We wish to thank Dr. Barina Zoltán from the Hungarian Natural
History Museum in Budapest for providing invaluable literature sources, Dr
Marjan Niketić from Natural History Museum in Belgrade for revision of
critical taxa and the staff of the Gymnasium of Sremski Karlovci.
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71
ПАНЧИЋЕВИ ПРИМЕРЦИ У ХЕРБАРИЈУМСКОЈ ЗБИРЦИ
АНДРЕАСА РАФАЕЛА ВОЛНИЈА
РАНКО ПЕРИЋ, КСЕНИЈА ЈАКОВЉЕВИЋ, ВИДА СТОЈШИЋ,
СНЕЖАНА ВУКОЈИЧИЋ
РЕЗИМЕ
Хербаријумска збирка Андреаса Рафаела Волнија у Гимназији у
Сремским Карловцима садржи и примерке чувеног српског ботаничара др Јосифа Панчића, које је он поклонио Гимназији. Такође се један
мали део ових примерака налази и у хербаријуму Покрајинског завода
за заштиту природе у Новом Саду. По први пут је ботаничкој јавности
представљен списак са 281 хербарским табаком и 295 таксона на
нивоу врсте (278) и подврсте (30) са подацима о месту и времену
сакупљања, коментарима о врстама и ревизијама.

PANČIĆ`S SPECIMENS IN THE HERBARIUM COLLECTION OF

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