H. longipes var. longipes

Register of Hosta Species and their Nomenclature • Part 2
For complete information refer to Species Update on the home page of Hostalibrary.org
© 2008 • The American Hosta Society/W.G. Schmid
H. ibukiensis Araki.
Acta Phytotaxonomica et Geobotanica, Vol. 11, 325
1942. イブキギボウシ = 伊吹宝珠 = Ibuki Gibōshi
(Mount Ibuki Hosta). Now considered a named
phenotypical variant of H. sieboldii as H. sieboldii
‘Ibukiensis’; in horticulture as the culton/cultivar H.
‘Ibukiensis’ (Schmid 2007)
AHS name: H. ‘Ibukiensis’
H. ‘Ibukiensis’ (in situ 1988) ►
Note Keeled Fertile Bracts
Ibukiyama; 伊吹山), Shiga-ken; 滋賀県. and Gifu
Prefecture (Gifu-ken; 岐阜). Hab.: Wet montane
meadows and forest margins. Kansai-chihō
(関西地方) central Honshū (本州).
H. jonesii M.G. Chung 1989
Annals of the Missouri Botanical Garden,
76-3:920–922. 1989
다도해비비추 = Da-do-hae-bi-bi-chu
(= Many Islands Hosta)
ジョーンズイギボウシ = Jonesii Gibōshi
AHS name: H. jonesii
H. jonesii (in situ) 1988 ►
UGA/Chung 1403 Voucher
Jin-do Island (진도) at 60 m (197 ft) AMSL
Hab.: In rocky and humus soil on southfacing slopes at 250 feet (76 m) AMSL, in
shady pine-oak forests near the ocean on
island along the southern coast of Chŏllanam-to, Korea: Jin-do (진도), Namhae-do
(남해도), and other islands. Chung (2005)
stated that some of the populations from
which type specimens were collected are
extinct due to human activity.
1
H. jonesii (cultivated) Branched raceme (left) and bell-shaped flowers (right)
UGA/Chung Voucher 1075 from Namhae Island (Namhae-do; 남해도)
Hosta Hill R.G. ● Photo W.G. Schmid July 1989 (left); August 1998 (right)
H. kikutii var. kikutii
Maekawa
J. Japanese Botany, 13:48, ic. f. 9 1937;
and J. of the Faculty of Science, Imp.
Uni. of Tokyo, Sect. 3 Bot., Vol. 5:375,
ic. 3, 4 p. 324 1940. ヒュウガギボウシ
= Hyūga Gibōshi (Hyūga Prov. Hosta)
AHS name: H. kikutii var. kikutii
AHS (hort.) name: H. kikutii
(with or without autonym).
H. kikutii (in situ)
Growing epiphytically on tree trunk.
Sendaigawa headwaters (川内川源流)
Kyūshū (九州). Court.: Hananomura
Hab.: Growing in-ground, lithophytically or epiphytically on wet rocks
or tree trunks in high rainfall areas.
Found in old Hyūga Province (日向国;
Hyūga-no kuni), hence the name
Hyūga Gibōshi. Largest populations on
Kyūshū (九州) and Shikoku (四国).
2
H. kikutii (cult.)
Raceme with large bracts ▲
Hosta Hill R.G. W.G. Schmid
Late August 1989
H. kikutii (cult.)
◄ Growing in sprays of waterfall
(river unknown)
H. kikutii var. kikutii f. leuconota W.G. Schmid
The Genus Hosta - Gibōshi Zoku (ギボウシ属); 61, 322. 1991.
ウラギロヒョウガギボウシ = Urajiro Hyūga Gibōshi (White-backed Hyūga Hosta)
細葉ウラギロヒョウガギボウシ = Hosoba Urajiro Hyūga Gibōshi (Narrow-leaf
white-backed Hyūga Hosta)
AHS name: H. kikutii var. kikutii
f. leuconota
AHS name: H. kikutii leuconota
(abbr. hort. name).
H. kikutii var. kikutii
f. leuconota (cult. ex HH/WGS)
Court.: HL/Christensen
Hab.: River valleys, wet exposed
rocky banks and moist forest
margins; Voucher, coll. 1970,
HH/SU No. 406; loc.cit.: Kagoshima-ken (鹿児島市) in Kyūshū
(九州) and Shikoku (四国).
3
H. kikutii var. polyneuron (Maekawa) Fujita.
Acta Phytotaxonomica et Geobotanica, Vol. 27, 3/4:78 1976.
スダレギボウシ = 簾擬宝珠 = Sudare Gibōshi (= rattan blind Hosta)
AHS name: H. kikutii
var. polyneuron
AHS name: H. kikutii
polyneuron (abbr. hort.
name).
H. kikutii var.
Polyneuron (in situ).
Loc. cit.: Mt. Kanpū
(寒風山); Court.:
Forest 萩生 2006
Hab.: Wet, exposed
rock faces and moist
forest margins; Kōchiken (高知市); Shikoku
(四国), Kyūshū (九州)
and Kansai region
(関西地方)
H. kikutii var. caput-avis
Maekawa
ウナズキギボウシ = 頷き擬宝珠 = Unazuki Gibōshi (bowing head Hosta)
H. kikutii var. tosana Maekawa
トサノギボウシ = 土佐の擬宝珠 = Tosano Gibōshi ([Province] Tosa Hosta)
Encyclopedia of Horticulture, 2:633 1950.
Combined in synonymy by Fujita (Acta Phytotaxonomica et Geobotanica, Vol. 27,
3/4:78;1976).
Both names are listed
as they represent locally
differentiated
populations with
distinct differences,
including disparities in
scape posture. In
montane loci the scapes
bend at ground level
and hang down; in flat
land areas the scape
bends further up (see
photos page 4 ►and
5▼).
4
AHS name: H. kikutii
var. caput-avis
AHS name: H. kikutii
caput-avis (abbr. hort.
name).
AND/OR:
AHS name: H. kikutii
var. tosana
AHS name: H. kikutii
tosana (abbr. hort.
name).
Photograph Page 4 ▲: H. kikutii var. caput-avis. Court.: Hototogi2 moto; Loc. cit.:
Mt. Kenzan (剣山); Tokushima-ken (徳島県); Shikoku (四国).
Photograph Page 5 ▲: H. kikutii var. tosana. Loc. cit.: Mt. Kajigamori (梶ヶ森山),
Kōchi-ken (高知県); Shikoku (四国) (ex HH/HM W.G. Schmid 1988). Hab.: wet,
exposed rock faces and moist forest margins; Kōchi-ken (高知市); Shikoku (四国)
and Kyūshū (九州).
H. kikutii var.
yakusimensis
(Masamune) Maekawa
Journal of the Faculty
of Science, Imperial
University of Tokyo,
Section 3 Botany, Vol.
5:374, 375, 376, ic. 42.
43, 44 1940.
ヒメヒョウガギボウシ
= Hime Hyūga Gibōshi
(small Hyūga Hosta)
AHS name: H. kikutii var. yakusimensis
AHS name: H. kikutii yakusimensis (abbr. hort.
name).
H. kikutii var. yakusimensis (cult. ex HH/WGS)▲►
Raceme and leaf mound, W.G. Schmid 1987 loc. cit.:
Mt. Yokogura (横倉山), Kōchi-ken (高知市);
Shikoku (四国). Hab.: Lower montane forest
margins, rock outcrops lithophytically or
epiphytically. Shikoku (四国) and Kyūshū (九州).
5
H. kiyosumiensis Maekawa
Jornal of Japanese Botany 11:689; ic. f. 15 1935
キヨスミギボウシ = 清澄擬宝珠 = Kiyosumi Gibōshi (Kiyosumi [mountain] Hosta)
H. kiyosumiensis var. petrophila Maekawa
Divisiones et plantae novae generis Hostae (2). J. Japanese Botany, 14:1:45–49. 1938.
イワマギボウシ = 岩間擬宝珠 = Iwama Gibōshi (liking [to be among] rocks Hosta)
Both names are listed,
but the differentiation
among populations is
slight and not distinct
enough to justify
separate rankings. Both
variants are now
considered synonymous
and use of the autonym
is no longer required.
AHS name:
H. kiyosumimensis
H. kiyosumimensis (►in
situ 1984) Oto River
(男川) Okazaki-shi (岡崎市), Aichi-ken (愛知県). Hab.: Along moist rocky river
banks, ravines, and forest margins at lower elevations and moist submontane
outcrops to 1000 m (2280 feet) in Kantō-chihō (関東地方), Kansai-chihō ( 関西地方)
and Chūbu-chihō (中部地方) in central Honshū (本州), and eastern Shikoku(四国)
in Awa-no kuni (阿波国).
H. laevigata W.G. Schmid
The Genus Hosta Giboshi Zoku, 64-65, 316. 1991
ラエウイガタギボウシ =
Laevigata Gibōshi (polished
Hosta).
Originated from seed
collected by B.A. Yinger, on
Taehuksan (Tae-huksan-do
(대흑산도 [企比至亀]) off
the southwestern coast of
Korea opposite the province
of Chollanam-do (verified
Yinger/Chung 1987/1988).
Maybe a natural hybrid with
low fertility. (see photos page
6 ►and 7▼).
6
◄ H. laevigata (right)
H. yingeri (left)
Spider-flowered tepals
▼ Perianth Grouping and
typical leaf mounds
(Cult. ex Yinger)
at Hosta Hill R.G.
W.G. Schmid 1989
AHS name: H. laevigata
AHS name: H. laevigata or H. ‘Laevigata’ (hort.)
Hab.: In shade near coast on rocky slopes on Taehuksan Island (대흑산도), Chollanam-do,; island
along the southwestern coast of Korea, province
Chollanam-do, southern Huksan-chedo.
The H. longipes Complex = 岩ギボウシ (The Rock Hostas)
H. longipes (岩ギボウシ) is considered one of the most variable (polymorphic)
groups of closely related Hosta populations. This is due to the wide-ranging habitat
that extends from the Kantō region (関東地方, Kantō-chihō) in the North to Kyūshū
region (九州地方, Kyūshū-chihō) to the South. Kantō is the area surrounding Tokyo
and Kyushu is the southernmost island where the large city of Nagasaki is located.
Nagasaki-shi (長崎市). This habitat spans almost one-half of the entire Japanese
Archipelago and areas of different climates and ecologies that caused different
forms to evolve. They are called rock hostas, because they are found mostly in the
mountains of this part of Japan where they have adapted to grow on rock outcrops
along the streams, splashed with water, while Others have evolved to inhabit sheer
rock faces in the mountains. Still other H. longipes variants cover huge boulders
with a contiguous layer of leafy hosta species. The multiplicity of phenotypes and
7
genotypes has evolved due to environmental conditions and relative isolation created
by tall mountain chains. Almost every geographic district has H. longipes
populations that are differentiated from others. Only taxonomically identified taxa
are included in this brief overview.
H. longipes var. longipes
(Franchet et Savatier) Matsumura
List Plant. Nikko Bot. G., 21. 1894
H. longipes var. vulgata
W.G. Schmid et G.S. Daniels
The genus Hosta – Gibōshi Zoku, pp.
71-73; 1991
イワギボウシ = 岩ギボウシ =
岩擬宝珠 = Iwa Gibōshi (Rock Hosta)
비비추 = Bi-bi-chu (Korean)
Including:
H. longipes f. alba
(Nakai) T.B.Lee 1980
シロバナイワギボウシ =
白花岩ギボウシ = Shirobana Iwa
Gibōshi (White-flowered Rock Hosta)
흰비비추 = Huin-bi-bi-chu (Korean)
H. longipes var. vulgata (in situ) ►
Loc. cit.: Chichibu-gun (秩父郡); in
Kantō-chihō (関東地方). Court.:
Rojiura-no-Giboshi 路地裏のギボシ
◄ H. longipes
in situ Tenryugawa (天竜川);
Shizuoka-ken
(静岡県). Streamside groups
showing affinity to
water and rock.
Court.: H. Sugita
HH/W.G. Schmid
1984. Hab.: Moist
rocks and cliffs
from Kantō (関東
地方) south to
Kyūshū region
(九州地方).
8
AHS name: H. longipes var.
longipes
AHS name: H. longipes (abbr.
hort. name).
AHS name: H. longipes var.
longipes var. vulgata (alt. bot.
name)
H. longipes var. caduca
Fujita: Acta Phytotax. et
Geobot., Vol. 27, 3/4:83, ic. 6
1976. サイコクイワギボウシ
= 西国岩擬宝珠 = Saikoku
Iwa Gibōshi (Rock hosta from
Saikoku) ウスバイワギボウシ
= ウスバ薄刃岩擬宝珠 =
▲ H. longipes var. caduca (in situ) ▲
Usuba Iwa Gibōshi (Slender
Mt. Kamegamori (瓶ヶ森) Ehime-ken (愛媛県).
leaf rock hosta) Hab.:
Court.: foresthagyu ▪ oka 新居浜市 愛媛県
Mountain-dwelling species,
growing mostly in higher elevations on moist rock out-crops and along ridge lines.
Population size is usually very small. Rarely seen in lower habitat along the river
valleys. H. longipes var. caduca is found on Shikoku in the Ishizuki and Akaishi
mountain ranges of Kochi Prefecture (Kochi-ken 高知県) and Ehime-ken (愛媛県);
also in Kyūshū in the Kyūshū Sanchi (九州山地) Mountain Range in following
prefectures: Miyazaki-ken (宮崎県), Fukuoka-ken (福岡県), Ōita-ken (大分県),
Kumamoto-ken (熊本県), and Kagoshima-ken (鹿児島県). Reportedly in
Hiroshima-ken (広島県).
H. longipes var.
longipes f. sparsa
(Nakai) W.G. Schmid 1991
The genus Hosta – Gibōshi
Zoku, pp. 67-68, 323; 1991
アキギボウシ = 秋擬宝珠
= Aki Gibōshi (fallblooming Hosta).
In syn: H. longipes var.
lancea Honda: Botanical
Magazine, Tokyo, 49:696
1935.
ホソバイワギボウシ=
細葉岩擬宝珠 = Hosoba
Iwa Gibōshi (Narrowleaved rock hosta)
▲H. longipes var. longipes f. sparsa (in situ) at Tenryu
River (天竜川 = Tenryugawa); Shizuoka-ken (静岡県)
9
This taxon is NOT H. ‘Tardiflora’.
Maekawa (1969) dropped the local native
name 秋擬宝珠 (= H. sparsa) and adopted
H. Tardiflora'. He also transferred the
Japanese name Aki Giboshi to H.
'Tardiflora'. Being incorrect, this transfer
is the reason why Japanese botanists have
mixed up the names H. ‘Tardiflora’, Aki
Giboshi (= H. longipes f. sparsa) and the local
name for Aki Giboshi, i.e., Hosoba Iwa
Giboshi (細葉 岩擬宝珠 = H. longipes var.
lancea). Schmid (1991. 2006) pointed out
that H. ‘Tardiflora’ has yellow anthers and
H. longipes f. sparsa has purple anthers
(◄◄ see photo to the left)
AHS name: H. longipes var. longipes
f. sparsa
AHS name: H. longipes sparsa
(abbr. hort. name).
H. longipes f. sparsa (in situ) ►
Tenryu River Area (天竜川 =
Tenryugawa) Shizuoka-ken
(静岡県) H. Sugita 1986
Hab.: Wet rocks on river banks
(Tenryugawa (天竜川) and
Otogawa (男川) in Shizuoka-ken
(静岡県) and Aichi-ken (愛知県)
and in mountain valleys in the
southern Chūbu Region
(中部地方).
H. longipes var. longipes f.
hypoglauca
J. of the Faculty of Science, Section 3
Botany, Vol. 5:388 1940
コフキイワギボウ = Kofuki Iwa
Giboshi (glaucous on the back
Hosta)
H. longipes hypoglauca (in situ) ►
Saitama-ken ((埼玉県;) Rojiura
路地裏のギボウシこと 後ノ上
憲一
10
AHS name: H. longipes var.
longipes f. hypoglauca
AHS name: H. longipes hypoglauca
(abbr. hort. name).
H. longipes var. longipes f.
hypoglauca ►
(in situ) Chichibu Gun (秩父郡)
Tochigi-ken (栃木県)
Rojiura-no-Giboshi (MDS 351
Type)
Hab.: Rocky forest areas in Nikko (
日光) and Kanagawa-ken (神奈川
県), Hakone-chō (箱根町), near
Mount Hakone, Kanagawa
Prefecture Kanto and Chubu (中部
地方).
H. longipes var. latifolia Maekawa
J. of the Faculty of Science, Imp. Uni. of Tokyo, Sect. 3 Bot., Vol. 5:388; 1940.
アマギイワギボウシ = 天城豆岩ギボウシ = Amagi Iwa Gibōshi (Amagi rock
hosta)
イズイワギボウシ= 伊豆岩ギボウシ = Izu Iwa Gibōshi (Izu rock hosta)
マルバイワギボウシ= 丸葉岩ギボウシ= 丸葉岩擬宝珠 = Maruba Iwa Gibōshi
(Round-leaf rock hosta)
AHS name: H. longipes var. longipes
f. latifolia
AHS name: H. longipes latifolia
(abbr. hort. name).
H. longipes var. latifolia ►
(in situ) Amagi-san (天城山), Izuhantō (伊豆半島).
Hab.: Wet rocks on river banks and
steep ravines in Izu Peninsula
伊豆半島; Izu-hantō); Shizuokaken (静岡県); on Izu Oshima Island
(伊豆大島); southern Chūbu
(中部地方); south-central Honshū
(本州).
11
H. longipes var. longipes f. viridipes Maekawa
J. of the Faculty of Science, Imp. Uni. of Tokyo, Sect. 3 Bot., Vol. 5:388 1940.
アオジクイワギボウシ = 緑色葉柄岩擬宝珠 = Aoziku Iwa Gibōshi (Green
leafstalk rock hosta)
AHS name: H. longipes var.
longipes f. viridipes
AHS name: H. longipes
viridipes (abbr. hort. name).
AHS name: H. longipes
‘Viridipes’ (cultivar name).
H. longipes f. viridipes (Loc.
cit.: Unspecified) ►
This taxon is here considered a
phenotypical variant of H.
longipes var. longipes and its
salient morphological traits are
identical those of the typical
species. Due to its presence in
horticulture, forma rank is maintained and its name may be written as H. longipes f.
viridipes or as a cultivar as H. longipes ‘Viridipes’.
H. longissima Honda
H. longissima var. brevifolia
Maekawa
J. Japanese Botany, 13:894 1937 and
Maekawa J. of the Faculty of Science,
Imperial Uni. Tokyo, Section 3 Botany,
Vol. 5:404 1940
ミズキボウシ = 水擬宝珠 = Mizu
Gibōshi (Swamp hosta)
サジキボウシ = 匙擬宝珠 = Saji
Gibōshi (Spoon-leaved hosta)
AHS name: H. longissima var. brevifolia
AHS name: H. longipes brevifolia (abbr.
hort. name).
H. longissima var. brevifolia (in situ) ►
Wetland Meadow near Nukata-chō
(額田町); Aichi-ken (愛知県). Court.: S.
Aoki. Hab.: Wet meadows and swampy
areas in parts of Honshū (本州),
Shikoku (四国) Kyūshū (九州).
12
H. longissima Honda
J. Japanese Botany, 13:894 1937 and Maekawa J. of the Faculty of Science, Imperial
University Tokyo, Section 3 Botany, Vol. 5:404 1940
H. longissima var. longifolia (Honda) W.G. Schmid
The genus Hosta: Gibōshi Zoku (ギボウシ属). 1991; pp. 72-74, 328, 339
ミズキボウシ = 水擬宝珠 = Mizu Gibōshi (Swamp hosta)
ナガバミズキボウシ = 長い葉身水擬宝珠 = Nagaba Mizu Gibōshi (Narrow-leaved
swamp hosta)
ホソバミズキボウシ = 細葉水擬宝珠 = Hosoba Mizu Gibōshi (Slender-leaved
swamp hosta)
AHS name: H. longissima var.
longifolia
AHS name: H. longipes longifolia
(abbr. hort. name).
H. longissima var. longifolia ►
(in situ)
Aichi-ken (愛知県) by Okayama
University of Science; Y. Hada
岡山理科大学 植物生態学.
Hab.: Wet meadows and swampy
areas in parts of Honshū (本州),
Shikoku (四国) Kyūshū (九州).
H. minor (Baker) Nakai
J. of the College of Science, Imperial
University Tokyo, 31:251; 1911
H. minor f. alba (Nakai) Maekawa
J. of the Faculty of Science, Imp. Uni.
Tokyo, Sect. 3 Bot., Vol. 5:418; 1940
좀비비추 = Jom-bi-bi-chu (Korean)
ケイリンギボウシ = Keirin Gibōshi
흰좀비비추 = Huin-jom-bi-bi-chu (Kor.)
シロバナケイリンギボウシ = Shirobana
Keirin Gibōshi
AHS name: H. minor
AHS name: H. minor ‘Alba’ (hort.)
H. minor (in situ) M.G. Chung ►
Koje Island, Kyongsangnam-do; 1988.
Hab.: Southern and south-eastern coast
provinces and coastal islands in Korea.
13
H. montana Maekawa
Journal of the Faculty of Science, Imp. Uni. of Tokyo, Sect. 3 Bot., Vol. 5:356–357, 361
ic. 21, 23 1940
オオバギボウシ = 大葉擬宝珠 = Ōba Gibōshi (Large-leaved hosta)
The following phenotypical variants of H. montana are now ranked as cultivar forms:
H. montana ‘Liliiflora’
H. montana ‘Praeflorens’ (used as an incorrect name for H. montana f. macrophylla).
H. montana ‘Transiens’
H. montana ‘Ovatolancifolia’
AHS name: H. montana (note the above
former taxa now considered cultivars
forms.
H. montana (in situ: Mt. Ibuki) Shigaken
(滋賀県); on the flanks of Mount Ibuki
(伊吹山). Court.: Fujiwa-yama. Hab.:
Low and high montane sites; large
populations spread on mountain tops, in
forests and higher montane meadows.
Common in Omote-Nihon (表日本),
Tohoku, Kanto, Chubu and Kansai regions, eastern Honshu; sporadically in other
areas of Japan. Cultivated as food in Honshū (本州) and a weed in rice-growing
areas.
H. montana in full bloom on Mt. Ibuki (Court.: nagozeki yama)
14
H. montana f. macrophylla W.G. Schmid
The Genus Hosta - Gibōshi Zoku (ギボウシ
属); 77, 317. 1991
キョダイバギボウシ = 巨大葉擬宝珠 =
Kyodaiba Gibōshi (Huge-leaved hosta)
AHS name: H. montana f. macrophylla
AHS name: H. montana macrophylla (abbr.
hort. name).
H. montana f. macrophylla (cult.) ►
Ex hort. Hosta Hill R.G. KR No. B–3 coll.
OSA, 1951 in Nara; Hab.: Low and high
montane sites; large populations spread on
mountain tops, in forests and higher
montane meadows in Nara-ken (奈良県);
Kyoto-fu (京都府) in west-central Honshū
(本州).
H. montana f. macrophylla (cult.)▼
Hosta Hill R.G. May 1988 B-3 ex coll.
W.G. Schmid
15