Digital Imaging of Biological Type Digital Imaging of Biological Type

Transcription

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18208F' */23)/F9' 82<' 8).' 3+.).@/).' 02483,4@8*3/);' @/)' 4*822,27' .-.2'
!
5[
!"#$%"$&'()*+,-.'!/*01.234'
1/<.)83.' 4,_.' 8)*+,-.49' :.*804.' 3+.;' ).I0,).' -,)308FF;' */23,20/04'
18208F' 40=.)-,4,/29' 82<' +8-.' -.);' F,1,3.<' 3+)/07+=03' )83.4H' S,7+'
-/F01.'=)/@.44,/28F'@F83:.<'4*822.)4'J,3+':0,F3B,2'</*01.23'@..<.)4'
8).' 8-8,F8:F.' @/)' =)/@.44,/28F' <,7,38F' 8)*+,-,27' 82<' </*01.23'
=)/*.44,27' 8==F,*83,/249' 82<' 3+.4.' 18;' 8F4/' :.' 1/).' 40,38:F.' @/)'
4*,.23,@,*'8)*+,-.'8*I0,4,3,/29'8F:.,3'83'8'*/43'V^BV^^'3,1.4'1/).'3+82'
:84,*' LC' 4*822.)4H' S/J.-.)9' =)/@.44,/28F' @F83:.<' 4*822.)4' 8).'
<.4,72.<' 3/' =)/*.44' 4382<8)<' */24,43.23' 1/<.)2' </*01.23' @/)1834'
K40*+'84').814'/@'(Y'=8=.)N9'82<'8).'/@3.2'028:F.'3/'+82<F.'8)*+,-.'
</*01.234' .@@,*,.23F;H' Z;' */1=8),4/2' J,3+' 1/<.)2' =8=.)9' 8)*+,-.'
</*01.234' 04.' 8' 10*+' J,<.)' -8),.3;' /@' =8=.)' 82<' *8)<' 4,_.49' 82<'
.A+,:,3' ,2*/24,43.2*;' 82<' -8);,27' I08F,3;' 82<' =8=.)' 3.A30).9' 84' J.FF'
84'@)87,F,3;9'J+,*+'18E.'.@@,*,.23'=8=.)'+82<F,27'8'<,@@,*0F3'=)/:F.1'3/'
/-.)*/1.H''
'
-#."(#+'(*/&01&()*#++&,)(
'
P243.8<' /@' 3)824=/)3,27' 8' F,2.8)' CC!' 8*)/44' 3+.' </*01.23' 3/'
=)/<0*.'8'4*822.<',187.9'3+.'</*01.23'*82':.'3)824=/)3.<'=843'8'
'
'
`,7H'VH'C+.I0.'4*822.)'K*/0)3.4;'UQ(CBZ($CSQNH'O='3/'5^'*+.I0.4'K/)'
,2<.A'*8)<4N'8).'F/8<.<',23/'3+.'@/23'+/==.)9'82<'8).'3+.2'@.<'/2.B:;B/2.'
=843'3J,2'-.)3,*8FF;'1/023.<'CC!4'K4/':/3+'4,<.4'/@'3+.'</*01.23'8).'
4*822.<'4,10F382./04F;NH'('04.)B4=.*,@,.<'3.A3'43),27'*82'8F4/':.'=),23.<'/2'
3+.'</*01.23'<0),27'4*822,27H'
!
a^'
!"#$%"$&'()*+,-.'!/*01.234'
@,A.<' F,2.8)' CC!H' %+,4' ,4' 3+.' =),2*,=F.' 04.<' ,2' 18,F' 82<' *+.I0.'
4*822.)49'J+.).'438*E4'/@'18,F'/)'*+.I0.4'8).'@.<'/2.B:;B/2.'=843'
3+.' CC!9' +.F<' :;' 1/-,27' :.F34' .,3+.)' 4,<.' /@' 3+.' </*01.23H' b8)7.'
4*8F.'4;43.14'04.<'@/)'=/438F'4/)3,27'3;=,*8FF;'4*82'82<'=)/*.44'V^'
F.33.)4'=.)'4.*/2<'@)/1'+/==.)4'@,FF.<'J,3+'3+/0482<4'/@'18,F=,.*.49'
J+,F.' 418FF' *+.I0.' 4*822.)4' K@,7H' VN' /@' 3+.' 3;=.' @/02<' 83' :82E' 3,FF4'
*82'4*82'8:/03'V'*+.I0.'=.)'4.*/2<'@)/1'8'+/==.)'+/F<,27'8:/03'5^'
*+.I0.4H'%+.'F8)7.'+,7+B4=..<'4;43.14'8).':0,F3'3/'/)<.)9'82<'3+04'
8).').F83,-.F;'.A=.24,-.9':03'418FF'*+.I0.'4*822.)4'8).'8-8,F8:F.'83'
1/).' 1/<.43' */43' K8)/02<' c]^^^N9' 82<' 8).' I0,3.' J.FFB40,3.<' 3/'
4*822,27' 8)*+,-.' *8)<' ,2<.A.4' /@' 3+.' 3;=.' 3;=,*8FF;' @/02<' ,2' 182;'
104.014'82<'F,:)8),.4'K4..'U.*3,/2'5NH''
'
2#+'3/&"'()*#++&,)(
'
(2/3+.)' =/44,:F.' 4*822.)' */2@,70)83,/2' ,4' 3+.' +82<B+.F<' 4*822.)H'
%;=,*8F' */11.)*,8F' ,1=F.1.2383,/24' 8).' 1/<.FF.<' /2' =.249' J,3+'
-8);,27' J,<3+' 4*822,27' +.8<4H' %+.;' 8).' 7.2.)8FF;' <.4,72.<' @/)'
4.F.*3,-.' 4*822,27' /@' </*01.23' */1=/2.2349'82<' /@3.2' ,23.2<.<' 3/'
4*82'/2.'/)'8'@.J'F,2.4'/@'3.A3'83'8'3,1.9'3+/07+'4/1.'1/<.F4'+8-.'
3+.' *8=8:,F,3;' 3/' 43,3*+' 3/7.3+.)' 8' 7)/0=' /@' /-.)F8==,27' )843.)' 4*824'
,23/'8'4,27F.'</*01.23'=87.H'U/1.'1/<.F4'8).'<.4,72.<'@/)'/=.)8B
3,/2' ,2' */2R02*3,/2' J,3+' 8' +/43' */1=03.)' 82<' ).*/)<' )843.)' ,187.4'
,23.)28FF;' 023,F' </J2F/8<.<' 3/' 3+.' */1=03.)' -,8' 82' 844/*,83.<' =.2B
+/F<.)d'/3+.)4'8).',23.2<.<'84',2<.=.2<.23'=.),=+.)8F49'82<',2*F0<.'
4/1.' "CD' *8=8:,F,3;' J,3+,2' 3+.' =.2H' !0.' 3/' 3+.,)' F,1,3.<' ,2=03' <838'
)83.'82<'=)/*.44,27'*8=8:,F,3,.49'40*+'<.-,*.4'8).'/@'F,1,3.<'-8F0.',2'
8)*+,-.' </*01.23' 8*I0,4,3,/29' 3+/07+' 3+.;' */0F<' :.' 04.<' @/)'
8*I0,),27' 418FF' ,187.4' @)/1' ,)).70F8)' </*01.23' @)871.234' 40*+' 84'
4=.*,1.2'F8:.F4H''
'
4556()*#++&,)(
'
%+.' 3.*+2/F/7;' <.4*),:.<' 4/' @8)' ,4' /=3,1,4.<' @/)' 4*822,27' 4,27F.'
</*01.23' =87.4' )83+.)' 3+82' :/02<' </*01.234' 40*+' 84' ://E4' /)'
F.<7.)4H' L.)+8=4' 3+.' 1/43' 40,38:F.' <.-,*.' @/)' 4*822,27' 8)*+,-.'
://E4' ,4' 3+.' e.)/A' L(DC' ://E4*822.)' K@,7H']NH' %+,4' 4*822.)' +84'
:..2' <.4,72.<' 4=.*,@,*8FF;' 3/' +82<F.' )8).' 82<' @)87,F.' ://E4' J,3+/03'
<8187,27' 3+.19' :;' 04,27' 8' 3J/B=82.<' 7F844' J.<7.' =F83.2' 3/'
!
aV
!"#$%"$&'()*+,-.'!/*01.234'
=/4,3,/2' .8*+' =8,)' /@' /==/4,27' =87.4' J,3+/03' /=.2,27' 3+.' ://E' 3/'
1/).' 3+82' [^' <.7)..4H' %+,4' 1,2,1,4.4' 7./1.3),*' <,43/)3,/2' 82<'
J8)=,27'/@'3+.'=87.',187.4'KJ+,*+'/3+.)J,4.'/**0)4'<0.'3/',28:,F,3;'
3/'@0FF;'@F833.2'3+.'://E'4=,2.NH'Z.3J..2'4*822,27'.8*+'=8,)'/@'=87.49'
3+.'=F83.2',4')8,4.<'82<'F/J.).<'82<'3+.'=87.'30)2.<'18208FF;&'2/3'
40)=),4,27F;'3+.'4*822,27')83.',4'3+04'4F/J'K]'=87.4G1,203.'*/F/0)d'X'
=87.4G1,203.'7).;B4*8F.NH''
'
'
`,7H']H'%+.'e.)/A'L(DC')8).'://E'4*822.)'K@)/1'
+33=&GGJJJ]H=8)*H*/1G.1FG1.1:.)4G).8<;G=8)*f://E4*822.)H+31N''
!
a]'
!"#$%"$&'()*+,-.'!/*01.234'
7#8&,#($&*/+5"59:(
'
!,7,38F'*81.)8'3.*+2/F/7;9':84.<'/2'3J/B<,1.24,/28F'CC!'8))8;49'
+84'<.-.F/=.<')8=,<F;',2'3+.'F843'<.*8<.9'J,3+'3+.').40F3'3+83' *81.B
)84'2/J'/@@.)'8'-,8:F.'8F3.)283,-.'3/'@F83:.<'4*822.)4',2'182;'</*0B
1.23',187.'8*I0,4,3,/2'8==F,*83,/24H'('a'T=,A.F'*81.)8'*82',2'=),2B
*,=F.',187.'82'(Y'=87.'J,3+'8').4/F03,/2'/@'2.8)F;'\^^'<=,9'82<'.-.2'
+,7+.)' ).4/F03,/2' CC!4' 8).' 2/J' ).8<,F;' 8-8,F8:F.' 83' 1/<.43' */43H'
L)/@.44,/28F'I08F,3;'*81.)84'8).'8-8,F8:F.'J,3+']^BY^'TL,A.F'CC!4H'
C81.)84'8).'@F.A,:F.9'=/)38:F.'82<'@843'*/1=8).<'J,3+'4*822.)49'82<'
@/)' 4/1.' 8:2/)18F' </*01.23' 4/0)*.4' KL8=;)049' L8F1' F.8-.49'
J//<.2' 82<' 43/2.' 38:F.34' .3*N' 18;' :.' 3+.' /2F;' -,8:F.' ,187,27'
4/0)*.H'U,72,@,*823').4.8)*+'+84'8F4/':..2'*8)),.<'/03'/2'3.*+2,I0.4'
@/)' 1/48,*,27' F8)7.' </*01.23' ,187.4' K.H7H' 0=' 3/' 3+.' 4,_.' /@' 8'
*/1=F.3.'<.4E3/=N'@)/1'8'4.),.4'/@'8<R8*.23'*81.)8',187.'@)81.4]H''
'
`,7H'\H'b.<7.)'=87.',187.4'@)/1'3+.'$830)8F'S,43/);'T04.019'4+/J,27'
.@@.*34'/@'8@@,2.'82<'=87.B*0)F'J8)=,279'82<',FF01,283,/2',23.24,3;'-8),83,/2'
<0.'3/'=/,23B4/0)*.'F,7+3,27H'
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
#
!!.4E3/='</*01.23'1/48,*,27$!
!
!"#$%"$&'()*+,-.'!/*01.234'
a\
%+.'<,48<-82387.'/@'04,27'*81.)84'3/'8*I0,).'</*01.23',187.4',4'
3+83',3',4'1/).'<,@@,*0F3'3/'8**0)83.F;'*/23)/F'7./1.3);9'F,7+3,279'@/*04'
82<',187.'2/,4.'@/)'*81.)8B:84.<'=,*30).49'3+82'@/)'@F83:.<'/)'F,2.8)'
4*822.)4H'g./1.3),*'<,43/)3,/24'8),4.'@)/1'2/2B/)3+/7/28F'8F,721.23'
/@'3+.'*81.)8'J,3+'3+.'4*822.<'=87.'82<',2*F0<.'=.)4=.*3,-.9'8@@,2.'
82<'2/2BF,2.8)'J8)=,27'/@'3+.'</*01.23',187.\9'4/1.3,1.4').I0,),27'
4/=+,43,*83.<' \!' 4+8=.' 1/<.F4' 3/' 02J)8=' *0)F.<' =87.4' K@,7H'\NH'
b,7+3,27'@/)'*81.)8B:84.<'</*01.23',187.'8*I0,4,3,/2',4'8F4/'1/).'
<,@@,*0F3'3/'*/23)/F9'82<'3;=,*8FF;'40@@.)4'@)/1'+,7+F,7+34'82<'4,72,@,*823'
,23.24,3;' -8),83,/2' <0.' 3/' F,2.8)' /)' =/,23B4/0)*.' ,FF01,283,/2' /@' 3+.'
</*01.23H' P2' -,.J' /@' 3+.' ,2*).84,27' =).-8F.2*.' /@' <,7,38F' *81.)849'
*/24,<.)8:F.' ).4.8)*+' ,4' 2/J' ,2' =)/7).44' /2' 3.*+2,I0.4' @/)' */)B
).*3,27' 7./1.3);9' F,7+3,27' 82<' 2/,4.' <.@.*34' ,2' *81.)8B:84.<'
</*01.23',187.4Y'H''
'
()"*+,$-'!)*.",/'
'
U*,.23,@,*' 8)*+,-.4' /),7,283.' ,2' 182;' <,@@.).23' 3;=.4' /@' </*01.2349'
:03' *82' :.' *F844,@,.<' ,23/' 3J/' 18R/)' *F844.49' ()%*+),- ./0)#!.' 82<'
%'1!2- )!#/)1.H' L),18);' 4/0)*.4' 8).' 8)*+,-.4' J+.).' 3+.' </*01.23'
,34.F@',4'3+.'4*,.23,@,*'4/0)*.9'@/)'.A81=F.9'F/7'://E4'82<'<.4*),=3,/24'
/@'3+.,)').4.8)*+'J),33.2':;'@81/04'4*,.23,43459'J+,*+'+8-.'-8F0.':/3+'
84' 4*,.23,@,*' 4/0)*.4' 82<' +,43/),*8F' ).*/)<4H' U0*+' 4/0)*.4' 8).' ,<,/B
4;2*)83,*8FF;'/)782,4.<':;'3+.'J),3.)9'82<'J,FF'2/3',2'7.2.)8F'+8-.'82;'
4.1823,*' 43)0*30).' :.;/2<' 3+.' 4/0)*.' ,34.F@' J+,*+' *82' :.' 04.<' 3/'
+.F=',2@.)'</*01.23'*/23.23H''
'
P2' */23)8439' ,2<.A' ).*/)<4' 8).' .44.23,8FF;' F,434' /@' =/,23.)4' 3/' /3+.)'
4*,.23,@,*' 8)3.@8*34' KJ+,*+' 18;' 3+.14.F-.4' :.' </*01.234N9' 82<'
+.2*.'J,FF',2+.).23F;'+8-.'82',1=F,.<',2<.A'40=.)43)0*30).H'%+.;'18;'
:.':84.<'0=/2'3+.'04.'/@'8'*/11/2'=+;4,*8F').*/)<,27'1.<,01'K.H7H'
,2<.A'*8)<4N9':03'3+,4'</.42?3'708)823..'8'*/24,43.23',2<.A'43)0*30).'
/)' F8;/039' 4,2*.' F8)7.' 4*,.23,@,*' ,2<.A.4' +8-.' 0408FF;' :..2' 8**010B
F83.<' /-.)' 182;' ;.8)4' :;' 4,72,@,*823' 7)/0=4' /@' ,2<,-,<08F49' .8*+' /@'
J+/1' 18;' +8-.' 04.<' 3+.,)' /J2' ,<,/4;2*)83,*' ).*/)<,27' 43;F.H'
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
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!O2J)8==,27'=87.'*0)F').@.).2*.H!
!C/1=.2483,27',FF01,283,/2'-8),83,/2!
'
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&
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aY'
!"#$%"$&'()*+,-.'!/*01.234'
$.-.)3+.F.449' 8' */24,43.23' =+;4,*8F' 43/)87.' 1.<,01',4' 3+.' @,)43'=).B
).I0,4,3.' @/)' .@@,*,.23' /2F,2.' 8*I0,4,3,/29' 4,2*.' ,3' ,23)/<0*.4' 3+.'
=/44,:,F,3;'/@'8'>=)/<0*3,/2BF,2.?'1.*+82,*8F'4*822,27'=)/*.44H''
'
$/3' 8FF' 4*,.23,@,*' 8)*+,-.4' *82' :.' ),7,<F;' *F844,@,.<' 84' 4/0)*.4' /)'
).*/)<4&'4/1.'+,43/),*'</*01.234'K.H7H'h.=4/2?4'34/)+-/"-5+4%"/)'%+6'
=0:F,4+.<' ,2' 4.-.)8F' -/F01.4' 4=822,27' <.*8<.49' @,7H' YN' @0F@,F' :/3+'
)/F.49' 82<' S.):8),8' 4+..34' 3;=,*8FF;' ,2*F0<.' :/3+' 4=.*,1.24' 82<'
</*01.2383,/2'/2'3+.'481.'4+..3H''
'
'
`,7H'YH'QA81=F.'/@'h.=4/2?4'34/)+-/"-5+4%"/)'%+-K@)/1'+33=&GG0*R.=4H:.)E.F.;H'
.<0GR.=4/2B=)/R.*3\H+31FN-
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
a
'+33=&GG.F,:H*4H:.)E.F.;H.<0G</*4G)8).H+31F'82<'+33=&GG0*R.=4H:.)E.F.;H.<0GR.=4/2B
=)/R.*3\H+31F''
!
!"#$%"$&'()*+,-.'!/*01.234'
a5
!/*01.23'8)*+,-.4'8).'8F4/'43/).<',2'8'-8),.3;'/@'<,@@.).23'=+;4,*8F'
@/)18349' 2/3' 8FF' /@' J+,*+' 8).' .84,F;' ,187.<' K4..' :.F/JNH' %+.).@/).'
3+.' =),18);' ,440.4' 3/' :.' ).4/F-.<' J+.2' =F822,27' 3/' */2-.)3' 8'
4*,.23,@,*'8)*+,-.',23/'82'/2F,2.'<838:84.'8).&''
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VH' #+83',187,27'3.*+2,I0.4'8).'=)8*3,*8:F.'82<'.@@,*,.23'@/)'3+.'
8)*+,-.i'
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40@@,*,.23'I08F,3;':/3+'@/)'*0)).23F;'=F822.<'82<'=/44,:F.'
@030).'04.4'/@'3+.'/2F,2.'8)*+,-.i'
\H' #+83'</*01.23',187.'828F;4,4'=)/:F.14'</.4'3+.'8)*+,-.'
=).4.239'82<'3/'J+83'.A3.23'</'3+.4.'.A*..<'3+.'*8=8:,F,3,.4'
/@'*0)).23'/@@B3+.B4+.F@'"CD'4;43.14i'
'
7#,'(;,*/.<&)(
'
C8)<' 8)*+,-.4' J.).' J,<.F;' 04.<' 3+)/07+/03' 3+.' ]^3+' C.230);' @/)'
/)782,4,27'4*,.23,@,*'82<'F,3.)8);'4/0)*.49'8')/F.'2/J'@0F@,FF.<':;'*/1B
=03.)' <838:84.4H' %+.' <.-.F/=1.23' /@' 43/)87.' :/A.4' 82<' 3)8;4' 3/'
8FF/J' *8)<' ,2<.A.4' 3/' :.' .84,F;' 4.8)*+.<' /2' 3+.,)' =),18);' E.;9'
3/7.3+.)'J,3+'3+.'@8*3'3+83'*8)<')83+.)'3+82'=8=.)'J84'04.<'@/)'.8*+'
).*/)<9' 1.824' 3+83' 1/43' ,2<.A.4' 8).' ,2' 7//<' =+;4,*8F' */2<,3,/2'
K8F:.,3'4/1.3,1.4'@8<.<N'82<'*82':.'4*822.<'J,3+/03'<8187,27'3+.'
/),7,28F'4/0)*.H''
'
S/J.-.)9' <,@@.).23' *8)<' ,2<.A.4' 18;' :.' 40:R.*3' 3/' -.);' <,@@.).23'
).*/)<,27'*/2-.23,/24H'"-.)'3+.'F843'*.230);9',2@/)183,/2'+84'0408FF;'
:..2'3;=.BJ),33.29':03'8'1,2/),3;'/@'*8)<4'K82<'4/1.3,1.4'*/1=F.3.'
,2<.A.4N'8).'.23,).F;'+82<BJ),33.2H'S82<BJ),33.2'822/383,/24'/2'3;=.<'
*8)<4'8).'8F4/'*/11/2'J+.).'3+.'*8)<'38A/2/1;'+84':..2'0=<83.<H'
U/1.'8)*+,-.4'+8-.',2<.A',2@/)183,/2'/2':/3+'4,<.4'/@'3+.'*8)<9'/)'
,2<.A',2@/)183,/2'/2'/2.'4,<.9'82<'4/0)*.'<838'K40*+'84'8'4=.*,1.2'
<)8J,27N'/2'3+.'/3+.)H''
'
C8)<'8)*+,-.4'*82':.'4*822.<'04,27'8':82E'*+.I0.'4*822.)9'1/<,B
@,.<' :;' 3+.' 8<<,3,/2' /@' 4=.*,8FF;' <.-.F/=.<' 4/@3J8).' J+,*+' 8FF/J4'
*/2@,70)83,/2'/@'3+.'4*822,27'=)/*.449'3/':0,F<'8'F8)7.',187.'8)*+,-.'
@)/1' 8' 4.),.4' /@' :83*+' 4*824' J+,*+' 18;' 38E.' =F8*.' /-.)' <8;4' /)'
J..E4H'P2<,-,<08F'*8)<4'8).'4*822.<'83'8')83.'/@'8:/03'V'*8)<G4.*/2<9'
:03'3+.'/-.)+.8<'/@'3)824@.)),27'418FF':F/*E4'/@'*8)<4':8*EJ8)<4'82<'
!
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:.*804.'/@'3+.,)'8</=3,/2'84'J.:'4382<8)<49':03'2.,3+.)',4'/=3,1,4.<'
@/)' </*01.23' ).=).4.2383,/2H' `,28FF;9' 3+.).' 8).' 4/1.' ,187.' */<,27'
4382<8)<4' 3+83' +8-.' :..2' <.-.F/=.<' 4=.*,@,*8FF;' @/)' </*01.23'
*/1=).44,/2'82<'*/<,27H''
'
"2.' +;:),<' 8F7/),3+1' ,4' !RW09' J+,*+' +84' :.*/1.' 8' <.' @8*3/'
*/11.)*,8F' 4382<8)<' KZ"%%"O' .3' 8FH' V[[XN9' 82<' ,4' :84.<' 0=/2'
4.71.23,27'3+.'</*01.23',187.',23/'3+)..'F8;.)49'@/).7)/02<9'184E'
82<' :8*E7)/02<H' %+.' 184E' F8;.)9' */<.<' 04,27' @8*4,1,F.' :,28);'
*/<,27' 3.*+2,I0.4' 4.=8)83.4' @/).7)/02<' 3.A308F' <.38,F' @)/1' 3+.'
:8*E7)/02<9' J+,*+' ,4' */<.<' 04,27' 3+.' hLQg]^^^' 2830)8F' ,187.'
J8-.F.3' */<,27' 1.3+/<H' "3+.)' </*01.23' ,187.' */<,27' 3.*+2,I0.4'
03,F,4.'3+.'D.7,/2B/@BP23.).43'1.*+82,41'J,3+,2'hLQg]^^^'3/',<.23,@;'
).*38270F8)' 8).84' /@' 3.A308F' <.38,F' J+,*+' *82' 3+.2' :.' =).@.).23,8FF;'
*/<.<'KjP$'.3'8FH']^^YNH'U3,FF'/3+.)'@/)1834'8).'.A=F,*,3F;'<.4,72.<'@/)'
1,A.<' ,187.B82<B3.A3' ).=).4.2383,/2' K.H7H' L!`9' S%Tb9' D%`' 82<'
L/43U*),=3N'/)'@/)'=0).'3.A3B/2F;'8==F,*83,/24'K.H7H'(UCPPGO2,*/<.NH''
'
%+.).@/).' ,3' J/0F<' :.' =).1830).' 3/' 84401.' 3+83' 3+.).' ,4' *0)).23F;'
82;' 4,27F.' >4382<8)<?' 3/' J+,*+' 4*822.<' 8)*+,-.' </*01.23' ,187.4'
4+/0F<'8<+.).9'82<'J+,*+'J,FF'708)823..'3/'8-/,<'/:4/F.4*.2*.H'%+.'
!
!"#$%"$&'()*+,-.'!/*01.234'
6\
).8F,3;' ,4' 3+83' *).83,27' /2F,2.' 8)*+,-.4' ,4' 2/3' 8' /2*.' 82<' @/)' 8FF'
/=.)83,/29':03')83+.)'82'.-/F03,/28);'=)/*.449'J+.).'8)*+,-.'*/23.23'
82<' @/)183' J,FF' */23,20.' 3/' <.-.F/=' /-.)' 182;' ;.8)4' 84' */1=03.)'
3.*+2/F/7;' /@@.)4' =)/7).44,-.F;' 1/).' 4/=+,43,*83.<' </*01.23'
828F;4,4'82<').*/72,3,/2'3//F4H'
'
%+.' */2*F04,/2' @)/1' 3+,4' ).-,.J' ,4' 3+839' J+,F.' @030).' 8)*+,-.'
</*01.23' ,187.' =)/*.44,27' 82<' 43/)87.' @/)1834' 8).' F,E.F;' 3/'
*/23,20.' 3/' .-/F-.9' 3+.' 18R/)' :/33F.2.*E' 83' =).4.23' ,4' ,2' 8*I0,),27'
4/0)*.'<838'3+)/07+'</*01.23'4*822,27H'S.2*.9'J+.).'4*822,27',4'
02<.)38E.29',3',4'.44.23,8F'3+83'I08F,3;'4382<8)<4'8).'1.3'3+83'J,FF'2/3'
*/1=)/1,4.' @030).' </*01.23' ,187.' 828F;4,49' "CD' 82<G/)' +82<B
J),3,27').*/72,3,/2H'('1,2,101'4=.*,@,*83,/2',4'3/'4*82'8)*+,-.4',2'@0FF'
*/F/0)' 83' \^^' <=,' /)' 7).83.)9' 4,2*.' OU' 40)-.;4' /@' */11.)*,8F' "CD'
4;43.14' +8-.' 4+/J2' 3+83' =.)@/)182*.' ,4' */1=)/1,4.<' :.F/J' \^^'
<=,' ).4/F03,/29' :03' ,4' 2/3' 4,72,@,*823F;' ,1=)/-.<' 83' +,7+.)').4/F03,/24'
KDPCQ' .3' 8FH' V[[aNH' O4.@0F' 70,<82*.' /2' :.43' */11.)*,8F' =)8*3,*.' ,2'
</*01.23'4*822,27'*82':.'@/02<'83'3+.'J.:4,3.'/@'3+.'(44/*,83,/2'/@'
P2@/)183,/2' 82<' P187.' T8287.1.23' K(PPTNV]9' @/)' .A81=F.' /2.' *0)B
).23',2,3,83,-.',4'3/'<.-.F/='82',23.)283,/28F'4382<8)<'3/'<.@,2.'3+.'04.'
/@'L!`'@/)'8)*+,-,27'82<'=).4.)-,27'</*01.234V\H'
'
()"*+,$-'0+#&,'7$#35/%/'#$6'?,")&$%-%)$'
'
U*,.23,@,*' 8)*+,-.' </*01.234' 8).' /@3.2' +,7+F;' 43)0*30).<9' J,3+' 3+.'
43)0*30).' .A3.2<,27' /-.)' 3+.' */1=F.3.' 8)*+,-.' )83+.)' 3+82' :.,27'
*/2@,2.<' 3/' .8*+' ,2<,-,<08F' </*01.23H' %+.' F827087.' 3+.;' */238,2'
18;' :.' 8)*82.' 82<' 40:R.*3B4=.*,@,*' K.H7H' b83,2' 4=.*,.4' 281.49'
=)/=.)'281.4' /)'=F8*.' 281.4N' 82<' ,187.' I08F,3;'18;' :.' =//)'<0.'
:/3+' 3/' =//)' @/)183,/2' /@' 3+.' /),7,28F' 3.A3' K.H7H' 04,27' .8)F;'
3;=.J),3.)4N'82<'40:4.I0.23'87.,27'82<'<.3.),/)83,/2H'
'
%/' 18A,1,4.' */1=03.)' ).*/72,3,/2' /@' 3+.' */23.23' /@' 40*+' </*0B
1.234').I0,).4'8'@/0)'4387.'=)/*.44,27'4;43.1'K<.4*),:.<':.F/JNH'(3'
=).4.239'/2F;'3+.'3+,)<'/@'3+.4.'4387.4'K"CDN',4').8<,F;'8-8,F8:F.'84'
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
V]
'(44/*,83,/2'@/)'P2@/)183,/2'82<'P187.'T8287.1.23'P23.)283,/28F9'VV^^'#8;2.'(-.H9'
U0,3.'VV^^9'U,F-.)'U=),279'T8);F82<9']^[V^9'+33=&GGJJJH8,,1H/)7'
V\
'(PPT'L!`B()*+,-.'*/11,33..9'+33=&GGJJJH8,,1H/)7G4382<8)<4H84=iP!m]5^V\'
!
6Y'
!"#$%"$&'()*+,-.'!/*01.234'
82' /@@B3+.B4+.F@' */11.)*,8F' 4/@3J8).' =8*E87.9' J,3+' 3+.' ).40F3' 3+83'
.@@.*3,-.' /2F,2.' 8*I0,4,3,/2' /@' 4*,.23,@,*' 8)*+,-.' <838' ).18,249' ,2'
7.2.)8F9'82'02).4/F-.<').4.8)*+'82<'<.-.F/=1.23'=)/:F.1H'
'
•' %+.' @,)43' 4387.9' %+#&,' =.,=.)",//%$&' ,4' ).4=/24,:F.' @/)' =,A.FB
F.-.F' /=.)83,/24' 8==F,.<' 3/' 3+.' 4*822.<' ,187.' 3/' 4.=8)83.'
@/).7)/02<' 3.A308F' K82<' =/44,:F;' F,2.B<)8J,27N' */23.23' @)/1' 3+.'
</*01.23':8*E7)/02<H'%+,4').<0*.4'3+.'40:4.I0.23'=)/*.44,27'
F/8<' 82<' 18A,1,4.4' 3+.' =.)@/)182*.' /@' 3+.' @/FF/J,27' </*01.23'
,187.' 828F;4,4' 82<' "CD' 4387.4H' U382<B8F/2.' "CD' =8*E87.4'
0408FF;' ,2*F0<.' ,23.)28F' 3+).4+/F<,27G:,28),483,/29' :03' 3+.4.' 18;'
:.' 40:B/=3,18F' @/)' 8)*+,-.' </*01.2349' 4,2*.' 3+.;' 8).' 1/).'
0408FF;' */2@,70).<' 3/' 18A,1,4.' =.)@/)182*.' /2' :04,2.44'
</*01.234H'
'
•' ()"*+,$-' %+#&,' #$#35/%/' @(07A' ,4' 04.<' 3/' +.F=' ,<.23,@;'
<,@@.).23'4.1823,*'*/1=/2.234'/@'3+.'8)*+,-.9'J+,*+'J,FF'3;=,*8FF;'
18=' 3/' <,@@.).23' @,.F<4' /@' 3+.' */)).4=/2<,27' /2F,2.' <838:84.H'
P<.23,@,*83,/2' /@' 3+.4.' */1=/2.234' 4.)-.4' 3J/' =0)=/4.4&' @,)43' ,3'
,<.23,@,.4' J+,*+' <838:84.' @,.F<' 8' =8)3,*0F8)'3.A3' 43),27' 4+/0F<' :.'
18==.<' 3/d' 82<' 4.*/2<9' ,3' 8FF/J4' "CD' /@' 3+83' @,.F<' 3/' :.'
/=3,1,4.<':;'3+.'8==F,*83,/2'/@'@,.F<B4=.*,@,*'<,*3,/28),.4H'
'
•' B=-%"#3' C2#.#"-,.' ?,")&$%-%)$' @BC?A' ,4' 8==F,.<' 3/' 4.1823,B
*8FF;BF8:.FF.<' 40:B,187.4' /@' 3+.' */1=F.3.' </*01.23H' P34' =.)B
@/)182*.' ,4' 18A,1,4.<' :;' /=3,18F' ,187.' 3+).4+/F<,27' 3/' */1B
=.2483.'@/)'-8),8:,F,3;',2'3+.'8)*+,-.'</*01.23':8*E7)/02<9'82<'
:;' 3+.' 8==F,*83,/2' /@' @,.F<B4=.*,@,*' <,*3,/28),.4' 3/' ,1=)/-.' J/)<'
).*/72,3,/2' )83.H' "CD' 18;' :.' :84.<' 0=/2' 3+.' 04.' /@' */2@,B
70)8:F.' /@@B3+.B4+.F@' 3.*+2/F/7;9' :03' *0)).23' /@@B3+.B4+.F@' "CD'
=8*E87.4' 8).' 028:F.' 3/' 8*+,.-.' 7//<' 4;43.1' =.)@/)182*.' 02B
F.44'8071.23.<':;'3+.'/3+.)'4;43.1'*/1=/2.234'<.4*),:.<'+.).H'
'
•' %+.' "CD' 4;43.1' =)/<0*.4' 8' )8J' 3.A3' 43),27' 84' ,34' /03=039' :03'
3+,4'43),27').I0,).4'@0)3+.)'-,D-*#3'=)/-<=.)",//%$&'3/'*/2@,70).'
,3'@/)',24.)3,/2',23/'3+.'/2F,2.'<838:84.H'`,)439').70F8)'.A=).44,/2'
183*+,27'18;':.').I0,).<'3/'.A3)8*3'3+.').I0,).<'40:B43),27'@)/1'
3+.' )8J' 3.A3' 43),27' */)).4=/2<,27'3/' 82' ,<.23,@,.<' 4.1823,*' @,.F<'
K@/)'.A81=F.'8';.8)'<83.'82<G/)'8'281.'18;'2..<'3/':.'.A3)8*B
3.<' @)/1' 8' 43),27' =)/-,<,27' 8' @0FF' F,3.)8);' ).@.).2*.NH' U.*/2<9'
!
!"#$%"$&'()*+,-.'!/*01.234'
65
@0)3+.)'8==F,*83,/2'/@'@,.F<B4=.*,@,*'<,*3,/28),.4'84'=8)3'/@'3+.'3.A3'
=/43B=)/*.44,27'*82':.'04.<'3/'*/)).*3'4/1.'"CD'.))/)4H'`,28FF;9'
=/43B=)/*.44,27'*82':.'04.<'3/'*/2@,70).'3+.'/03=03'3.A3'43),274'
@/)' <,).*3' ,24.)3,/2' ,23/' 3+.' 38)7.3' /2F,2.' <838:84.9' 04,27' 8'
4382<8)<' <838:84.' ,2=03' @/)183' 40*+' 84' CUW9' 40,38:F.'
.1:.<<.<'Unb'*/1182<49'/)'82'eTb').=).4.2383,/2H'
'
7$'ED#+=3,'!5/-,+'F'G07(BC!'
!
!5/-,+'C)$",=-'
'
WP(!"CU' KW.)483,F.' P23.)8*3,-.' ()*+,-.' !/*01.23' C/2-.)4,/2'
U;43.1N' J84' 8' ).4.8)*+' =)/R.*3' 3/' <.-.F/=' 8' 04.)B*/2@,70)8:F.'
8)*+,-.' </*01.23' ).*/72,3,/2' 4;43.19' @02<.<' :;' 3+.' OM' Z,/F/7,*8F'
82<' Z/382,*8F' U*,.2*.4' 82<' Q27,2..),27' 82<' L+;4,*8F' U*,.2*.4'
D.4.8)*+' C/02*,F4' KZZUDC' 82<' QLUDCN' 02<.)' 3+.,)' R/,23' Z,/,2@/)B
183,*4' ).4.8)*+' =)/7)811.H' P3' ,23.7)83.4' ,187.' 828F;4,4' 82<' 3.A3'
=/43B=)/*.44,27'3//F4'J,3+'8'*/2@,70)8:F.'*/11.)*,8F'"CD'=8*E87.9'
3/' 7.2.)83.' 3.A3' */23.23' 3+83' *82' :.' @.<' <,).*3' ,23/' 8' 38)7.3' /2F,2.'
<838:84.9' 82<' ,2*F0<.4' 82' .@@,*,.23' 8)*+,-.' *8)<' 4*822.)' @/)' ,2,3,8F'
</*01.23',187.'8*I0,4,3,/2':84.<'/2'8'*/11.)*,8F'*+.I0.'4*822.)H'
'
O4.)' */2@,70)8:,F,3;' ,4' .44.23,8F' 3/' 8FF/J' 3+.' 4;43.1' 3/' :.' ).B
38)7.3.<' 3/' =)/*.44' <,@@.).23' 8)*+,-.49' 82<' 8F4/' -8),8:F.' F8;/034'
J,3+,2' 3+.' 481.' 8)*+,-.H' %+.' =833.)2' ).*/72,3,/2' 84=.*34' /@' 3+.'
4;43.1' K)827,27' @)/1' */F/0)' 4.71.2383,/29' 3/' </*01.23' 43)0*30).'
*F844,@,*83,/29' 3/' 8)3.@8*3' ,<.23,@,*83,/2' 82<' ).1/-8FN' 8).' 02,@/)1F;'
,1=F.1.23.<' 04,27' 8' @0__;' *F844,@,*83,/2' 4*+.1.' J+,*+' ,4' =8)8B
1.3.),4.<' J,3+,2' 3+.' 04.)' ,23.)@8*.H' "CD' =.)@/)182*.' ,4' /=3,1,4.<'
04,27' */2@,70)8:F.' 04.)' <,*3,/28),.4' F,2E.<' 3/' 4.1823,*8FF;' F8:.FF.<'
3.A3'@,.F<4',<.23,@,.<'04,27'3+.'</*01.23',187.'828F;4,4'40:B4;43.1H'
%+.' )8J' /03=03' /@' 3+.' "CD' 4;43.1' */)).4=/2<,27' 3/' .8*+' F8:.FF.<'
40:B@,.F<' /@' 3+.' </*01.23' ,187.' ,4' 3+.2' =/43B=)/*.44.<' 04,27' 8'
).70F8)' .A=).44,/2' .27,2.' 3/' */2-.)3' ,3' ,23/' 3+.' ).I0,).<' <838:84.'
@/)183' K.H7H' b.=P2<.Ad' ZQCC(b"$P' .3' 8FH' ]^^5NH' %+.' 4;43.1' J84'
<.-.F/=.<' ,2' */2R02*3,/2' J,3+' 3+.' OM' $830)8F' S,43/);' T04.01'
K$STN9'82<'+.2*.'+84'F8)7.F;':..2'.-8F083.<'/2'3+.,)'8)*+,-.'<8389'
8F3+/07+' ,3' ,4' 8F4/' ,23.2<.<' 3/' :.' 1/).' J,<.F;' 8==F,*8:F.' 3/' /3+.)'
43)0*30).<'</*01.23'8)*+,-.4H'
!
6a'
!"#$%"$&'()*+,-.'!/*01.234'
HIJ'"#.6'#."2%8,/'
!
()*+,-.4'83'3+.'OM'$830)8F'S,43/);'T04.01'K$STN'8).').*/)<.<',2'
*8)<' ,2<.A.49' J+,*+' */238,2' :,:F,/7)8=+,*8F' <838' 82<' /3+.)'
,2@/)183,/2' @/)' /2.' 4*,.23,@,*' 281.' /2' .8*+' *8)<9' F8,<' /03' ,2' 8'
4382<8)<,4.<' @/)183' K@,7H' XN' @/)' .8*+' 8)*+,-.H' S/J.-.)9' <,@@.).23'
8)*+,-.4' 18;' :.' 40:R.*3' 3/' -.);' <,@@.).23' ).*/)<,27' */2-.23,/24H'
P2@/)183,/2',4'0408FF;'3;=.BJ),33.29':03'8'1,2/),3;'/@'*8)<4'8).'.23,).F;'
+82<BJ),33.2'82<'+82<BJ),33.2'822/383,/24'8).'*/11/2H''
'
C8)<' 8)*+,-.4' 8).' 4*822.<' 04,27' 8' :82E' *+.I0.' 4*822.)' K@,7H' VN9'
1/<,@,.<' :;' 3+.' 8<<,3,/2' /@' 4=.*,8FF;' <.-.F/=.<' 4/@3J8).' J+,*+'
8FF/J4'*/2@,70)83,/2'/@'3+.'4*822,27'=)/*.449'3/':0,F<'8'F8)7.',187.'
8)*+,-.'@)/1'8'4.),.4'/@':83*+'4*824'J+,*+'18;'38E.'=F8*.'/-.)'<8;4'
/)'J..E4H'O4,27'3+,4'4;43.19'/-.)'^H5'1,FF,/2'b.=,</=3.)8'KT/3+4'82<!
Z033.)@F,.4N'82<'C/F./=3.)8'KZ..3F.4N'*8)<4'+8-.'4/'@8)':..2'4*82.<'
!
`,7H'XH'(2',2<.A'*8)<'J,3+'10F3,=F.'+82<'=),23'82<'+82<J),3,27'822/383,/24'
4+/J,27'*/1=/2.234'3/':.'.A3)8*3.<H'
!
!"#$%"$&'()*+,-.'!/*01.234'
66
83'3+.'$STH'%+.'4*822.)'+84'3+.'*8=8:,F,3;'3/'4*82':/3+'4,<.4'/@'8'
*8)<'4,10F382./04F;',2'*/F/0)'82<G/)'1/2/*+)/1.'83']^^'=,A.F4G,2*+'
).4/F03,/2'83'8')83.'/@'8:/03'V'*8)<G4.*/2<9'82<'43/).4'3+.').40F3,27'
,187.4',2'hLQg'@/)183H'P3',4'8F4/'8:F.'3/'=),23'8').@.).2*.'@,F.'201:.)'
/2' 3+.' :8*E' /@' .8*+' *8)<' @/)' *)/44B*+.*E,27' 878,243' 3+.' .F.*3)/2,*'
@,F.'8)*+,-.H''
'
Z.*804.' <,@@.).23' *8)<' 8)*+,-.4' ).*/)<' -8);,27' 4.1823,*' ,2@/)18B
3,/2' J,3+' <,@@.).23' F8;/0349' 04.)' */2@,70)83,/2' /@' 3+.' 4;43.1' ,4' ).B
I0,).<':.@/).'*/11.2*,27'828F;4,4'/@'82'8)*+,-.9'3/'<.@,2.'40,38:F.'
*8)<'3.1=F83.'F8;/03K4N9',<.23,@;'<,*3,/28),.4'3/':.'04.<'J,3+'3+.'"CD'
4;43.19' 82<' 4=.*,@;' 82;' 3.A3' =/43B=)/*.44,27' ).I0,).<' 3/' ,23.)@8*.'
3+.' 4;43.1' /03=03' 3/' 8' 38)7.3' <838:84.H' %+.' 4;43.1' ,4' */2@,70).<'
04,27' @0__;' 3.1=F83.4' 4=.*,@,.<' 04,27' 8' 7)8=+,*8F' 04.)' ,23.)@8*.9'
J+,*+'8).'<.4*),:.<',2'<.38,F'.F4.J+.).VYH'
'
!5/-,+'B8,.8%,K'
'
%+.'/-.)8FF'4;43.1'K@,7H'[N'*/24,434'/@'@/0)'18,2'*/1=/2.2349'=).B
=)/*.44,279'</*01.23'828F;4,49'"CD'K04,27'8'*/11.)*,8F'"CD'.2B
7,2.N' 82<' =/43B=)/*.44,27H' L).B=)/*.44,27' ).8<4' 3+.' /),7,28F' hLQg'
</*01.23',187.49'82<'*/2-.)34'3+.1'@)/1'.,3+.)'*/F/0)'/)'7).;'F.-.F'
,23/':,28);'@/)'4.1823,*'F8:.FF,27'=0)=/4.4H'!/*01.23'828F;4,4'3+.2'
4.71.234' 82<' 4.1823,*8FF;' F8:.F4' ,1=/)3823' 3.A3' @,.F<4H' %+.' /03=03'
2/)18FF;' */24,434' /@' F8:.FF.<' 3.A3' @,.F<' */F/0)' /)' 7).;BF.-.F' 40:B
,187.4',2'3+.'4;43.1?4',23.)28F',187.'@/)183'KL$TNH'S/J.-.)9'=).B
=)/*.44,27'*82'8F4/':.').*8FF.<'8@3.)'!/*01.23'(28F;4,4'4/'3+83'3+.'
F8:.FF.<' 3.A3' @,.F<' 40:B,187.4' *82' :.' */2-.)3.<' ,23/' :,28);' 02<.)'
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Digital Imaging of Biological Type Digital Imaging of Biological Type

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