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NAOSITE: Nagasaki University's Academic Output SITE
Title
Palynomorphs from the Santonian Uge Member of the Taneichi Formation,
Northeast Japan
Author(s)
Takahashi, Kiyoshi; Sugiyama, Ryozo
Citation
長崎大学教養部紀要. 自然科学篇. 1990, 30(2), p.133-573
Issue Date
1990-01
URL
http://hdl.handle.net/10069/16606
Right
This document is downloaded at: 2016-10-20T14:45:30Z
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Bull. Faculty of Liberal Arts. Nagasaki Univ.CNatural Science),30(2),133-573 (JanuarY,1990)
Palynomorphs from the Santonian Uge Member of
the Taneichi Formation, Northeast Japan
Kiyoshi T AKAHASHI* and Ryozo
SUCIYAMA **
(Received October 13,1989)
Abstract
The basal Uge Member of the Taneichi Formation consisting mainly of terrigenous
sediments, viz. basal conglomerate, rhyolitic tuff, sandstones, coaly shales etc, was palynologically studied. A total of 303 species of palynomorphs which consist of 115 spores,
90 gymnospermic pollen, 3 gymnospem-angiosperm incertae sedis, 85 angiospermic pollen,
7 phytoplankton and 3 incertae sedis were described and illustrated.
The following forms are new: Appendicisporites giganti/ormis n. sp., A. kanukaensis
n. sp., Baculatisporites giganticus n. sp., Balmeisporites nipponicus n. sp., Camarozonosporites (Camarozonosporites) similis n. sp., Cicatricosisporites minuticanaliculatus n. sp.,
C. senonicus n. sp., Cingulatisporites iwatensis n. sp., Foveosporitesper/ossus n. sp., Gleicheniidites verrucatus n. sp., Laevigatosporites longus n. sp., L. nitidulus n. sp., Leiotriletes giganticus n. sp., Lycopodiacidites circularis n. sp., Lygodiidites tohokuensis n. sp., Punctatisporites granulatus n. sp., Todisporites grandi/ormis n. sp., Toroisporis (Duplotoroisporis) triangulus n. sp., Trachysporites microverrucatus n. sp., Trilites pulchellus n. sp.,
T. pustulosus n. sp., Triplanosporites giganteus n. sp., T. rikuchuensis n. sp., T. taneichiensis n. sp., Uruhtlatisporites /lexuosus n. sp., Verrucatosporites verruculosus n. sp., Abiespollenites minus n. sp., Alisporites enormis n. sp., Callialasporites ugensis n. sp., Cedripites sanrikuensis n. sp., Classopollis grandissimus n. sp., C. taneichiensis n. sp., Cycadopites mirus n. sp., Ephedripites (Spiralipites) elongatus n. sp., lnaperturopollenites rugatus n~ sp., Phyllocladidites globulosus n. sp., Piceapollis grandi/ormis n. sp., Pityosporites
cretaceus n. sp., Podocarpidites senonicus n. sp., Rugubivesiculites japonicus n. sp., R.
sphaericus n. sp., Clavatipollenites variabilis n. sp., Cupuli/eroidaepollenites lanceolatus
n. sp., Foveotricolpites /astidiosus n. sp., F. globosus n. sp., Foveotricolporites gloriosus
n. sp., F. grandiformis n. sp., Ilexpollenites minus n. sp., Potamogetonacidites senonicus
n. sp., Retitrescolpites pseudoazemae n. sp., Rousea elegantula n. sp., R. reticosa n. sp.,
R. triangulata n. sp., R. ugensis n. sp., Satishia pomposa n. sp., S. tri/ormis n. sp., S.
*
**
Department of Geology, Faculty of Liberal Arts, Nagasaki University, 1-14 BunkyoCho, 852 Nagasaki, Japan.
Morioka Daiichi Senior High School, Ueda 3 Chome 2-1, 020 Morioka, Japan.
134
K.
TAKAHASHI
& R.
SUGIYAMA
uni/ormis n. sp., Symplocacites microreticulatus n. sp., Tricolpites ellipsoideus n. sp., T.
ovi/ormis n. sp., T. sphaeroides n. sp., and Tricolpopollenites baculatus n. sp.
Furthermore, new combinations are proposed: Appendicisporites auri/er Verbizkaja
n. comb., A. cr. bellus Markova n. comb., A. macrorhyzus Maljavkina n. comb., A. cf.
pseudomacrorhyzus Markova n. comb., Ephedripites (Spiralipites) longus Song & Zheng n.
comb., E. (S.) perlatus Wang&Zhao n. comb., E. (S.) praeclarus Chlonova n. comb.,
Pityosporites alatipollenites Rouse n. comb., P. microsibiricus Zaklinskaja n. comb., P.
cf. pini/ormis Zaklinskaja n. comb., Ilexpollenites claviger Takahashi n. comb. and Tricolpites rudis Takahashi n. comb.
The middle Okonai Member consists mostly of brown to greenish gray sandstone with
Crassostrea fossil bank in the lower part and fine sandstone which yields Sphenoceramus
sanrikuensis Matsumoto & Sugiyama together with other fossils in the upper part.
Formerly the Taneichi Formation was considered as the Neogene,but later revised by
Terui et al. (1975) to the middle Upper Cretaceous (Santonian) on the evidence of ammonoids and inoceramid. The Uge Member resembles lithologically the basal Tamagawa Formation of the Kuji Group. Crassostrea fossil bank in the lower horizon of the Okonai
Member is closely similar to that of the Tamagawa Foramation. Accordingly, the Uge
Member and the lower part of the Okonai Member are correlated with the Tamagawa
Formation. The upper part of the Okonai Member is like to the Kunitan Formation of
the Kuji Group in which Inoceramus (Platyceramus) japonicus Nagao & Matsumoto occurs
in yielding Sphenoceramus sanrikuensis Matsumoto & Sugiyama which is an effective member of the Platyceramus japonicus zone and indicates the lowest Campanian in age.
The Santonian Uge playnoflora is characterized by the following spores and pollen
grains: Aequitriradites verrucosus, Appendicisporites spp., Balmeisporites nipponicus,
Camarozonosporites spp., Cicatricosisporites spp., Cyathidites spp., Gleicheniidites senonicus, Jimboisporites senonicus, Lygodiidites spp., Patellasporites spp., Trilites spp., Zlivisporis novomexicanus, Araucariacites australis, Callialasporites ugensis, Classopollis spp.,
Cupressacites spp., Ephedripites spp., Phyllocladidites spp.. Pristinuspollenites microsaccus, Rugubivesiculites spp., Vitreisporites pallidus, Clavatipollenites variabilis, Asteropollis clavatus, Callistopollenites radiatostriatus, Fibulapollis spp., Rousea spp., Satishia
spp., and Tricolpites spp.
Contents
Introduction
········ .. ······································135
Stratigraphic and palaeontologie notes of the Taneichi Formation ···························135
Materials and method
138
Palynologic assemblage··
140
Palynomorphs from the Uge Member
·
·.. ··
·..· ·.. ·147
Spores
· ·.. ·· .. ·
·· .. ·.. ··147
·.. ·
Gymnospermic pollen· .. ·· .. ·· .. ·· .. ·· .. ·.. ·.. ·
Gymnosperm-angiosperm incertae sedis ·.. ·..·· .. ··
·· .. ·.. ·.. ·.. ·.. ···· ·.. ·
·
·· .. ·.... ·.. ·.. ·.. ·· .. ·· ·.. ·.. ··
151
154
Palynomorphs from the Santonian Uge Member
Angiospermic pollen
Phytoplankton
Incertae sedis
Systematic description
Spores
·.. ············· .. ··
Gymnospermic pollen·· .. ·.. ·.. ·.. ·.. ··· ..···
Gymnosperm-angiosperm incertae sedis
Angiospermic pollen
Phytoplankton
Incertae sedis
References
·· .. ·
135
154
157
157
157
·.. 157
·· .. ······ .. ·· .. ·.. ·.. ···· .. ······· .. ·.. ··· .. ·.. 234
296
299
351
355
356
Introduction
The Taneichi Formation, outcropping in a long and narrow belt area along
Rikuchu Coast in northeastern Honshu, consists predominantly of sandstones
of marine facies in middle and upper horizons and coaly shales-sandstones
of non-marine facies in lower horizon. The rocks of the middle horizon
contain some macrofossils, marine faunas and silicified woods, by contrast
the lower yield many microfossils, spores and pollen grains, including some
microphytoplankton. The abundant spore-pollen contents from the lower
horizon are described here with the purpose that they will play an important
role in constructing the Upper Cretaceous palynobiostratigraphy of Japan
and be useful in determining the age.
Miki (1972) described many well preserved palynomorphs from the Kuji
Group in the vicinity of Kuji city, being situated in south ca. 20 km of Taneichi
town. These palynomorphs are compared with those of the Taneichi Formation. The spore-pollen data of Santonian sediments of Japan have been
published only from the Futaba Group [Miki, 1972 ; Takahashi 1973 (971),
1988J and the Kuji Group (Tokunaga & Takase, 1968; Miki, 1972).
Stratigraphic and palaeontologie notes of
the Taneichi Formation
The Taneichi Formaion, lying unconformably on the Lower Cretaceous
granite, consists of the basal Uge Member, the middle Okonai Member, and
the upper Yagi Member (Sugiyama, 1982, 1983; Matsumoto & Sugiyama, 1985,
1986). This is composed predominantly of marine and non-marine sediments
yielding macro- and microfossils. The basal Uge Member consists predomi-
136
K.
TAKAHASHI
& R.
SUGIYAMA
nantly of terrigeous sediments, viz. basal conglomerate, rhyolitic tuff, sandstone, coaly shales or sandstones etc. and abounds in carbonaceous matter
on the whole. The basal conglomerate consists mostly of insufficiently rounded cobbles of granite, hornfels, chert etc., less than 20 cm in diameter and is
ca. 2 m in thickness. A bluish gray rhyolitic tuff (ca. 1 m in thickness) on the
basal conglomerate is pursued without difficulty in all the areas in which the
Taneichi Formation is distributed. Moreover, in upper horizon than the tuff
~--+---1--
36
···· 1
0.....
oo
......
2
I:
0
:~ 3
k+++~4
X
5
,......
HACHINOHE
Text-fig. 1.
Situation and geologic map of the Taneichi Formation entered localities
eX) and numbers of the collected samples.
137
there are coaly shales or sandstones and sandstones with closs-laminae. All
the samples studied palynologically were provided from these coaly shales or
sandstones. Sugiyama (1982) illustrated and described preliminarily some
palynomorphs obtained from these rocks.
The Okonai Member (ca. 140 m in maximum thickness) consists mostly of
brown to greenish gray sandstone with Crassostrea fossil banks in the lower
part and fine sandstone which yields a new inoceramid, Sphenoceramus san:.
.
"
.,'.
"
'.'
'. :.o : .. ':
:.... :'.. ','.
~
..
.',',.
· .....
.'.: .':
"
Vagi M.
~-:~.~~
":.. ',
'
~
.
._ --..........
.....:.-. Sphenoceramus sanrtkuensls
"
'""~7
',": . "' .
.. .. : ::.. :
,"
::".:/<:.::
'.:.~
'
-: ..
..:',
.,
·
•
.
.
;.' ...
,,'
"
"
.,
...
.'
..
,,'.
Okonai M.
J::
I"
: . 0'
......
0-
U
"
0-
·
....
..
'
:',' , '
o~:'~;";'~.~~
e
d
c
b
a
Crassostrea
..
. "
':.':.' :,,:.:
:," :,' :.', ...
::.::.:
~.
",
:",
15.36
34
::...........
.... ~ 06.33
17,31,39,40
Uge M.
i=' +
+ +
+ +
+ +
+ +
Text-fig. 2.
Granite (Lower Cretaceous)
Synthetic columnar section of the Taneichi Formation and horizons of the
collected samples and fossils.
138
K.
TAKAHASHI
& R.
SUGIYAMA
rikuensis Matsumoto & Sugiyama (1985, 1986), together with other bivalvia,
shark teeth, silicified woods, amber, trace fossils a sedimentary environment
of shallow sea near land is expected. Many silicified woods yield in the fossil
banks of the upper part of the Member. Sugiyama (1982) disriminated already
the following species of the silicified woods: Pityoxylon spp., Araucarioxylon
kiiense Ogura, Cupressinoxylon vectense Barber, Taxodioxylon albertense
Penhallow, a tree fern which was newly named as Tempskya iwatensis Nishida (1986).
The uppermost Yagi Member consists predominantly of a massive sandstone, ca. 25 m in thickness, only in the Yagi area and almost is continuous
under the surface of the sea.
The Taneichi Formation was formerly considered as the Neogene, but
later revised by Terui et al. (1975) to the middle Upper Cretaceous (Urakawan, K6 -Santonian) on the evidence of ammonoids [Polyptychoceras cf. subundatum (Yokoyama) etc. ] and inoceramid (Inoceramus naumanni Yokoyama).
This is approximately correlated with the Upper Cretaceous Kuji Group in
south ca. 20 km of Taneichi, and the Okonai Member resembles generally the
Kunitan Formation, middle part of the Kuji Group, in which Inoceramus
(Platyceramus) japonicus Nagao et Matsumoto occurs among others.
Recently, Toshimitsu (1988) suggested that the Sphenoceramus sanrikuensisS. cristatus-Inoceramus (Cordiceramus) kanmerai Subzone occupies the main
part of the Platyceramus japonicus Zone which indicates the lowest Campanian age. Therefore, the Uge Member and the lower part of the Okonai
Member with the Crassostrea fossil bank are correlated with the Tamagawa
Formation of the Kuji Group.
Materials and method
All the samples were collected by one of the authors, Sugiyama, from
several localities along Rikuchi Coast of north Iwate Prefecture, as follows,
with their relative positions and horizons shown in text-figs. 1-3.
Sample no.! slide no.
locality
south of Kanuka
C 31 a-I
C 33 a-k
north of Uge station (A)
C34 a-e
Uge
C 36 a-f
north of Uge station (B)
C 39 a-I
Uge harbor (A)
C 40 a-k
Uge harbor (B)
139
C 6d-e
C 15 d-e
C 17 k-l
cliff near a private house, north of Uge station
north of Uge station
Uge harbor
Yogi
.
"
M.
"
.
.'
".'
'0"
-~
:~:.
~"..;..
r:-.-....
.
"
.
'.
•
~;....:
.~:
·· " ", ' : '
· : .: .
~.
....
. '." ..
•
'0
'
••
'. .
'
",
0'"
•
.o
........
"
.
LL.
'0
••
'
'
:.
......
.
••
.. ' .
...
\
Okano;
M.
,
. ':
.c
...
'
.0 •..
,
"
,
"
.',
c::
"
. '.'
"
u
Q)
.
,',
'
..
.
..
.
,
.
"
--.",
.
.'
,"
~ 1---------t\2~
~~r------J.;..._-:....., , ,. 1536
Uge
- ---
M.
,
..:...
''':'''-
','
•
~
~
C
n1
'-
C)
Text-fig. 3.
'"
+++-j
+ +-
~:'
,
"
~1,~
.
'".
' . '
06,33
",,'
to
m~t7.39,40 ~--I
~~
34'
:.:
: .
0"
.
",
: ..
-:-:
b
~+
+ -I
++r-l
A
B
d
c
a
"
+ i
+
+++
f
e
c
o
Columnar sections of the Taneichi Formation in four localities and horizons
of the collected samples.
A : Uge harbor (south of Uge)
B : North of Uge station (Uge-Okonai)
C : Yagi-Shukunohe
D : South of Kanuka
a : conglomerate; b : sandstone; c : tuff; d : coal or coaly shale;
e : siltstone; f : marine fossils.
140
K.
TAKAHASHI
& R.
SUGIYAMA
They were relatively abundant in sprre-pollen contents except some samples. One of the authors, Takahashi, examined several strewn slides, which
were prepared by Sugiyama, under the Nikon Apophot microscope with Plan
and Apo objectives and studies several specimens using a JEOLCO JSM T200
scanning electron microscope (SEM).
All the slides containg specimens studied are kept in the Department of
Geology, Faculty of Liberal Arts, Nagasaki University.
Palynologic assemblage
The total assemblage of identified and unidentified palynomorphs from 9
samples of the Uge Member amounts to 303 form-species, namely 115 spores
(ca. 38%),90 gymnospermic pollen (ca. 30%), 3 gymnospem-angiosperm
incertae sedis (ca. 1 %), 85 angiospermic pollen (28%), 7 phytoplankton (2. 3
%) and 3 incertae sedis (ca. 1 %). Within this palynoflora about 47%043
species) could be referred to previously described palynomorphs mainly from
Europe, Japan, Siberia, North America, Australia and New Zealand. Although
the Taneichi Formation is distributing in the area not far from Kuji city in
where the Kuji Group is developed and Miki (972) described and illustrated
102 palynomorphs from the Tamagawa and Sawayama Formations of the
Kuji Group, 71 species (about 23 %) were newly named. Many forms (approximately 28 %) which were either very infrequently recorded within the
present assemblage or in an imperfect state of preservation, had yet to be
excluded from identification below generic level.
Tab. 1 Species composition of the Uge palynoflora.
i
GymnospermI
Gymnospermic
Angiospermic
Incertae
angiosperm
Phytoplankton
Total
:Pteridophyte
pollen
pollen
sedis
!
(? )
Number of
species
. 115(37.9")
I
90(29.7")
Formerly __
cribed species
(18.5")·
56-(48.7")
(15.8")
48-(53.3")
New species
(10.2")
31-(27")
(5.3")
16(17.8")
• (" for total number of species)
3(0.99")
0
(0.33")
1-(33.3")
upper row
(" for number of species in each section) •.• lower roW
85(28")
7(2.3")
36 (11.9")
(42.3")
(0.99")
3-(42.8")
(7.6")
23-·
(27")
0
3(0.99")
303
0
143
0
71
141
1) Pteridophyte
Among the trilete spores, the following species occurred frequently or
abundantly: Appendicisporites distocarinatus Dettmann & Playford, A.
giganticus n. sp., A. kanukaensis n. sp., A. taneichianus n. sp., Camarozonosporites (Camarozonosporites) similis n. sp., C. (Hamulatisporis) hamulatis
Krutzsch, Cicatricosisporites minuticanaliculatus n. sp., C. senonicus n. sp.,
Cyathidites australis Couper, C. minor Couper, Deltoidospora diaphana Wilson
& Webster, Dictyophyllidites harrisii Couper, Gleicheniidites senonicus Ross,
Leiotriletes giganticus n. sp., Monoleiotriletes gracilis Krutzsch, M. grandissimus n. sp., Patellasporites polyverrucifer n. sp., P. verrucatus n. sp., Toroisporis (Duplotoroisporis) triangulus n. sp., Trachysporites microverrucatus n. sp., Trilites pulchellus n. sp., T. pustulosus n. sp., and Zlivisporis
novomexicanus (Anderson) Leffingwell.
Several trilete spores which are morphologically remarkable make splendidly the Uge palynoflora: Aequitriradites verrucosus (Cookson & Dettmann)
Cookson & Dettmann, Appendicisporites spp., Balmeisporites nipponicus n.
sp., Camarozonosporites spp., Cicatricosisporites spp., Jimboisporites senonicus Miki, Lycopodiacidites circularis n. sp., Lygodiidites spp., Patellasporites
spp., Trachysporites microverrucatus n. sp., Trilites spp. and Zlivisporis novomexicanus (Anderson) Leffingwell.
With respect to the monolete spores, Laevigatosporites dehiscens Takahashi and L. nitidulus n. sp. appeared frequently.
Palynomorphs from the Uge Member which were formerly recorded in
sediments of Europe, Japan, North America, Australia etc. vary no doubt in
their stratigraphic distribution. There are several species (e. g. Aequitriradites verrucosus, Cyathidites australis etc.) which appeared for the first time
during the Jurassic or the Lower Cretaceous and cover all the Upper Cretaceous
stages whereas others (e. g. Deltoidospora nodaensis, Jimboisporites senonicus
etc.) reveal a much more restricted range in the Upper Cretaceous (Tab. 2 ).
Almost 66% of the dispersed spores of the Uge Member can be associated
with recent spore families, in some cases even with the genera: Athyriaceae
(Athyrium), Cyatheaceae (Cyathea), Dipteridaceae (Dictyophyllum), Gleicheniaceae (Gleichenia), Hymenophyllaceae (?), Lycopodiaceae (Lycopodium),
Osmundaceae (Osmunda, Todites) Polypodiaceae, Schizaeaceae ( Anemia,
Mohria, Lygodium), and Selaginellaceae CSelaginella).
Tab. 2 Range chart of Cretaceous spores noted in the Uge Member.
~
Albian
Cenoman.
Turonian
Coniac.
Santon.
Campan.
Maastrich.
Cicatricosisporites dorogensis
:::t;:::::::::::::::::::::::::::::: ::::::::::::::::::::::::::::~:::::::::: ::::::::::::::::::::::~:f.::::::::::: ':::::::::::::::::::::::::::::::::::::::: ::::::::::::::::::::::::::::::::::::::::: :::::::::::::::::::::::::::::::::::::::: .:~::::::::::::::::::::::::::::::::::::: :::
Gleicheniidites senonicus
~ ~t~ ~ ~ ~ ~ ;~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~;~;~ ~ ~;~;~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ g~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ i~ ~ ~ ~ ~ g~ ~ g ~ i~ ~i i~ i~ ~ ~ i~ ~ ~ ;~ ~ ~;~ ~;~;(;~;~;~i;~;~;~;~;~;~;~;~;~i ;~;~ ~;n~
Aequitriradites verrucosus
Dictyophyllidites harrisii
Leiotriletes rotundiformis
Appendicisporites exilioides
Deltoidospora cascadensis
Deltoidospora diaphana
Cyathidites australis
cyathidites minor
Deltoidospora psilostoma
Cicatricosisporites australiensis
Appendicisporites distocarinatus
tt:::;;::~;g:~ .~:~;:;:::::::~:~;: ;: :=: ;:; :;:;:;~;: ::.:::.::;;::::::::::::::::::::::::: :~::~~:::.:~::~:, X,::;:X{{@:;:::<i,:::!:;
. .· . . · . .i.. ·.. ·..
:~ J~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ t~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ t~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ t~ ~;~;~ ;~;~;~;~;~;~;~;~;~;~;~;~;~;~l;~;~;~ ~;~;~;~ ;~ ;~ ~ ~ ~;~ ~ ~ t: : : : : : : : :~: : : : :~: : : : ,
I··.. ····..··········l······..··..········f·..········..······
J;;;;;;;;;;;;;; ;~;;;;;;;';~';~;; ;;.~:~;;~;~~~;~~~:r~;~;,~;~~;;;~~;~r~:~~~;~;~;;~~
:J~~~~~~~~~~~~~~~~~~~~gg~~;~ ~ ~ ;~;~;~;~;~;~;~;~;~ ~ ~ ~ ~ ;~;~ ~ ;~: ~ ;~ ~;~;g~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~;~;~:1
~
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:>
~
5;
:>
rn
:::::::::::::::::::: ::::::::::::::::::::'
::s
~
.
,
f'
conveX1-' orm1-S
Laevigatos porite s dehiscens
.
.
.
ovo"deus
v
Laev ~gatospo~~tes
,L-t;
lJ
.
.
.
Laev1-gatospor1-tes
prom1-nens
Laev ~·gatospo~·tes
'"
J,.
'"
senon~·cus
'"
Triplanosporites minutulus
Camaro2onosporites (H.) hamulatis
Laevigatosporites probatus
Jimboisporites senoniaus
Foveotriletes scrobiculatus
Zlivisporis novomexicanus
Biretisporites incrassatu8
t>.:>
~;f;~;~;~ ~ ~ ~;~ ~;~ ~ ~ ~ ;~: ~ ~ ~ ~ ~ ~ ~ ~ ~:~ ~ ~ ~ ~ ~ ~ :~:~ ~ ~:~:~:~:~ ~:~ ~:~ ~:~ ~ ~:~:~:~ ~ ~ :~: ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ :~:~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ .~ ~ ~ ~ ~ ~ ~ ~ :~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ :;~ ~ ~ ~ ~ ~ ~ ~ ~ ; ~ ~;~ ~ ;~ ~ ;~ ; :
Appendicisporites pseudomaarOrhY2Us~i»:»»:»~~:~»»»»»»»»»» ~»»»»»»»»:~
.. .............. ~
"
..
De l toidospora nodaensis
.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:~:~:~:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:~:.:.:..:.:.:~:~:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.: :.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:. .:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.: :.:.:.:.:
· t r1-. 1"e t es
Le '1-0
~
I
I
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.
::::::::::::::::::::::::::::::::::::::::: ::::::::::::::::::::::::::::::::::::::::: ::::::::::::::::::::::::::::::::::::::::: ::::::::::::~:::~::::::::~::~:::::::::
.
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•••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••
.....................
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..........................
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...........................
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.
.................
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.
'.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.,:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:••It.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.l
,
'
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.
.
1••••••••• • •••••••••• 1
.::::::::::::::::::::
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:
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l
,
en
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3:
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Tab. 3 Range chart of Cretaceous gymnospermous pollen noted in the Uge Member.
Albian
Cenoman.
Turonian
Santon.
Coniac.
Campan.
Maastrich.
Inaperturopollenites dubius
t:::::::::~::::::::::::::::::: .::::::::::::::::::::::::::::::::::::::: :::::::::::::::::::::::::::::::::::::::: :::::::::::::::::::::::::::::::::::::::: :::::::::::::::::::::::::::::::::::::::: :;:::::::::::::::::::::::::::::::::::::: :::::::::::::::::::::::::::::::::::::::~ •.
Araucariacites australis
Cedripites medius
t;1;:~!.f;' ;!.;:'i!.~f;~; ::'[;;;E,;; i;:'~i;';:'i~i; '~ ~r;[;i~[;:J.;~ 1;'i;?~[;!.~~ ~;;';?~!!.;';ft
Cedripites microsaccoides
Ephedripites (E.) notensis
Ephedripites (S.) longus
Phyllocladidites mawsonii
Podocarpidites ellipticus
Ephedripites (S.) praeclarus
Ephedripites (S.) perlatus
Psophosphaera aggereloides
Prisnnuspollenites
':"::.'-::::.-:::::::. :::::::::::::::::::. :::::::......: ::::::..: : .:::::::::::::::::::. :::::::..:::::::::::: .:::::::::::::::::::. ::::::::::::::::::::,
E~~~ri~E~~~~~~?~~i~~E~~E~fi~~~~~~~~~:~~~~~~~ifi~j;~j
f:::::::::::::::::::::::::::::·:::::::::::::::::::::::::::::::::::::::.:::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::
....................................................................................................................... ·
·:1
microsaccjr:i:':i:i~::~:i;:':':;:';:'::if::i:' ::~::=:::~ :;~~:':'f:;::' :::;'::::;:1: :~~:~;~;~z;;;~=*:
~~~:~::
Vitreisporites
Araucariacites
Podocarpidites bifo~is
Ephedripites (E.) chaloneri
,~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ;~ ~ ~ ~ :~ ~ ~ ~;~ ~ ~; ; ;~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ .~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ .~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ ~ : : : : : : : : : : : : : I
Pityosporites scopulipites
Ephedripites (E.) regularis
1
Ephedripites (D.) scabridus
3
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t:::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::: :::::::::::::::::::::::::::::::::::::::: ::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::::.
Sciadopityspollenites antiquus
Psophosphaera pseudotsugoides
Cupressacites cU8pidataefo~is
Ephedripites (S.) ellipsoideus
Inaperturopollenites parvus
Pityosporites siegburgensis
Pityosporites alifo~is
Phylloeladidites ovatus
Inaperturopollenites laevigatus
'"d
e.
'<
~
::l
;::::::::::::::::::::::::::::::::::::::::i::::::::::::::::::::::::::::::::::::::::±::::::::::::::::::::::::::::::::::::::::i::::::::::::::::::::::::::::::::::::::::±::::::::::::::::::::::::::::::::::::::::f.::::::::::::::::::::::::::::::::::::::::+:.
C
aq
(I)
'::::::::::::::::::::::::::::::::::::::::i::::::::::::::::::::::::::::::::::::::::±::::::::::::::::::::::::::::::::::::::::f.::::::::::::::::::::::::::::::::::::::::+;
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:::::::::::::::::::::::::::::::::::::::::1::::::::::::::::::::=:::::::::::::::::::i:::::::::::::::::::::::::::::::::::::::::
....................
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·::::::::::::::::::::::::::::::::::::::::t:::::::::::::::::::::::::::::::::::::::l::::::::::::::::::::::::::::::::::::::::.
··· ·..···· .. ··l ::,::::::..: : ..:·..:..·t··
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:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.:.~.
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c.:>
144
K.
TAKAHASHI
& R.
SUGIYAMA
2) Gymnospermic pollen
The gymnospermic pollen assemblage of the Uge Member include some
morphologcially characteristic forms: Alisporites spp. (Pinaceae), Araucariacites spp. (Araucariaceae), Callialasporites ugensis n. sp. (Araucariaceae
or Podocarpaceae), Classopollis spp., (Pagiophyllum or Brachyphyllum),
Phyllocladidites spp. (Podocarpaceae, Phyllocladus), Pristinuspollenites
microsaccus (Couper) B. D. Tschudy, Rugubivesiculites spp. (Podocarpaceae),
and Vitreisporites pallidus (Reissinger) Nilsson (Caytoniales, Caytonanthus).
Disaccate conifer pollen are Pinaceae (Abiespollenites, Alisporites, Cedripites, Piceapollis, Pityosporites), Podocarpaceae (Phyllocladidites, Podocarpidites, Rugubiuesiculites) , and Caytoniales (Vetreisporites). Non-saccate
conifer pollen consist of Araucariaceae (Araucariacites, Callialasporites),
Taxodiaceae-Cupressaceae (Cupressacites, Inaperturopollenites, Sciadopityspollenites, Sequoiapollenites) and Pinaceae (Psophosphaera).
Monosulcate and polyplicate pollen forms of the Cycadaceae and Ephedraceae are a minority (about 24% of species number) of the gymnospermic
pollen group of the Uge Member.
Many gymnospermic pollen range from the Jurassic or Lower Cretaceous to Tertiary or Upper Cretaceous. 4 species, Ephedripites (E.) chaloneri
Brenner, E. (E.) regularis Hoeken-Klinkenberg, E. (D.) scabridus Song &
Zheng, and Phyllocladidites ovatus Takahashi, reveal a much more restricted
range which does not cross the Santonian age (Tab. 3).
3) Gymnosperm-angiosperm incertae sedis
Clavatipollenites variabilis n. sp. and 2 species of Clavainaperturites were
distinguished. Clauatipollenites uariabilis n. sp. occurred abundantly from
the Uge Member and resembles closely Clavatipollenites tenellus Phillips &
Felix (1972) from the Albian of Lousisiana and Kansas (U.S.A.). The genus
Clavatipollenites possesses a sulcus-like furrow and clavate sculpture.
Therefore, this pollen provides both gymnospermic and angiospermic characteristics.
4) Angiospermic pollen
The angiospermous pollen grains from the Uge Member include 85 species,
but the ratios of abundances are not so high.
The following species are remarkable in respect of their morphological
features: Atriopollis cf. indivisus Agasie, Asteropollis clauatus (Phillips &
Felix) Ward, Callistopollenites radiatostriatus (Mtchedlishivili) Srivastava,
145
Fibulapollis enodatus (Chlonova) Takahashi, Fibulapollis evanidus (Chlonova)
Takahashi, Foveotricolpites spp., Foveotricolporites spp., Phimopollenites
pannosus (Dettman & Palyford) Dettmann, Rousea spp., Satishia spp.,
Symplocacites micropunctatus n. sp. and S. microreticulatus n. sp.
Not only the monocolpate but monoporate pollen groups appear rarely.
Only three pollen grains of potamogetonaceous plant were found. The tricolpate
and tricolporate pollen groups include many genera and species which are of
almost Japanese and European types and much lesser North American or
Australian ones, and in which 22 new species are mentioned. Two tricolpate
pollen genera with heterobrochate sculpture, Rousea Srivastava (1969) and
Satishia Ward (1986), were distinguished from the genus Tricolpites with
homobrochate sculpture.
Asteropollis clavatus (Phillips & Felix,1972) Ward (1986) with a triradiate
sulcus is reported so far from Albian to lower Cenomanian of U.S.A. and Australia and Callistopollenites radiatostriatus (Mtchedlishivili, 1961) Srivastava
(1969) is hitherto a Maastrichtian type from Siberia, Japan, and Canada.
In a systematic arrangement the dispersed angiospermous pollen grains
except unknown botanical affinity can be related to the following plant families:
Monocotyledoneae: Potamogetonaceae, Gramineae, Palmae, ? Liliaceae.
Dicotyledoneae: Cyrillaceae, Santalaceae or Loranthaceae, Chloranthaceae (?), Aquifoliaceae, Nyssaceae, Fagaceae, Oleaceae, Salicaceae, Juglandaceae, Symplcaceae (?), Hamamelidaceae.
5) Phytoplankton
Some phytoplankton, a dinoflagellate cyst, 3 Pterospermataceae, and 3
acritarchs, were found rarely from the Uge Member. The authors considered
that the Uge Member is influenced by a non-marine environment of sedimentation, because it abounds in coaly matters and yields no marine fossil. However, the above-mentioned phytoplankton show clearly their marine influence.
Two species of Pterospermella range from the Lower Cretaceous to Palaeogene i~ appearance and Cymatiosphaera reticulosa occurs from the Oligocene to Pleistocene.
Miki (1972) described and illustrated 102 palynomorphs from the Tamagawa
and Sawayama Formations of the Kuji Group being situated in south ca. 20
km of Taneichi town.
The Santonian Tamagawa palynoflora is closely similar to the lower
146
K.
TAKAHASHI
& R.
SUGIYAMA
Campanian Sawayama palynoflora except some palynomorphs which are not
common to each Formation: e. g. Balmeisporites minutus, Cyathidites australis, Deltoidospora nodaensis, Uveaesporites simplex, Jimboisporites kujiensis,
Schizosporis scabratus, Foveosporites sawayamensis, Azonia (al. Ocellipollis)
obliquus etc.
15 species of all palynomorphs from the Kuji Group are common to the
Uge palynoflora, namely Gleicheniidites senonicus, Cicatricosisporites australiensis, C. dorogensis, Cyathidites australis, Deltoidospora cascadensis, D.
psilostoma, D. nodaensis, Jimboisporites senonicus, Vitreisporites pallidus,
Araucaracites australis, A.· limbatus, Monocolpopollenites kyushuensis,
Arecipites pflugii, Tricolpites vulgaris, and Fibulapollis evanidus.
After all, a similarity of the Uge palynoflora to the Kuji palynoflora is not
so strong. This fact is indicated by 71 new species in the Uge palynoflora.
Miki (1972) reported more than 120 palynomorphs from the Coniacian and
Santonian Futaba Group. 15 species of them appear commonly in the Uge
Member: Todisporites minor, Cicatricosisporites dorogensis, C. australiensis,
Cyathidites minor, Deltoidospora nodaensis, D. cascadensis, Jimboisporites
senonicus, Vitreisporites pallidus, Araucariacites australis, A. limbatus, Cupuliferoidaepollenites ditis, Tricolpites rudis, Monocolpopollenites kyushuensis
and Arecipites pflugii.
Takahashi (1973, 1988) reported and described 115 palynomorphs from the
Futaba Group and 30 species of them are common to the Uge palynoflora:
Deltoidospora psilostoma, Cyathidites australis, C. minor, Triplanosporites
minutulus, Monoleiotriletes gracilis, Camarozonsporites hamulatis, Cicatricosisporites australiensis, Laevigatosporites dehiscens, L. senonicus, L. ovoideus,
L. prominens, Inaperturopollenites dubius, 1. laevigatus, 1. parvus, Psophosphaera pseudotsugoides, Pityosporites aliformis, P. siegburgensis, P. scopulipites, P. microinsignis, Phyllocladidites mawsonii, P. ovatus, Vitreisporites
pallidus, Ephedripites (S.) ellipsoideus, Monocolpopollenites kyushuensis, Cup-
uliferoidaepollenites fallax, C. ditis, Tricolpites retiformis, Cyrillaceaepollenites
minor and Rhoipites minus.
From the above-mentioned comparision between the Uge and Kuji (or
Futaba) palynofloras the authors have to regard the Uge palynoflora as a
representative palynoflora of Japan in the Santonian stage.
In conclusion, many spores and pollen grains occurred in the Uge Member,
have respectively each long range in appearance and only a few ones with
restricted range are effective to palynostratigraphy (see Tab. 2 and 3).
147
Palynomorphs from the Uge Member
Spores:
( 1) Aequitriradites verrucosus (Cookson & Dettmann) Cookson &
Dettmann (pI. 30, fig. 7)
( 2) ? Aequitriradites sp. (pI. 35, fig. 2)
( 3) Appendicisporites aurifer Verbizkaja n. comb. (pI. 41, figs. 3a - b;
pI. 45, fig. 3)
( 4) Appendicisporites cf. bellus Markova n. comb. (pI. 34, fig. 3)
( 5) Appendicisporites distocarinatus Dettmann & Playford (pI. 36, figs.
2a - b; pI. 37, figs. 2a - b; pI. 38, figs. 2a - b; pI. 42, figs. 4a - b; pI.
46, fig. 3)
( 6) Appendicisporites exilioides (Maljavkina) Takahashi (pI. 35, fig. 4)
( 7) Appendicisporites gigantiformis n. sp. (pI. 36, figs. la- b; pI. 37, figs.
la - b; pI. 39, figs. la - b; pI. 43, fig. 3(cf.))
( 8) Appendicisporites kanukaensis n. sp. (pI. 40, figs. 3-4 ; pI. 41, figs.
la-b; pI. 42, figs. 1-2; pI. 43, figs. 1-2; pI. 44, figs. 2a-b; pI. 46,
figs. 1, 3)
( 9) Appendicisporites macrorhyzus Maljavkina n. comb. (pI. 45, fig. 2)
(10) Appendicisporites cf. pseudomacrorhyzus Markova n. comb. (pI. 35,
figs. 3a - b)
(11) Appendicisporites rarius n. sp. (pI. 38, figs. la - b; pI. 45, figs. lab)
(12)
(13)
(14)
(15)
(16)
(17)
(18)
(19)
(20)
(21)
(22)
Appendicisporites sellingii Pocock (pI. 41, fig. 2)
Appendicisporites taneichianus n. sp. (pI. 39, figs. 2a -d; pI. 40, figs.
1-2; pI. 44, figs. 1, 3-4; pI. 46, figs. 4-5)
Appendicisporites sp. (pI. 41, figs. 4a - b)
Baculatisporites giganticus n. sp. (pI. 22, figs. 3-4)
Balmeisporites nipponicus n. sp. (pI. 32, figs. 5-6; pI. 33, figs. 1-4;
pI. 34, figs. la - b; pI. 35, figs. la-b)
? Balmeisporites sp. (pI. 34, fig. 2)
Biretisporites incrassatus Takahashi & Shimono (pI. 11, fig. 4)
Camarozonosporites (Camarozonosporites) similis n. sp. (pI. 28, figs.
5-7; pI. 29, figs. 1-4; pI. 30, figs. 1-2)
Camarozonosporites (C.) semilevis Krutzsch (pI. 30, fig. 4)
Camarozonosporites (Hamulatisporis) hamulatis Krutzsch (pI. 29, figs.
5-8; pI. 30, fig. 5)
Camarozonosporites (H.) insignis Norris (pI. 30, fig. 3)
148
(23)
(24)
K.
TAKAHASHI
& R.
SUGIYAMA
Camarozonosporites (H.) sp. (pI. 28, fig. 4)
Cardioangulina trichacantha Maljavkina (pI. 4, fig. 7; pI. 7, figs. 12)
(25)
(26)
(27)
(28)
(29)
(30)
(31)
(32)
(33)
(34)
(35)
(36)
(37)
(38)
(39)
(40)
(41)
(42)
(43)
(44)
(45)
(46)
(47)
(48)
(49)
(50)
(51)
(52)
(53)
Cibotiidites sp. (pI. 24, figs. 2a - b)
Cicatricosisporites australiensis (Cookson) Potonie (pI. 46, fig. 6)
Cicatricosisporites brevilaesuratus Couper emend. Kemp (pI. 43, figs.
4a-b)
Cicatricosisporites dorogensis Potonie & Gelletich (pI. 45, fig. 5)
Cicatricosisporites cf. hallei Delcourt & Sprumont (pI. 45, fig. 4)
Cicatricosisporites minuticanaliculatus n. sp. (pI. 47, figs. 3 - 4; pI.
48, fig. 1; pI. 49, figs. 2-3)
Cicatricosisporites pseudotertiarius Krutzsch (pI. 42, fig. 3)
Cicatricosisporites senonicus n. sp. (pI. 47, figs. 2,5; pI. 48, figs. 23; pI. 49, fig. 1)
Cicatricosisporites subrotundus Brenner (pI. 47, figs. la-b)
Cingulatisporites iwatensis n. sp. (pI. 23, figs. 2-3)
Clavatisporites sp.(pi. 23, figs. 7a - b)
Cyathidites australis Couper (pI. 8, figs. 6-7; pI. 9, figs. 1-2,5)
Cyathidites minor Couper (pI. 8, figs. 8-10; pI. 9, figs. 3-4.6)
Cyathidites splendens Harris (pI. 2, fig. 1)
Deltoidospora cascadensis Miner (pI. 4, fig. 2)
Deltoidospora diaphana Wilson & Webster (pI. 4, fig. 5(?); pI. 7, fig.
3; pI. 10, figs. 2-3)
Deltoidospora psilostoma Rouse (pI. 4, figs. 3-4)
Deltoidospora nodaensis Miki (pI. 10, fig. 4)
Densoisporites cf. microrugulatus Brenner (pI. 25, fig. 6)
Dictyophyllidites harrisii Couper (pI. 9, figs. 7-9; pI. 10, figs. 7-8)
Endoculeospora cf. delicata Burger (pI. 24, fig. 1)
Extrapunctatospora /ayumensis Takahashi & Jux (pI. 53, fig. 2)
Extrapunctatospora microalveolatus Krutzsch (pI. 49, fig. 4)
Extrapunctatospora sp. (pI. 51, fig. 7)
Foveosporites per/ossus n. sp.(pI. 28, figs. 2-3)
Foveosporites sp. (pI. 27, figs. 4a - b)
Foveotriletes cf. scrobiculatus (Ross ex Weyland & Krieger) Potonie
(pI. 28, fig. 1)
Gleicheniidites cf. con/ossus Hedlund (pI. 14, fig. 3)
Gleicheniidites senonicus Ross (pI. 12, figs. 8-10; pI. 13, figs. 1-10;
pI. 14, figs. 1- 2)
(54)
(55)
(56)
(57)
(58)
(59)
(60)
(61)
(62)
(63)
149
Gleicheniidites verrucatus n. sp. (pI. 23, figs. la - b, 5a - b)
Ischyosporites sp. (pI. 27, fig. 3)
Jimboisporites senonicus Miki (pI. 21, figs. 3a -c; pI. 22, figs. 1- 2)
Laevigatosporites dehiscens Takahashi (pI. 49, figs. 5-11; pI. 51, figs.
8-9; pI. 52, figs. 1-4)
Laevigatosporites longus n. sp. (pI. 50, figs. 8-9)
Laevigatosporites nitidulus n. sp. (pI. 50, figs. 4-7; pI. 51, figs. 1-3)
Laevigatosporites ovoideus Takahashi (pI. 48, fig. 7; pI. 52, fig. 7)
Laevigatosporites probatus Takahashi (pI. 50, fig. 3; pI. 51, fig. 4)
Laevigatosporites prominens Takahashi (pI. 50, figs. 1- 2; pI. 52, figs.
5-6)
Laevigatosporites senonicus Takahashi (pI. 48, figs. 4, 6; pI. 51, fig.
6)
(64)
(65)
(66)
Laevigatosporites sp. (pI. 53, fig. 1)
Latosporites rotundus Takahashi & Jux (pI. 48, fig.5 ; pI. 51, fig. 5)
Leiotriletes cf. convexiformis Chlonova (pI. 1, figs. 6a - b; pI. 3, fig.
6)
(67)
(68)
Leiotriletes giganticus n. sp. (pI. 1, figs. 1-3; pI. 2, figs. 2-4)
Leiotriletes maxoides Krutzsch maximus (Pflug) Krutzsch (pI. 4, fig.
1)
(69)
(70)
(71)
(72)
(73)
Leiotriletes rotundiformis (Maljavkina) Chlonova (pI. 1, figs. 4-5;
pI. 2, fig. 5(cf.))
Leiotriletes wolffi Krutzsch wolffi (pI. 3, fig. 4(cf.); pI. 10, fig. 1)
?Leiotriletes sp. (pI. 11, fig. 2)
Leptolepidites sp. (pI. 23, fig. 4)
Lycopodiacidites circularis n. sp. (pI. 26, figs. 6a - b; pI. 30, figs. 6ab)
(74)
(75)
(76)
(77)
(78)
(79)
(80)
Lygodiidites lal;vigatus Pocock (pI. 32, fig. 4)
Lygodiidites tohokuensis n. sp. (pI. 31, figs. 1-2; pI. 32, figs. 1-3)
Monoleiotriletes gracilis Krutzsch (pI. 2, figs. 6a - b; pI. 3, fig. 7; pI.
5, figs. 3-5; pI. 10, fig. 5)
Monoleiotriletes grandissimus n. sp. (pI. 4, fig. 6; pI. 5, figs. 1- 2;
pI. 6, fig. 1; pI. 10, fig. 6)
Monoleiotriletes minimus Krutzsch (pI. 8, figs. 1-2)
Patellasporites polyverrucifer n. sp. (pI. 19, figs. 2-5; pI. 20, figs. 14; pI. 21, figs. la - b)
Patellasporites verrucatus n. sp. (pI. 15, figs. 4-5; pI. 18, figs. 4-5;
pI. 21, figs. 2a - b)
150
(81)
(82)
(83)
(84)
(85)
(86)
(87)
(88)
(89)
(90)
(91)
(92)
(93)
(94)
(95)
(96)
(97)
(98)
(99)
K.
TAKAHASHI
& R.
SUGIYAMA
Patellasporites sp. (pI. 18, figs. 2a - b)
"Perinomonoletes" tropicalis Salard (pI. 53, fig. 10)
Punctatisporites granulatus n. sp. (pI. 22, figs. 6-7; pI. 23, fig. 6)
Punctatosporites sp. (pI. 53, fig. 3)
? Punctatosporites sp. (pI. 53, figs. 6-7)
Retitriletes d. saltimaniolus Srivastava (pI. 25, figs. 7a - b)
Retitriletes sp. (pI. 27, figs. 2a - b)
Rugulatisporites sp. (pI. 25, fig. 4)
Saxosporis sp. (pI. 22, fig. 5)
Todisporites grandi/ormis n. sp. (pI. 5, fig. 6; pI. 6, figs. 2-3)
Todisporites cf. minor Couper (pI. 3, fig. 5)
Toroisporis (Duplotoroisporis) triangulus n. sp. (pI. 12, figs. 3-7)
Trachysporites microverrucatus n. sp. (pI. 24, figs. 4-5; pI. 25, figs.
1-3)
Trachysporites sp. (pI. 24, fig. 3)
Trilites pulchellus n.sp.(pi. 14, figs. 5-7; pI. 15, figs. 1-2; pI. 16,
figs. 1-3; pI. 17, figs. 1-8)
Trilites pustulosus n. sp. (pI. 15, figs. 3a - b; pI. 16, figs. 4-5; pI. 18,
fig. 1)
Trilites sp. (pI. 18, fig. 3)
? Trilites sp. (pI. 19, figs. 1a - b)
Trilobosporites (Trilobosporites) cr. weylandii Doring (pI. 11, figs. 5ab)
(100)
Trilobosporites (Tuberosisporites) sp. (pI. 25, fig. 5)
(101) ? Trilobosporites sp. (pI. 14, fig. 4)
(l02) Triplanosporites giganteus n. sp. (pI. 3, figs. 1-3)
(l03) Triplanosporites microsmuosus pflanzl (pI. 7, figs. 8-9; pI. 8, fig.
5)
(104)
(l05)
(l06)
(l07)
(l08)
(l09)
(lID)
(l11)
(112)
Triplanosporites minutulus Takahashi (pI. 8, figs. 3-4)
Triplanosporites rikuchuensis n. sp. (pI. 7, figs. 4-7)
Triplanosporites taneichiensis n. sp. (pI. 5, figs. 7-8)
Triplanosporites varius n. sp. (pI. 6, figs. 4-7)
Undulatisporites flexuosus n. sp. (pI. 12, figs. 1- 2)
Undulatisporites d. pseudo brasiliensis Krutzsch (pI. 11, figs. 7a - b)
Undulatisporites sp. (pI. 11, fig. 1)
? Undulatisporites sp. (pI. 11, fig. 3)
Verrucatosporites verruculosus n. sp. (pI. 53, figs. 5.9; pI. 54. figs.
1-4)
(113)
(114)
(115)
151
Verrucatosporites sp. a (pI. 53, fig. 4)
Verrucatosporites sp. b (pI. 53, fig. 8)
Zlivisporis novomexicanus (Anderson) Leffingwell (pI. 26, figs. 1- 5;
pl. 27, fig. 1)
Gymnospermic pollen:
(116) AbiespoUenites cL microsaccoides Krutzsch (pI. 68, fig. 3(cL))
(117) AbiespoUenites minusn. sp. (pI. 66, fig. l;pI. 67, figs. 1, 2(cL),7;pI.
69, fig. 2; pI. 70, fig. 3)
(118) AbiespoUenites sp. (pI. 69, fig. 1)
(119) ? Abiespollenites sp. (pI. 65, fig. 7)
(120) Alisporites bilateralis Rouse (pI. 63, fig. 4)
(121) Alisporites bisaccus Rouse (pI. 60, figs. 1- 2;pl, 61, figs. 1a - b)
(122) Alisporites enormis n. sp. (pI. 60, fig. 4; pI. 66, fig. 2)
(123) Araucariacites australis Cookson ex Couper (pI. 57, figs. 2, 4-6; pI.
58, figs. 1-5)
(124) Araucariacites limbatus (Balme) Habib (pI. 56, fig. 12; pI. 57, figs. 3,
7)
(125)
(126)
(127)
(128)
(129)
(130)
(131)
(132)
(133)
(134)
(135)
(136)
(137)
(138)
?Araucariacites sp. (pI. 58, fig. 6)
Callialasporites ugensis n. sp. (pI. 56, figs. 10-11)
Cedripites medius (Zauer) Krutzsch (pI. 69, fig. 4; pI. 70, figs. 2, 4)
Cedripites microsaccoides Song & Zheng (pI. 69, fig. 3; pI. 70, fig. 1)
Cedripites sanrikuensis n. sp. (pI. 62, figs. 4-5)
Classopollis grandissimus n. sp. (pI. 79, figs. 4-8)
Classopollis taneichiensis n. sp. (pI. 78, figs. 2-8; pI. 79, figs. 1-3, 9)
Cupressacites cuspidatae/ormis (Zaklinskaja) Krutzsch (pI. 54, fig. 11;
pI. 56, fig. 6; pI. 59, fig. 9(cf.))
Cupressacites insulipapillatus (Trevisan) Krutzsch (pI. 54, figs. 9-10,
12; pI. 55, figs. 1-2, 3(cf.))
Cycadopites mirus n. sp. (pI. 86, figs. 6-7, 9-11)
Cycadopites sp. a (pI. 86, fig. 8; pI. 98, fig. 2)
Cycadopites sp. b (pI. 86, fig. 13)
Cycadopites sp. c (pI. 98, fig. 1)
Ephedripites (Distachyapites) scabridus Song & Zheng (pI. 83, figs. 12)
(139)
(140)
(141)
Ephedripites (D.) sp. (pI. 83, fig. 3)
Ephedripites (Ephedripites) chaloneri Brenner (pI. 83, fig. 12)
Ephedripites (E.) notensis (Cookson) Krutzsch (pI. 83, figs. 7-11)
152
(142)
(143)
(144)
(145)
(146)
(147)
(148)
(149)
(150)
(151)
(152)
(153)
(154)
(155)
(156)
(157)
(158)
(159)
(160)
(161)
(162)
(163)
(164)
(165)
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(170)
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K.
TAKAHASHI
& R.
SUGIYAMA
Ephedripites (E.) cf. regularis Hoeken-Klinkenberg (pI. 83, fig. 6)
Ephedripites (E.) sp. a (pI. 83, fig. 4)
Ephedripites (E.) sp. b (pI. 83, fig. 5)
Ephedripites (8piralipites) cf. contaxtus Gao & Zhao n. comb. (pI. 83,
fig. 15)
Ephedripites (8.) ellipsoideus Takahashi (pI. 84, fig. 6)
Ephedripites (8.) elongatus n. sp. (pI. 84, figs. 7-9)
Ephedripites (8.) longus Song & Zheng n. comb. (pI. 84, figs. 4-5)
Ephedripites (8.) perlatus Wang & Zhao n. comb.( pI. 83, figs. 13-14)
Ephedripites (8.) praeclarus Chlonova n. comb. (pI. 84, figs. la- b, 3a b(ef.) )
Ephedripites (8.) sp. aff. E. (8.) longus Song & Zheng n. comb. (pI. 84,
fig. 10)
Ephedripites (8.) sp. a (pI. 83, fig. 16)
Ephedripites (8.) sp. b (pI. 84, fig. 2)
Inaperturopollenites dubius (Potonie & Venitz) Thomson & Pflug (pI.
54, figs. 13-14; pI. 55, figs. 7-10; pI. 59, figs. 3-6)
Inaperturopollenites laevigatus Takahashi (pI. 54, figs. 7-8)
Inaperturopollenites parvus Takahashi (pI. 54, figs. 5 -6)
Inaperturopollenites rugatus n. sp. (pI. 55, figs. 12 -13)
Inaperturopollenites sp. (pI. 56, fig. 1)
? Microcachryidites sp. (pI. 75, fig. 8)
Phyllocladidites globulosus n. sp. (pI. 74, figs. 3-5; pI. 75, fig. 4)
Phyllocladidites mawsonii Cookson (pI. 75, fig. 7;pI. 76, figs. 6-9)
Phyllocladidites cr. ovatus Takahashi (pI. 69, fig. 16; pI. 70, figs. 7ab; pI. 75, figs. 5-6)
Piceapollis grandiformis n. sp. (pI. 67, fig. 3;pI. 70, fig. 6)
Piceapollis misellus n. sp. (pI. 67, figs. 4-6; pI. 68, figs. 1-2)
Piceapollis praemarianus Krutzsch (pI. 64, fig. 3)
Piceapollis sp. a (pI. 69, fig. 5)
Piceapollis sp. b (pI. 77, fig. 2)
? Piceapollis sp. (pI. 77, fig. 3)
Pityosporites alatipollenites Rouse n. comb. (pI. 62, figs. 1-3, 6 -7;
pI. 76, figs. 1-4; pI. 77, fig. 1)
Pityosporites aliformis Takahashi (pI. 60, fig. 3)
Pityosporites cedrisacciformis Krutzsch (pI. 64, fig. 2)
Pityosporites ef. constrictus Singh (pI. 61, fig. 2)
Pityosporites cretaceus n. sp. (pI. 63, figs. 1-3, 7; pI. 65, fig. 2)
153
(174) Pityosporites macroinsignis Krutzsch (pI. 64, fig. 1)
(175) Pityosporites microsibiricus Zak1inskaja n. comb. (pI. 61, fig. 3)
(176) Pityosporites minutus (Zaklinskaja) Krutzsch (pI. 61, fig. 8; pI. 63,
figs. 5-6)
(177)
(178)
Pityosporites d. pini/ormis Zaklinskaja n. comb. (pI. 65, fig. 6)
Pityosporites scopulipites (Wodehouse) Krutzsch (pI. 63, fig. 8; pI.
76, fig. 5)
(179)
(180)
(181)
(182)
(183)
(184)
(185)
(186)
(187)
(188)
Pityosporites siegburgensis Takahashi & Jux (pI. 65, fig. 5; pl. 66,
figs. 5, 7; pl. 77, figs. 4-5, 7-8)
Pityosporites sp. a (pI. 61, fig. 7)
Pityosporites sp. b (pI. 62, fig. 8)
Pityosporites sp. c (pI. 65, fig. 1)
Pityosporites sp. d (pI. 70, fig. 5; pI. 73, fig. 7)
Podocarpidites bi/ormis Rouse (pI. 68, figs. 4a - b)
Podocarpidites cf. ellipticus Cookson (pI. 68, fig. 5)
Podocarpidites embryonalis Krutzsch (pI. 61, fig. 6)
Podocarpidites piniverrucatus Krutzsch (pI. 66, fig. 4)
Podocarpidites cf. podocarpoides (Thiergart) Krutzsch (pI. 73, figs.
6a-b)
(189) Podocarpidites senonicus n. sp.(pI. 64, figs. 4-5)
(190) PristinuspoUenites microsaccus (Couper) B. D. Tschudy (pI. 68, fig. 6;
pI. 71, figs. 2-5; pI. 72, figs. 1-3)
(191) Pristinuspollenites sp.(pi. 71, fig. 1)
(192) Psophosphaera aggereloides (Ma1javkina) Ch1onova (pI. 55, fig. 6; pI.
58, fig. 8)
(193)
(194)
(195)
(196)
Psophosphaera pseudotsugoides Krutzsch (pI. 55, figs. 4-5; pI. 57,
fig. 1; pI. 59, figs. 1- 2)
Psophosphaera sp.(pI. 58, fig. 7)
Rugubivesiculites japonicus n. sp. (pI. 72, figs. 4-5; pI. 73, figs. 15; pI. 75, fig. 1; pI. 78, figs. la-b)
Rugubivesiculites sphaericus n. sp. (pI. 74, figs. 1- 2; pI. 75, figs. 23)
(197) Sciadopityspollenites antiquus Krutzsch (pI. 56, fig. 9)
(198) SciadopityspoUenites quintus Krutzsch (pI. 56, fig. 8)
(199) Sciadopityspollenites tuberculatus (Zak1inskaja) Krutzsch (pI. 56, fig.
7)
(200) Sequoiapollenites gracilis Krutzsch (pI. 56, figs. 3-4; pI. 59, fig. 7)
(201) Sequoiapollenites sp. a (pI. 56, fig. 2)
154
K.
TAKAHASHI
& R.
SUGIYAMA
(202) Sequoiapollenites sp. b (pI. 56, fig. 5)
(203) Sequoiapollenites sp. c (pI. 59, fig. 8)
(204) ? Sequoiapollenites sp. (pI. 55, figs. lla - b)
(205) Vitreisporites pallidus (Reissinger) Nilsson (pI. 60, figs. 5-6; pI, 61,
figs. 4-5)
Gymnosperm - angiosperm incertae sedis:
(206) Clavatipollenites variabilis n. sp. (pI. 80, figs. 1-8; pI. 81, figs. I ll; pI. 82, figs. 1-5)
(207) Clavainaperturites sp. ,a (pI. 97, figs. 7)
(208) Clavainaperturites sp. b (pI. 97, figs. 8)
Angiospermic pollen:
(209) Arecipites pjlugii (Takahashi) Krutzsch (pI. 86, fig. 16)
(210) Arecipites sp. a (pI. 86, fig. 14)
(211) Arecipites sp. b (pI. 86, fig. 15)
(212) Arecipites sp. c (pI. 87, fig. 1)
(213) Arecipites sp. d (pI. 87, fig. 2)
(214) Arecipites sp. e (pI. 87, figs. 3-4)
(215) Artiopollis cf. indivisus Agasie (pI. 95, figs. 5a -c)
(216) Asteropollis clavatus (Phillips & Felix) Ward (pI. 82, figs. 6-7)
(217) Callistopollenites radiatostriatus (Mtehedlishivili) Srivastava (pI. 88,
figs. 27a - b)
(218) Cricotriporites sp. (pI. 95, fig. 8)
(219) Cupulijeroidaepollenites ditis (Takahashi) Takahashi (pI. 87, fig. 10;
pI. 98, fig. 7)
(220) Cupulijeroidaepollenites jallax (Potonie) Potonie (pI. 87, figs. 20 - 25;
pI. 98, figs. 4, 8-10)
(221) Cupulijeroidaepollenites lanceolatus n. sp. (pI. 87, figs. 26-28; pI. 98,
fig. ll(eL))
(222) Cupulijeroidaepollenites vulgaris (Takahashi) Takahashi (pI. 87, figs.
11-14; pI. 88, figs. 16-18; pI. 97, figs. 9-10)
(223) Cyrillaceaepollenites minor (Takahashi) Takahashi (pI. 94, figs. 6-8)
(224) Fibulapollis enodatus (Chlonova) Takahashi (pI. 85, figs. 8-9; pI. 86,
figs. 1-5)
(225) Fibulapollis evanidus (Chlonova) Takahashi (pI. 85, figs. 1-7)
(226) Foveotricolpites concinnus Singh (pI. 91, fig. 16)
(227) Foveotricolpites fastidiosus n. sp. (pI. 93, fig.6; pI. 94, figs. 1a - b)
(228)
(229)
(230)
(231)
155
Foveotricolpites globosus n. sp. (pI. 92, figs. 1a - b, 5, 6a - b; pI. 93,
figs. 4-5)
Foveotricolpites sp. a (pI. 89, fig. 6)
Foveotricolpites sp. b (pI. 100, fig. 11)
Foveotricolporites cr. elegantulus Takahashi & Jux (pI. 94, figs. 17ab)
(232) Foveotricolporites gloriosus n. sp. (pI. 95, figs. 1- 2)
(233) Foveotricolporites grandi/ormis n. sp. (pI. 95, figs. 3-4)
(234) ? Foveotricolporites sp. (pI. 98, fig. 13)
(235) Gothanipollis sp. (pI. 96, figs. 1-2)
(236) Graminidites cf. laevigatus Krutzsch (pI. 95, fig. 7)
(237) Graminidites sp. (pI. 95, fig. 6)
(238) Hammenia sp. (pI. 91, fig. 1a)
(239) Ilexpollenites claviger Takahashi n. comb. (pI. 93, fig. 9)
(240) Ilexpollenites miniclavatus n. sp. (pI. 93, figs. 7-8)
(241) Ilexpollenites minusn. sp. (pI. 93, figs. 10-11)
(242) Intrabaculitricolporites consularis (Takahashi) Takahashi consularis
(pI. 94, fig. 2)
(243) Monocolpopollenites kyushuensis Takahashi (pI. 86, fig. 12)
(244) Nyssapollenites pseudocruciatus (Potonie) Thiergart (pI. 94, figs. 4ab)
(245)
(246)
(247)
(248)
(249)
(250)
(251)
(252)
(253)
(254)
(255)
(256)
(257)
(258)
Phimopollenites pannosus (Dettmann & Playford) Dettmann (pI. 91,
figs. 1b, 2)
Phimopollenites sp. (pI. 87, fig. 19)
Potamogetonacidites senonicus n. sp. (pI. 79, figs. 10-12)
Quercoidites cf. henrici (Potonie) Potonie (pI. 87, fig. 17)
Quercoidites umiensis (Takahashi) Takahashi (pI. 87, figs. 5 - 9, 15;
pI. 98, fig. 3)
Retitrescolpites pseudoazemae n. sp. (pI. 89, figs. 9-10)
Rhoipites kitakamiensis n. sp. (pI. 94, figs. 12-16; pI. 101, figs. 12, 7)
Rhoipites minus Takahashi & Jux (pI. 94, figs. lla - b)
? Rhoipites sp. (pI. 94, fig. 9)
Rousea elegantula n. sp. (pI. 91, figs. lla - b)
Rousea cr. prosimilis (Norris) Srivastava (pI. 89, figs. 5a - b)
Rousea reticosa n. sp. (pI. 89, figs. 1-3)
Rousea triangulata n. sp. (pI. 91, figs. 12-15; pI. 92, figs. 4a - b)
Rousea ugensis n. sp. (pI. 92, figs. 2-3)
156
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(260)
(261)
(262)
(263)
(264)
(265)
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TAKAHASHI
& R.
SUGIYAMA
Rousea sp.(pI. 93, figs.3a-b)
? Rousea sp. (pI. 101, fig. 4)
Satishia cf. compacta (Norton) Ward (pI. 90, fig. 22)
Satishia glyceia Ward (pI. 89, figs. 8a - b)
Satishia pomposa n. sp. (pI. 93, figs. 1-2)
Satishia triformis n. sp. (pI. 90, figs. 18-19)
Satishia uniformis n. sp. (pI. 99, fig. 5; pI. 100, figs. 5-7)
Satishia sp. a (pI. 90, fig. 20)
Satishia sp. b (pI. 90, fig. 23)
Satishia sp. c (pI. 100,. fig. 9)
Satishia sp. d (pI. 100, fig. 10)
Striatopollis striatellus (Takahashi) Takahashi (pI. 87, fig. 16)
Subtriporopollenites kyushuensis Takahashi (pI. 95, fig. 9)
Symplocacites micropunctatus n. sp. (pI. 96, figs. 3-8; pI. 101, fig. 8)
Symplocacites microreticulatus n. sp. (pI. 96, figs. 10-11; pI. 97, figs.
1-4)
? Symplocacites sp. (pI. 96, figs. 9a - b)
Tricolpites ellipsoideus n. sp. (pI. 90, figs. 13-17; pI. 91, figs. 3-4,
6-9; pI. 99, figs. 1-4)
Tricolpites ellipticus Takahashi & Jux (pI. 91, fig. 5)
Tricolpites nemejcii Pacltova (pI. 89, figs. 4a - b; pI. 90, fig. 2l(cf.))
Tricolpites ouiformis n. sp. (pI. 89, figs. 11-15; pI. 101, figs. 3, 5-6)
Tricolpites retiformis (Pflug & Thomson) Takahashi & Jux (pI. 88,
figs. 19-21, 23)
Tricolpites sphaeroides n.sp.(pI. 90, figs. 2-12; pI. 99, figs. 6-12;
pI. 100, fig. 8)
Tricolpites rudis Takahashi n. comb. (pI. 91, fig. 17; pI. 92, fig. 7)
Tricolpites uulgaris (Pierce) Srivastava (pI. 90, fig. 1)
Tricolpites sp. (pI. 89, fig. 16)
? Tricolpites sp. (pI. 91, fig. 10)
Tricolpopollenites asper Pflug & Thomson (pI. 88, figs. 3a - b, 12, 15;
pI. 98, fig. 5)
Tricolpopollenites cf. augustinensis Takahashi & Jux (pI. 88, fig. 24)
Tricolpopollenites baculatus n. sp. (pI. 88, figs. 25-26)
Tricolpopollenites chikushiensis Takahashi grandiformis Takahashi
(pI. 88, figs. 1- 2)
Tricolpopollenites cf. pseudoeuphorii Pflug (pI. 88, fig. 22)
Tricolpopollenites subasper Takahashi (pI. 88, figs. 4-11, 13-14; pI.
(291)
(292)
(293)
157
98, figs. 6, 12)
Tricolporopollenites sp. a (pI. 94, fig. 3)
Tricolporopollenites sp. b (pI. 94, figs. 5a - b)
Tricolporopollenites sp. c (pI. 94, fig. 10)
Phytoplankton:
(294) Ascodinium sp. (pI. 102, fig. 8)
(295) Cymatiosphaera reticulosa Takahashi (pI. 102, figs. 5a - b)
(296) Cymatiosphaera sp. a (pI. 102, figs. 4a - b)
(297) Cymatiosphaera sp. b (pI. 102, figs. 6-7)
(298) Pterospermella cf. aureolata (Cookson & Eisenack) Eisenack (pI. 102,
fig. 1)
(299) Pterospermella cf. australiensis (Deflandre & Cookson) Eisenack (pI.
102, fig. 3)
(300) Pterospermella sp. (pI. 102, fig. 2)
Incertae sedis:
(301) ? Ovoidites sp. (pI. 87, figs. 29a - b)
(302) Indeterminable monolete spore (?) (pI. 97, fig. 5)
(303) Indeterminable inaperturate pollen with roughly rugulate sculpture
(pI. 97, figs. 6a - b)
Systematic description
The genera and species described here are arranged alphabetically under
the broad heading of spores, gymnospermic pollen, gymnosperm-angiosperm
incertae sedis, angiospermic pollen, phytoplankton, and incertae sedis.
Spores
Genus Aequitriradites Delcourt & Sprumont 1955 emend. Cookson
& Dettmann 1961.
Type species:Aequitriradites dubius Delcourt & Sprumont 1955.
Aequitriradites uerrucosus (Cookson & Dettmann) Cookson & Dettmann
PI. 3D, fig. 7.
158
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& R.
SUGIYAMA
1958 Cirratriradites verrucosus Cookson & Dettmann, Prec. Roy. Soc. Victoria, N. S., 70,
pt. 2, p. 112, pl. 18, figs. 2-6.
1959 Cirratriradites verrucosus Cookson & Dettmann, Bolchovitina,l29, pI. 8, fig. 118.
1961 Cirratriradites verrucosus Cookson & Dettmann, Chlonova, Trudy Inst. Geol. Geophys., Acad. Sci. USSR, Siberian Br., 7, pp. 52-53, pI. 5, figs. 35, 35a.
1961 Aequitriradites verrucosus (Cookson & Dettmann) Cookson & Dettmann, Palaeontology, 4, pt. 3, p. 427, pI. 52, figs. 1- 6
1969 Aequitriradites verrucosus (Cookson et Dettmann) Cookson et Dettmann, Chlonova,
Trudy Inst. GeoI. Geophys., Acad. Sci. USSR, Siberian Br., 91, p. 53, pI. 7, fig. 6.
1972 Aequitriradites verrucosus (Cooskon & Dettmann) Cookson & Dettmann 1961, Srivastava, Palaeontographica, B, 139, Lfg. 1-4, p. 4, pI. 3, figs. 1-4.
1980 Aequitriradites verrucosus (Cookson & Dettmann) Cookson & Dettmann, Burger,
BMR Bull., 189, p. 62, pl. 16, figs. 3-4.
Description: See Cookson & Dettmann (1958).
Measurements: Overall equatorial diameter 65 - 98 /.lm, equatorial diameter
of spore body 48 -70 /.lm (Cookson & Dettmann, 1958); overall diameter 60(66) -75 /.lm, body 48-(49)-54 /.lm (Chlonova, 1961); overall diameter 60 -77
/.lm, body 51-58 /.lm (Chlonova, 1969); equatorial diameter of spore 40 - 70
/.lm, equatorial diameter of central cavity 34-60 /.lm (Burger, 1980); present
specimen: equatorial diameter of spore body 68 X 65 /.lm.
Occurrence: Uge Member, south of Kanuka (C 31).
Previous records: Lower Cretaceoua (Aptian-Albian), south Australia, Victoria, New South Wales, and Queensland, Australia (Cookson & Dettmann, 1958);
Upper Cretaceous, Viliuisk basin, USSR (Bolchovitina, 1969); Upper Cretaceous
in eastern area of the western Siberian Lowland (Chlonova,1961, 1969); Wealden, northern Germany (Doring, 1964); Cretaceous, Santa Cruz, Argentina
(Archangelsky & Gamerro, 1965); Maastrichtian, Alberta, Canada (Srivastava,
1972); Lower Cretaceous (Aptian-Albian), Surat basin, Queensland and New
South Wales, Australia (Burger, 1980).
Remarks: Three species of triradiate microsporess with equatorial flange
were referred by Cookson & Dettmann (1958) to the Palaeozoic form-genus
Cirratriradites Willson & Coe (1940) : C. spinulosus, C. tilchaensis, and C. verr
ucosus. After that, they (1961) recognized their close agreement with the Cr
etaceous form-genus Aequitriradites Delcourt & Sprumont (1955) and then
emended it. Recently recognizable species of Aequtriradites are as follows:
Aequtriradites dubius Delcourt & Sprumont, A. inconspicuus Delcourt & Sprumont, A. spinulosusCCookson & Dettmann)Cookson & Dettmann, A. tilchaensis
(Cookson & Dettmann) Cookson & Dettmann, A. verrucosus CCookson &
159
Dettmann) Cookson & Dettmann A. salebrosaceus (Maljavkina) Nilsson, A.
interruptus(Bolchovitina) Cookson & Dettmann, and A. infrapunctatus Lantz.
Botanical affinity: Unknown.
?Aequitriradites sp.
PI. 35, fig. 2.
Description: Trilete (?) spore, with a broken membranous zona. Exine 1. 5
ttm thick, with small echini (1-2 ttm high) and bacula (2.2-3 ttm high).
Proximal Y mark indistinct.
Measurements: Equatorial diameter of spore body 60 X 55 ttm.
Remarks: A single specimen was observed. It is not clear, whether this belongs
to A. spinulosus or not, because the specimen is in incomplete preservation.
Genus Appendicisporites Weyland & Krieger 1953.
Type species: Appendicisporites tricuspidatus Weyland & Greifeld 1953.
Remark: The form-genus Appendicisporites Weyland & Krieger (1953) is different from the form-genus Cicatricosisporites R. Potonie & Gelletich 1933 emend. R. Potonie 1966 in having apical exinous horn-like thickenings (appendices), and from the form-genus Costatoperforosporites Deak (1962) in having
apical appendices and in lacking perforate ribs.
Srivastava (1975) stated that the genus Plicatella Maljavkina (1949) has
priority over Appendicisporites Weyland & Krieger (1953), according to Ariele
37 of the ICBN, but is abondoned here because its original circumscription is
ambiguous and the species are poorly described and illustrated.
However, type species of the genus Plicatella has oblong appendices
('caps') in polar view and that of the ginus Appendicisporites possesses very
long ones. Potonie (1960, p. 50) described in his synopsis as follows:"
,
but with appendices on the corner shorther than in Appendicisporites,
"
Appendicisporites can be maintained as a separate form-genus only on the
basis of the longer appendices; possibly this is not sufficient justification
(Jansonius & Hills, 1976, genera file). Accordingly, it is very difficult to distinguish between Plicatella and Appendicisporites. However, the genus Appendicisporites is adopted here, according to Srivastava's opinion (Srivastava,
1975).
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Appendicisporites aurifer Verbizkaja n. comb.
PI. 41, figs. 3a - b; pI. 45, fig. 3.
1958 Anemia aurifer Verbizkaja.
1961 Anemia aurifer Verbizkaja,
p. 59, pI. 18, figs. 2a-c.
Trudy Lab. GeoI. Acad. Sic., USSR, pI. 2, fig. 38.
Bolchovitina, Trudy Geol.Inst., Acad. Sci. USSR, 40,
Diagnosis: Trilete spore; amb triangular with convex sides and apical hornlike appendices at the three corners (11 -12 J.1m long). Trilete rays nearly
reach equator. Endexine thin and smooth; ectexine thick, ornamented on the
proximal surface by nine ribs, 2-3 J.1m wide, spaced 0.5-1 J.1m apart, running parallel to the sides of the spore. Distal ribs running parallel to the
sides except in a triangle area (16 X 10 J.1m) across over the distal pole.
Measurements: Present specimen: 75-83 J.1m X 60 /lID in approximately equatorial diameter; 67 J.1m in height.
Previous records: Cretaceous (Cenomanian), Suchan basin and Lower Cretaceous (Barremian-Aptian), Suifun basin, Primorskii krai (USSR)(Verbizkaja,
1958; Bolchovitina, 1961).
Remarks: Two specimens were observed. They resemble colsely Appendicisporites (al. Anemia) aurifer Verbizkaja n. comb. from the Maritime Province
of Siberia (Primorskii krai), USSR.
Botanical affinity: Schizaeaceae, Anemia.
Appendicisporites cr. bellus Markova n. comb.
PI. 34, fig. 3.
1961 Anemia bella Markova,
Samoilovitch et aI.. VNIGRI, 177, p. 70, pI. 17, figs.
Sa-b.
Diagnosis: Trilete spore. Amb triangular with convex sides and slightly projected apices (2.5 J.1m long). Trilete laesurae distinct, straight, almost reaching corners. Endexine thin and smooth; ectexine thick, ornamented on the
proximal surface by two (?) ribs, ca. 5 J.1m wide, spaced ca. 1 J.1m apart, running parallel to the sides of the spore; three distal ribs also running parallel
to the sides except in the innermost triangle area of the distal pole, ca. 5 - 9
J.1m wide.
Measurements:Diameter45.4 J.1m, number of ribs 2-3, width of ribs 4-5 J.1m
161
(Samoilovitch et al., 1961); present specimen: 65 X 58 /-lm in equatorial diameter, ribs 5-9 /-lm in width.
Occurrence:Uge Member, south of Kanuka (C 31).
Previous record: Lower Cretaceous (Aptian-Albian), Omsk province, Siberia
(USSR)(Samoilovitch et aI., 1961).
Remark: The present specimen is resemble Appendicisporites (al. Anemia) bellus Markova n. comb. in Samoilovitch et ai. (1961) from the Cretaceous of
Omsk Province, Siberia, but the Uge specimen is somewhat larger in size.
Appendicisporites distocarinatus Dettmann & Playford
PI. 36, figs. 2a - b; pI. 37, figs. 2a - b; pI. 38, figs.
2a - b; pI. 42, figs. 4a - b;46, fig. 3.
1986 Appendicisporites distocarinatus Dettmann & Palyford, Proc. Roy. Soc. Victoria,
81, no. 2, p. 75, pl. 6, figs. 16-18.
1976 Appendicisporites distocarinatus Dettmann & Playford, Burger, BMR Bull., 151,
pp. 124-125, pI. 20, figs. 7a-b;pl. 21, figs. 1a-b,4.
Description: See Dettmann & Plalyford (1968) and Burger (1976).
Measurements: Equatorial diameter 38-(56)-82 /-lm (Burger, 1976);present
specimens: (40) 75-90 /-lm in eqatorial diameter, 43 /-lm in height, appendices
7-13 /-lm long.
Occurrence: Uge Member, south of Kanuka (C 31) and north of Uge station
(C 33, C15).
Previous records: Lower Cretaceous (late Albian), eastern Australia (Dettmann & Playford, 1968); early Late Cretaceous (Cenomanian), southeastern
Australia (Dettmann & Playford, 1969); Upper Cretaceous (Cenomanian-Turonian), Bathurst Island, Northern Territory, Australia (Burger, 1976).
Remarks: The present specimens from the Uge Member are quite identified
with Appendicisporites distocarinatus Dettmann & Playford from the Cretaceous (Albia~-Turonian)of Australia. This species is first reported in Japan.
Appendicisporites exilioides (Maljavkina) Takahashi
PI. 35, fig. 4.
1949 Plicatella trichacantha
~
exilijormis Maljavkina,
Trudy VNIGRI, n. s., 33, p. 61,
162
1953
1961
1961
1982
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pI. 12, fig. 2.
Anemia exilioides Bolchovitina, Trudy Geoi. lnst., Acad. Sic. USSR, 145, Geoi. Ser.
61, p. 37, pI. 4, figs. 7- 8.
Anemia exilioides CMaljavkina) Bolchovitina, Trudy Geoi. lnst. Acad. Sci. USSR,
40, p. 51, pI .14, figs. 2a-b; pI. 17, figs. la-d; pI. 40, fig. 3.
Anemia exilioides CMaljavkina) Bolchovitina f. sibirica Chlonova, Chlonova, Trudy
lnst. GeoI. Geophys., Acad.Sci. USSR, Siberian Br., 3, pp. 22-23, pI. 2, figs. 1517.
Appendicisporites exilioides CBolchovitina) Takahashi, Takahashi & Shimono, Bull.
Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 22. no. 2, pp. 29-30, pI. 3, fig. 11.
Description: See Maljavkina (1949) and Bolchovitina (1953, 1961).
Measurements: Diameter 100 /.lm (Maljavkina, 1949); diameter 50 - (55) - 65
pm (Bolchovitina, 1961); diameter 30. 5-(57)-65 pm (Chlonova, 1961); diameter 60 pm (Takahashi in Takahashi & Shimono, 1982); present specimen: 67 X
55 pm in equatorial diameter.
Occurrence: Uge Member, south of Kanuka(C 31).
Previous records: Cretaceous (Albian-Cenomanian), western Siberia (USSR)
(Maljavkina, 1949); lower Cretaceous, Crimea; Aptian, Russian platform; Albian Kazakhstan and central Ural; Cenomanin-Turonian, Yakut (Bolchovitina,
1953, 1961); Cenomanian-Turonian and Danian-lower Palaeogene, ChulymEniseisk, Siberia (USSR) (Chlonova, 1961); Maastrichtian, Hida district (Japan)
(Takahashi in Takahashi & Shimono,1982).
Appendicisporites gigantijormis n. sp.
PI. 36, figs. 1a - b; pI. 37, figs. 1a - b;
pI. 39, figs. 1a - b; pI. 43, figs. 3 (cf.).
Description: Trilete spores. Equatorial outline triangular with convex sides
and slightly rounded angles. Trilete laesurae long, extending two-thirds or
three-quarters to equatorial margin, sometimes enclosed within thin elevated
lips, borderd by 5 - 7 pm wide margo (see pI. 36, fig. 1a); margo smooth,
broader in polar area than in the apical areas. Exine thick, canaliculate;
ribs 4-6 in number, ca. 5 -12 pm wide separated by 1 /.lm + wide canals on
the proximal side and by ca. 4-8 pm wide canals on the distal side; appendices
on the radial areas, smooth, regular crest-like in appearance in lateral view,
ca. 10-15 pm long.
Proximal sculpture consists of ca. 2-4 ribs parallel to the side of the
163
amb in each inter-radial area; distally 2-4 ribs running parallel to the side of
the amb; ribs on the distal pole forming a simple triangle area. Three or
four inner ribs coalesce with margo and outer equatorial ribs coalesce with
adjacent ribs forming appendices on the radial areas, radial thickening on the
proximal surface gradually increase towards the equator.
Measurements: 90-117 J,J.m X 90-107.5 J.Lm in approximately equatorial diameter (four specimens), 53 J,J.m in height (one specimen).
(A)(C 33).
Holotype: PI. 37, figs. la - b; 107.5 X 100 J.Lm in approximately equatorial
diameter; ribs 6 in number, ca. 5-12 /.lID wide separated by ca. 1 J,J.m + wide
canals on the proximal side and ca. 3 - 6 /l-m wide canals on the distal side;
appendices 15 IJ.m long; slide: C 33-a.
Name derivation: gigant-Oat,) = gigantic; forma Oat.) = figure, shape.
Comparison: The Uge specimens are similar to Plicatella triauguliformis Maljavkina (1949) (90 IJ.m in diameter) and Plicatella trichacantha Maljavkina
(1949) (110-140 J.Lm in diameter) from the Lower Cretaceous (Aptian) of western Siberia, USSR, but the specimens are different from the two latter in
number and width of ribs and in form of appendices.
Appendicisporites trichacanthus newly combined by Singh (1964) from
Anemia trichacantha (Maljavkina, 1949) Markova (1961) is not validly published,
because a full and complete reference to the basionym is not provided, as
required by Article 33.2 of the ICBN (1983).
Furthermore, the Uge specimens recemble Appendicisporites giganticus
Groot & Groot (1962) from the Aptian and Cenomanian of Portugal and Appendicisporites cf. A. giganticus Groot & Groot (Burger, 1976, pp. 9-10, pI. 3,
fig. 3; pI. 4, fig. 1 ;pI. 5, fig. 1) fro.Q1 the Lower Cretaceous (Neocomian to
basal Aptian of the Great Artresian Basin in Queensland, Australia, but the
former has a smaller number and broader width of ribs than those of the
two latters.
Botanical affinity: Schizaeceae, Anemia.
Appendicisporites kanukaensis n. sp.
PI. 40, figs. 3-4; pI. 41, figs. 1a-b;pl. 42, figs. 1-2; pI. 43,
figs. 1-2; pI. 44, figs. 2a-b; pI. 46, figs. 1,3.
Description: Trilete spores. Amb triangular with convex sides and slightly
rounded angles in polar view and with straight or slighty convex sides on the
164
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proximal surface and well-rounded side on the distal surface in lateral view.
Trilete laesurae long, slender, straight, almost reaching the equatorial corners. Exine thick, canaliculate; ribs 5-7 in number, ca. 3-6 IJ,m wide, with
more or less sinuous margin, separated by O. 5-1 fJ..m wide canals on the proximal surface and by ca. 2-5 IJ,m wide canals on the distal surface. The
proximal and distal ribs form a protrusion on the radial ridges at the point
of their coalescence. The appendices appear 4-5 lobed in oblique and lateral
views, depending on the prominence of the protrusions on the radial ridges.
They appear smooth and bluntly conical, and are ca. 4-7 IJ,m long. A triangular area on the distal pole is relatively small, simple, and ca. 12 -13 IJ,m in
diameter.
Measurements: 57-85 IJ,m in equatorial diameter (7 specimens), 71 IJ,m in
height (one specimen).
(A)(C 33).
Holotype: PI. 41, figs. la - b; 85 X 85 IJ,m in equatorial diameter; ribs 7 in
number, ca. 4-5 IJ,m wide; margin of rib undulate; appendices 5-7 IJ,m high;
slide C 31 - b.
Name derivation: After the place name Kanuka.
Comparison: The new species is comparable with Appendicisporites potomacensis Brenner from the Lower Cretaceous of U. S. A., Canada and the Netherlands, but differs in its larger size and form of ribs.
Appendicisporites macrorhyzus Maljavkina n. comb.
PI. 45, fig. 2.
1949 Plicatella trichacantha var. macrorhyza Maljavkina, Trudy VNIGRI, n. s., 33, p.
62, pI. 12, fig. 5.
1953 Anemia macrorhyza (Maljavkina) Bolchovitina, Trudy Geol. Inst., Acad. Sci. USSR,
145, GeoI. Ser., 61, p. 39, pI. 4, fig. 16.
1961 Anemia macrorhyza (Maljavkina) Bolchovitina, Trudy Geol. Inst., Acad. Sci. USSR,
40, pp. 54-55, pI. 15, figs. 7a-e; pI. 17, figs. 9a-d.
Description: See Maljavkina (1949) and Bolchovitina (1953, 1961).
Measurements: 60-71 fJ..m in diameter (Bolchovitina, 1961); present specimen:
70 X 53 IJ,m in equatorial diameter; ribs 3-5 IJ,m wide.
165
Previous records: Lower Cretaceous (Albian), western Siberia (Maljavkina,
1949); Lower Cretaceous and Late Jurassic, Yakut; Lower Cretaceous (Early
and Middle Albian), Chylumo-Eniseisk, Ural, Kazakhstan, Crimea, north Aral
(Bolchovitina,1961).
Remarks: Only one specimen was observed. This belongs to Plicatella trichacantha var. macrorhyza Maljavkina (Anemia macrorhyza) and the genus
Plicatella is transferred to the genus Appendicisporites .
Appendicisporites cf. pseudomacrorhyzus Markova n. comb.
Pl. 35, figs. 3a - b.
1961 Anemia pseudomacrorhyza Markova in Samoilovitch et aI.,
p. 67, pI. 17, figs. 2a - b.
Trudy VNIGRI. 177,
Description: See Markova in Samoilovitch et al.(l961).
Measurements: 39.6- (45) - 50 /.lm in diameter (Markova in Samoilovitch et
al., 1961); present specimen: 46 X 40 /.lm in diameter (one specimen).
Occurrence: Uge Member, north of Uge station (A)(C 33).
Previous record: Lower Cretaceous (Aptian-Albian), Tyumensk; Upper Cretaceous, Tomsk province; Lower Cretaceous (Albian), Omsk province; Upper
Cretaceous (Cenomanian), Omsk, Tyumensk, and Tomsk provinces; Upper
Cretaceous(Cenomanian-Turonian), Tomsk and Kemerobsk provinces (Markova in Samoilovitch et al., 1961).
Remarks: Anemia pseudomacrorhyza is now transferred to the genus Appendicisporites Weyland & Krieger, because the genus Anemia is epithet given to
a recent plant.
Appendicisporites rarius n. sp.
Pl. 38, figs. 1a - b; pI .45, figs. 1a - b.
Description: Trilete spores. Amb triangular with convex sides and rounded
corners in polar view and distal outline semicircular in lateral view. Trilete
laesurae distinct, long, straight, reaching nearly the periphery. Exine thick,
canaliculate; ribs 6 -7 in number on the distal side, ca. 3 - 6 J1.m wide, with
smooth or slightly sinuous margins, separated by ca. 1-5 J1.m wide canals,
running parallel to one another and to the equatorial sides. The alternate
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ribs of adjacent sides coalesce in the distal radial regions. The ribs form
each other a protrusion and sometimes a large crest, into which 6 - 7 ribs
merged together, on the radial ridges. A triangular area centered on the
distal pole is simple and large (ca. 20-37 fJ,m in diameter). The contact area
around the proximal pole are smooth. The appendices are smooth and
small, 7-15 fJ,m long.
Measurements: 88-105 j.lm in diameter; 70 j.lm in height.
Occurrence: Uger Member, south of Kanuka (C 31).
Holotype: PI. 38, figs. 1a - b; 92 X 88 fJ,m in diamater; proximal surface
smooth; ribs on the distal surface 6-7 in number, 5-6 fJ,m wide; appendices
7 J.lffi high; slide C 31- d.
Name derivation: rarius (Iat.) = more or less rare or scarce.
Remarks: The ribs of this new species appear quite feebly. The specimen
(pI. 45, figs. la - b) possesses many small pits on the ribs.
Appendicisporites sellingii Pocock
PI. 41, figs. 2.
1964 Appendicisporites sellingii Pocock,
pI. 3. figs. 5-8.
Grana palynologica, 5, no 2. pp. 163-164.
Description: See Pocock (1964).
Measurements: Equatorial diameter 60-66 fJ,m (including appendix) (Pocock,
1964); present specimen: 70 X 65 fJ,m in diameter (one specimen).
Previous record: Late Middle Albian, upper Mannville strata, Saskatchewan
(Canada)(Pocock,1964).
Remarks: The specimen coincides very well with Appendicisporites sellingii
Pocock in large form of appendices. Six ribs regularly disposed in both
proximal and distal faces merged each other to form large and broad appendices in polar view and regular crest-like in appearance in lateral view.
Appendicisporites taneichiensis n. sp.
PI. 39, figs. 2a - d; pI. 40, figs. 1- 2;
pI. 44, figs. 1, 3-4; pI. 46, figs. 4-5.
167
Description: Trilete spores. Equatorial outline triangular with convex sides
and rounded corners. Outline in lateral view semicircular on distal face and
more or less flat on proximal face. Trilete laesurae distal, straight, reaching the periphery on proximal face. Exine thick, canaliculate; ribs 4 -7 in
number on the distal face, ca. 5 -10 /.lm wide, separated by ca. 1-1. 5 J.lm
wide canals; proximal face smooth, more or less flat; appendices on the radial areas smooth, relatively small, 5-12 /.lm high. Some ribs coalesce partially
on the radial areas in some cases. A triangular form on the distal pole is
simple and 15-16 X 20-30 /.lm in diameter.
Measurements: 68 - 86 J.lm in equatorial diameter, 64 - 74 J.lm in height (6
specimens).
Holotype: PI. 40, figs. 2a - b; 84 X 82 Jlm in equatorial diameter; ribs 4 - 5 in
number, 6-9 J.lm wide; appendices 7 /.lm high; slide C 31- b.
Name derivation: After Taneichi town.
Remarks: Pocock (1964) altered Plicatella trichacantha f. typica Maljavkina
(1949) to Appendicisporites trichacanthus trichacanthus (Maljavkina) in accordance with the ICBN and recognized that his specimens are identical to
that described as Anemia trichacantha (Maljavkina) Markova by Markova in
1961 by his acceptance of varieties of this species, notwithstanding his specimens are about half the size of the Maljavkina's original specimen.
The Uge specimens are alike Appendicisporites trichacanthus trichacanthus and A. erdtmanii Pocock from upper Mannville strata (late Middle Albian), Canada, but differ from A. trichacanthus trichcanthus in size and width
of rib and from A. erdtmanii in size, width of ribs, and disposition of the ribs
at the distal pole.
Appendicisporites sp.
PI. 41, figs. 4a - b.
Description: Trilete spore. Amb triangular with straight or slightly concave
sides and narrowly rounded corners in polar veiw. Trilete rays slender, sinuous, ca. 4 J.lm high, extending four-fifths to equatorial corners. Exine thick,
canaliculate; ribs 2-3 in number, ca. 4-5 /.Lm wide; appendices small, ca. 35 J.lm long.
Measurements: 48 X 40 J.lm in equatorial diameter (one specimen).
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Remarks: The single specimen which was observed, was not specifically identifiable.
Genus Baculatisporites Pflug & Thomson 1953.
Type species: Baculatisporites primarius (Wolff 1934) Thomson & Pflug 1953.
Baculatisporites giganticus n. sp.
PI. 22, figs. 3-4.
Description: Trilete miospores. Amb circular or subcircularin polar view.
Trilete laesurae indistinct, straight, long, reaching nearly equatorial periphery. Exine thin, 1. 2 -1. 5 fJ,m thick; sculptural elements more or less densely
distributed, ranging from single bacula or spines to clavae and warts, 1. 54.5 fJ,m long.
Measurements: 73-87 X 68-70 fJ,m in diameter.
Holotype: PI. 22, fig. 4; 87 X 68 fJ,m diameter; exine 1. 5 /.lm thick; ornamentation baculate, clavate, echinate, and gemmate or verrucate, 1. 5-4. 5 /.lI'!llong;
Y mark indistinct, straight, long, reaching the equator; slide C 31- b.
Name derivation: giganticus (Iat.) = gigantic, gigantean.
Remaks: The form-genus Baculatisporites Pflug & Thomson was proposed by
Thomson & Pflug in March, 1953 for all the trilete spores with osmundoid
character of ornamentation. On the other hand, Couper (Fall, 1953) instituted
the form-genus Osmundacidites for Osmunda-spores.
Accordingly, as Krutzsch (1967) pointed out, the genus Bacualtisporites has a priority of nomenclature.
The Uge specimens are colsely similar to Osmundacidites wellmanii Couper (1953) from the Jurassic to Lower Cretaceous of New Zealand, but differ
from O. wellmanii in having larger size. They resemble also to Baculatisporites primarius major Raatz (1937) and B. primarius crassiprimarius Krutzsch(1967) from the Tertiary of Europe, but differ from B. primarius major
in having more densely distributed sculptural elements and B. primarius crassiprimarius in size.
Botanical affinity: Osmundaceae, Osmunda.
Genus Balmeisporites Cookson & Dettmann 1958.
Type species: Balmeisporites holodictyus Cookson & Dettmann 1958.
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Balmeisporites nipponicus n. sp.
PI. 32, figs. 5 -6; pI. 33, figs.
1-4; pI. 34, figs. la - b; pI. 35, figs. la - b.
Description: Trilete megaspores; spore-body circular in outline, covered by
double-layered exospore which is psilate to faintly pitted. The triletelaesurae
are indistinct. The perine is reticulate. The lumina are arranged generally
in pentagons or hexagons and ca. 20-35 IJ,m in diameter. The perine is thrust
upward as a spine with truncated tip and broad base at the angles or the
center of polygons. The muri range generally from 7 IJ,m to 14 IJ,m in length
and the maximal size is ca. 50 IJ,m long. The inner layer in the lumen is composed of small circular and elongated sculptures. The acrolamella is very
finely punctate or chagrenate in sculpture. A mast-like prop which develops
from the spore-body into the acrolamella is 10 /.lm wide at the base, ca. 90
/.lm long, and tapers gradually.
Measurements: Diameter of spore-body (exclusive of the perine and acrolamella)100-120 /.lm ; acrolamellae: 100-140 ).Lm in length and 133 ).Lm in width.
Occurrence: Uge Member, north of Uge station (C 15).
Holotype: PI. 33, figs. la - b; diameter of the spore-body, exclusive of the
perine and acrolamella 120 ).Lm, acrolamella 11 0 -140 /.lm in length and 133
/.lm in width; slide C 15 - e.
Name derivation: After Nippon.
Comparison: All species of Balmeisporites described up to now, are as follows:
Balmeisporites holodictyus Cookson & Dettmann (1958, Lower Cretaceous, Albian), B. tridictyus Cookson & Dettmann (1958, Lower Cretaceous, Albian), B.
glenelgensis Cookson & Dettmann (1958, Upper Cretaceous), B. auriculatus
Hall (1963, Upper Cretaceous, Cenomanian; Bergad, 1973, Cenomanian), B.
diversispinulatus Kondinskaja (1966, Senonian -? Danian), B. rarus Kondinskaja (1966, Senonian-? Danian; Bergad, 1973, Maastrichtian), B. granulatus
Kondinskaja (Senonian -? Danian), B. striatellus Kondinskaja (1966, Senonian -? Danian), B. longirimosus Kondinskaja (1966, Senonian - ? Danian; Bergad ,1973, ::ty1aastrichtian), B. bellus Kondinskaja (1966, Senonian -? Danian;
Srivastava & Binda, 1969, Maastrichtian), B. mollis Kondinskaja (1966, Senonian -? Danian), B. canadensis Srivastava & Binda (1969) emend. Bergad (1973)
(Srivastava & Binda, Maastrichtian; Bergad, Campanian), B. kondinskajae
Srivastava & Binda (1969, Maastrichtian; Bergad, 1973, Campanian), B. dettmannii Srivastava & Binda (1969, Maastrichtian), B. major (Norton 1967)
Bergad (1973) (Maastrichtian), B. minutus Brenner (1968, Albian), and B.
170
K.
TAKAHASHI
& R.
SUGIYAMA
rigidus Bergad (1973, Maastrichtian).
The new species of Uge is different from all species above-mentioned in
morphological features.
? Balmeisporites sp.
Pi. 34, fig. 2.
Description: Trilete megasore (?); spore-body subcircular in outline, covered
by tow-layered exospore which is psilate. Trilete laesurae are indistinct.
The exine is solid and psilate, and 5 /.lID thick. Details of sculptural character
of the spore-body and form of acrolamella are unexplained, because the specimen is very badly preserved.
Measurements: Spore-body 81 X 67 /.lm in diameter.
Occurrence: Uge Member, Uge harbor (C 17).
Remarks: Only one specimen was discovered and is very badly preserved.
An existence of the ridge-like muri parallel to the polar axis of the spore lets
us estimate a comparison with Balmeisporites kondinskajae Srivastava & Binda or B. rarus Kondinskaja.
Genus Biretisporites Delcourt & Sprumont 1955.
emend. Delcourt, Dettmann & Hughes 1963.
Type species: Biretisporites potoniaei Delcourt & Sprumont 1955.
Biretisporites incrassatus Takahashi & Shimono
Pi. 11, figs. 4.
1982 Biretisporites incrassatus Takahashi & Shimono, Bull. Fac. Liberal Arts, Nagasaki
Univ., Nat. Sci., 22, no. 2, p.23, pI. 2, figs 4a-b,5.
Description: See Takahashi & Shimono (1982).
Measurements: 26. 5 - 30 /.lm in equatorial diameter (Takahashi & Shimono,
1982); present specimen: 32 X 24 p,m in equatorial diameter; width of trilete
laesurae 2. 5-3 p,m.
Previous record: Upper Cretaceous (Maastrichtian), Hida (central Japan)
171
(Takahashi & Shimono, 1982).
Remarks: The single specimen encountered has a thin and chagrenate exine.
Genus Camarozonosporites Pant 1954 ex R. Potonie 1956
emend. Klaus 1960.
Subgenus Camarozonosporites (Camarozonosporites)
Krutzsch 1936.
Type species: Camarozonosporites (Camarozonosporites) cretaceous (Weyland & Krieger, 1953) R. Potonie 1956.
Remarks: Krutzsch (1963) classified the genus Camarozonosporites Pant ex
R. Potonie emend. Klaus into three subgenera - Camarozonosporites, Hamulatisporisand Inundatisporis- on the basis of the proximal sculpture. However, Srivastava (1972) considered the subgeneric classification unsuitable,
because the frequent use of form-subgenera creates confusion and has little
stratigraphic use. In spite of the Srivastava's proposal the authors adopt
here the Krutzsch's subgenera classification.
Camarozonosporites CCamarozonosporites) similis n. sp.
PI. 28, figs. 5-7; pI. 29, figs. 1-4; pI. 30, figs. 1- 2.
Description: Trilete microspores. Amb nearly circular with a narrow cingulum that is much narrower at the radial corners of the ambo The cingulum
along the equator between the radial corners is ca. 2.2-5 p,m wide. Proximal surface smooth; trilete rays distinct, straight, and extending almost to
the equator. Distal exine relatively thick, with heavily rugulose sculpture
(hamulate) that may be strongly curved, sinuous and does not extend onto
the proximal side.
Measurements: 42-67 /.lm in equatorial diameter.
(A)(C 33).
Holotype: PI. 3D, figs. 1; 50 X 47 /.lm in equatorial diameter; proximal face
smooth; distal face with hamulate sculpture; maximum width of cingulum
between the radial corners 5 p,m; slide C 31- b.
Name derivation: similis Oat.)= resembling, similar.
Comparison: This new species is closely similar to Camarozonosporites
(Camarozonosporites) decorus (Wolff, 1934) Krutzsch (1959) (Krutzsch, 1963,
K.
172
TAKAHASHI
& R.
SUGIYAMA
p. 126, pI. 44, figs. 1-13) from the Oligocene and Neogene of Germany, but
can be distinguished by the much smoother equatorial contour and more or
less larger size.
Botanical affinity: Lycopodiaceae, Lycopodium.
Camarozonosporites (Camarozonosporites) semilevis Krutzsch
PI. 30, fig. 4.
1963 Camarozonosporites (Camarozonosporites) semileuis Krutzsch, Atlas, Lfg. II, p.124,
pI. 43. figs. 1-11.
1986 Camarozonosporites (Camarozonosporites) semileuis Krutzsch, Takahashi & Jux,
Bull. Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 26. no. 2, p. 58, pI. 4, figs. 5-
8.
Description: See Krutzsch (1963).
Measurements: 23-38 /lm in diameter (Krutzsch, 1963); 33-38 /lm in equatorial diameter (Takahashi & Jux, 1986); present specimen: 38 X 38 /lm in equatorial diameter; cingulum 4 /lm wide.
Occurrence: Uge Member, cliff in north of Uge station (C 6 ).
Previous records: Late Eocene-Oligocene, Germany (Krutzsch, 1963); Late Oligocene, Germany (Takahashi & Jux, 1986).
Remarks: The specimen with small and weak hamulate sculpture on the distal face can be identified with Camarozonosporites (Camarozonosporites)
semilevis Krutzsch from the Eocene to Oligocene of Germany.
Subgenus: Camarozonosporites (Hamulatisporis) (Krutzsch, 1959)
Krutzsch 1963.
Type species: Camarozonosporites (Hamulatisporis) hamulatis Krutzsch 1959.
Camarozonosporites (Hamulatisporis) hamulatis Krutzch
Pl. 29, figs. 5-8; pl. 30, fig. 5.
1959 Hamulatisporis hamulatis Krutzsch, Geologie, Jrg. 8, Beih. 21/22, p. 157, pI. 29,
figs. 326-328.
1963 Camarozonosporites (Hamulatisporis W. Kr. 1959b) Krutzsch, Atlas, Lfg. II, p. 22.
1965 Hamulatisporis hamulatis Krutzsch, Stanley, Bull Amer. Paleont. 49. no. 222, pp.
242- 243, pI. 29, figs. 7-8.
1986 Hamulatisporis hamulatis Krutzsch 1959, Farabee & Canright, Palaeontographica,
173
B, 199, Lfg. 1-3, pp. 19-20, pi. 4, fig. 3.
Description: See Krutzsch (1959, 1963) and Stanley (1965).
Measurements: Ca. 30 J,tm in size (Krutzsch, 1959); 27-34 J,tm in equatorial
diameter (Stanley,1969); 24.2-(28.5)-35 jJ.m in equatorial diameter (Farabee
& Canright, 1986); present specimen: 27-30.5 J,tm X 22.4-27.6 J,tm in equatorial diameter.
Previous records: Middle Eocene, Geiseltal (Germany) (Krutzsch, 1959); Upper
Cretaceous, northwestern South Dakota (Stanley, 1965); Maestrichtian, Wyoming CFatabee & Canright, 1986).
Remarks: All illustrations obtained from the SEM-observation are identical
with e.CH.) hamulatis Krutzsch from the Eocene of Germany.
This apecies is different from the following species of Camarozonosporites CHamulatisporis) : e. CH. ) amplus Stanley n. comb. Cal. Hamulatisporis
amplus Stanly, 1965, p. 242, pI. 29, figs. 1-6) in much smaller size, C.(H.)
albertensis Srivastava n. comb. (aI. Hamulatisporis albertensis Srivastava,
1972, p. 16, pI. 10, figs. 3-8) in much smaller size, C.CH.) loeblichii Srivastava n. comb. Cal. Hamulatisporis loeblichii Srivastava, 1972, p. 16, pI. 10,
figs. 9-10; pI. 11, figs. 1-4) in much smaller size and finer sculpture, C. (H.)
rugulatus Couper n. comb. (al. Perotrilites rugulatus Couper, 1958, p. 147,
pI. 25, figs. 7-8) in much smaller size, and C.CH.) bonitellicola Srivastava n.
comb. Cal. Hamulatisporis bonitellicola Srivastava, 1975, p. 42, pl. 19, figs.
1-4) in much finer sculpture.
Camarozonosporites CHamulatisporis) insignis Norris
Pl. 30, fig. 3.
1967 Camarozonosporites insignis Norris, Palaeontographica, B, 120, Lfg. 1- 4, pp.
96-97, pI. 13, figs. 12-16.
1975 Camarozonosporites insignis Norris 1967, Srivastava, PaleobioI. Contin. , 6, no. 2,
pp. 21-22, pi. 10, figs. 5-14.
Description: See Norris (1967).
Mesurements: Equatorial diameter 30-55 jJ.m (holotype 43 J,tm) (Norris, 1967);
24-33 J,tm in equatorial diameter (Srivastava, 1975); present specimen: 43
jJ.m in equatorial diameter.
174
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TAKAHASHI
& R.
SUGIYAMA
Occurrince: Uge Member, south of Kanuka (C 31).
Previous records: Albian, Maryland [(Brenner, 1963) - Norris, 1967J; Cenomanian, Oklahoma [(Hedlund, 1965)-Norris, 1967J; Late Albian, Alberta (Canada)
(Norris, 1967); Albian, southern United States (Srivastava, 1975).
Remarks: Only one specimen was found. This is identical with Camarozonosporites insignis Norris with indistinct rugulae on the proximal surface and
closely spaced hamulate sculpture (rugulae) on the distal one.
Camarozonosporites CHamulatisporis) sp.
PI. 28, fig. 4.
Description: Trilete microspore. Amb originally circular or subcircular.
The cingulum is slightly thicker between the radial corners than at the corners,
ca. 2-3.5 pm thick.
Hamulate sculpture on the proximal face except the
contact area of the Y mark. The Y mark is distinct, straight, and nearlyextend to the equatorial corners.
Measurements: 65 X 52 /.lm in equatorial diameter.
Remarks: The only one specimen observed is comparable with Camarozonosporites (Hamulatisporis) rugulatus Couper n. comb. (al. Perotrilites rugulatus
Couper, 1958, p. 147, pI. 25, figs. 7-8) and Hamulatisporis rugulatus (Couper
Srivaatava,1972, p. 17, pI. 11, figs. 5-7;pI. 12, figs. 1-3), but differs in
having much finer sculpture.
Genus Cardioangulina Maljavkina 1949 ex Potonie 1960.
Type species: Cardioangulina trichacantha Maljavkina 1949 ex Potonie 1960.
Cardioangulina trichacantha Maljavkina
PI. 4, fig. 7; pI. 7, figs. 1-2.
1949 Cardioangulina trichacantha Maljavkina, Trudy VNIGRI, N. 8., 33, pI. 37, pI. 2,
fig. 8.
1960 Cardioangulina trichacantha Maljavkina emend. Potoni~ , Beih. Geo!. Jb., 39, pp.
28-29, pI. 1, fig. 2.
Description: See Maljavkina (1949).
175
Measurements: 80 /.lm in diameter (Maljavkina, 1949); present specimen: 8493 IJ,m X 82 - 90 IJ,m in equatorial diameter; exine 2. 5 - 5.5 IJ,m thick. (three
specimens).
(A)(C 33).
Previous record: Lower Cretaceous, western Siberia (USSR) (Maljavkina,
1949).
Remarks: The Uge specimens are identified with Cardioangulina trichacantha
Maljavkina (1949) from the Lower Cretaceous, western Siberia, USSR, despite
they are somewhat larger than the holotype. Cardioangulina major Maljavkina (1949) is larger than C. trichacantha and has much concaver equatorial
sides.
Genus Cibotiidites Ross 1949.
Type species: Cibotiidites zonatus Ross 1949.
Cibotiidites sp.
PI. 24, figs. 2a - b.
Description: Trilete spore. Amb triangular with more or less convex sides a
nd slightly pointed corners in polar view. Exine 2-3 Jim thick, with roughly
rugulate sculpture. The trilete laesurae are framed with lips or crests (ca.
5 /.lm wide) and reach the angles.
Remarks: As one specimen was recognized, more specific identification was
omitted.
Genus Cicatricosisporites Potonie & Gelletich 1933.
Type spocies: Cicatricosisporites dorogensis Potonie & Gelletich 1933.
Remarks: The genus Cicatricosisporites is distinguished easily from Appendicisporites in having no appendix and Costatoperforosporites in having smooth
or unperforated ribs.
Cicatricosisporites australiensis (Cookson) Potonie
176
K.
TAKAHASHI
& R.
SUGIYAMA
Pl. 46, fig. 6.
1953 Mohrioisporites australiensis Cookson, Australian Jour. Bot., 1. no. 3, p. 470, pI.
2, figs. 31-34.
1956 Cicatricosisporites (al. Mohrioisporites) australiensis (Cookson) Potoni~, Beih. Ge01. Jb., 23, pp. 47-48.
1967 Cicatricosisporites australiensis (Cookson) Potoni~, Norris, Palaeontographica, B,
120, Lfg. 1-4, p. 92, pI. 11, fig. 14.
1972 Cicatricosisporites australiensis (Cookson) Potoni~, Miki, Jour. Geol. Soc. Japan,
73, no. 5, pI. 1, fig. 4.
1972 Cicatricosisporites australiensis (Cookson) Potoni~, Miki, Jour. Fac. Sci., Hokkaido Univ.,Ser. IV, Geol.&Mineral., 15, nos. 3-4, p.539, pI. 2, figs. 13-14.
1973 Cicatricosisporites australiensis (Cookson) Potoni~, Miki, Jour. Geol. Soc. Japan,
79, no. 3, pI. 1, fig. 4.
1973 Cicatricosisporites australiensis (Cookson) Potoni~, Burger, Spec. pubIs. GeoI. Soc.
Australia, no. 4, pI. 1, fig. 3.
1980 Cicatricosisporites australiensis (Cookson) Potoni~, Burger, BMR Bull., 189, p.
55, pI. 9, figs. 4, 7, 8, (cf. 9. 10).
1981 Cicatricosisporites cf. australiensis (Cookson) Balme, Song et aI., p. 53, pI. 10,
fig.6.
1985 Cicatricosisporites australiensis (Cookson) Potoni~, Yu et aI., p. 82, pI. 10, fig.
14.
1988 Cicatricosisporites cf. australiensis (Cookson) Potonie, Takahashi, Bull. Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 28, no. 2, pp. 94-95, pI. 7, figs. 4 a-c.
Description: See Cookson (1953).
Measurements: 33-50 X 29-34 pm in diameter (Cookson, 1953); 55-63 pm
in equatorial diameter (Miki, 1972); 33-(47)-62 pm in equatorial diameter
(Burger, 1980); 35 jJ,m in diameter (Song et al., 1981); 36 /.lm in diameter (Yu
et al., 1985); 41. 6 /.lm in equatorial diameter (Takahashi, 1988); present specimen (SEM): 36.4 /.lm in equatorial diameter, 25.3 /.lm in height.
Previous records: Pre-Tertiary, South Australia (Australia) (Cookson, 1953);
late Albian, Alberta (Norris, 1967); Upper Cretaceous (Coniacian-$antonian),
northeast Japan (Miki, 1972; Takahashi, 1988); Upper Cretaceous (CenomanianTuronian), Hokkaido (Japan) (Miki, 1973); Lower Cretaceous, Australia, Papua
New Guinea; Cenomanian, Bathurst Island; Neocomian-Albian, Surat Basin
(Burger, 1980); late Lower Cretaceous, Jiangsu (China) (Song et al., 1981);
Lower Cretaceous, Jiangxi (China) (Yu et al., 1985).
Remarks: The present picture was taken by the SEM. This is identified with
Cicatricosisporites australiensis (Cookson)Potonie by size of the spore and
177
number of rib.
Botanical affinity: Schizaeaceae, Mohria.
Cicatricosisporites brevilaesuratus Couper emend. Kemp
Pl. 43, figs. 4a - b.
1985 Cicatricosisporites brevilaesuratus Couper, Palaeontographiea, B, 130, Lfg. 4 - 6,
p. 136, pI. 18, figs. 1-3.
1970 Cicatricosisporites brevilaesuratus Couper, emend. Kemp, Palaeontographica, B, 131,
Lfg. 1-4, pp. 94-95, pI. 13, figs. 12-14; pI. 14, figs, 1-4; text-fig. 11.
1985 Cicatricosisporites brevilaesuratus Couper, Yu et al., p. 79, pI. 9, fig. 4.
Description: See Couper (1958) and Kemp (1970).
Measurements: 70-(90)-120 IJ.m in equatorial diameter (Couper, 1958); 52(72) -92 IJ.m in equatorial diameter (Kemp, 1970); 80 IJ.rn in diameter (Yu et
al., 1985); present specimen: 75 IJ.ffi in equatorial diameter, 64 IJ.m in height,
rib 5-8 IJ.rn wide.
Previous records: Wealden and Aptian, British Isles (Couper, 1958); AptianAlbian, Portugal (Groot & Groot, 1962); Neocomian-Albian, eastern U. S. A.
(Brenner, 1963); Barremian-Lower Aptian, southern England (Kemp, 1970);
Lower Cretaceous (Hauterivian- Barremian), Jiangxi (China) (Yu et al., 1985).
Remarks: The single specimen observed is compared thoroughly with Cicatricosisporites breuilaesuratus Couper emend. Kemp, especially with the Kemp's
illustrations (1973, pI .13, figs. 12-14).
Botanical affinity: Schizaeaceae, M ohria.
Cicatricosisporites dorogensis Potonie & Gelletich
Pl. 45, fig. 5.
1933 Cicatricosisporites dorogensis Potoni(3 & Gelletich, Sitzungsber. Ges. Naturf. FreundeBerl~n, p. 552, pI. 1, figs. 1-5.
1934 Sporites dorogensis (Potoni(i & Gelletich 1933) Potoni~, Pall:\obot. Petrogr. Brennst.,
Preuss. Geoi. Landesanstalt Berlin, 4, p. 40, pI. 1, fig. 21 (p. p.).
1951 Mohrioisp. dorogensis R. Pot.& Gell., R. Potoni(\, Palaeontographica, B, 91, pI. 20,
fig. 14.
1953 Cicatricosisporites dorogensis Potoni(J & Gelletieh, Thomson & Pflug, Palaeontographiea, B, 93, p. 48, pI. 1, figs. 1-12.
1955 Cicatricosisporites efr. dorogensis Pot. & Gel!., Delcourt & Sprumont, Me. Soc. BeIge
178
1956
1962
1967
1969
1969
1972
1972
1972
1973
1977
1982
K.
TAKAHASHI
& R.
SUGIYAMA
Geol.Paleont.et d'HydroI.,N.S.,in 4°, no. 5, pp. 21-22, pI. 1, figs. 3a-b.
Cicatricosisporites dorogensis R. Pot. & GeII., R. Potoni~, Beih. Geol. Jb., 23, p. 47,
pI. 7, fig. 60.
Cicatricosisporites dorogensis R. Potoni~ & Gelletich, Pocock, Palaeontographica,
B, 111, Lfg. 1-3, p. 39, pI. 2, figs. 35-36; pI. 3, figs. 37-41.
Cicatricosisporites dorogensis R. Pot. & Gell. 1933, Krutzsch, Atlas, Lfg. IV & V, p.
BO, pI. 22, figs. 1-6.
Cicatricosisporites dorogensis R. Potoni~ & Gelletich, Trudy Inst. Geol. Geophy.,
Acad. Sci. USSR, Siberian Br., 91, pp. 45-46, pI. 2, figs. 3-5.
Cicatricosisporites dorogensis R. Pot. & Gell. Bratzeva,Trudy Geol. Inst., Acad. Sci.
USSR, 207, pi. 20, fig. 8.
Cicatricosisporites dorogensis Pot.& Gell., Miki, Jour. GooI. Soc. Japan, 78, no. 5,
pI. 1, fig. 3.
Cicatricosisporites dorogensis Potoni~ & Gelletich, Miki, Jour. Fac. Sci., Hokkaido
Univ., Ser. IV, Geol. & Minerl., 15, nos. 3-4, p. 540, pI. 4, figs. 2-3.
Cicatricosisporites dorogensis R. Potoni~ & Gelletich, 1933, Srivastava, Rev. Palaeobot. Palynol., 14, p. 227, pI. 3, figs. 6-7; pI. 4, figs. 1-11.
Cicatricosisporites dorogensis R. Pot. & Gell. 1933 subsp. dorogensis, Kedves, Studia
BioI. Hung., 12, p. 43, pI. 12, figs. 5-8.
Cicatricosisporites dorogensis R. Pot. & Gell.1933, Krutzsch & Vanhoorne, Palaeontographics, B, 163, p. 17, pI .6, figs. 11-14.
Cicatricosisporites dorogensis Potoni~ & Gelletich, Takahashi & Jux, Bull. Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 23, no. 1, p. 32, pI. 3, figs. 5a- b.
Description: See R. Potoni~ & Gelletich (1933).
Measurements: 59-68 p.m CR. Potonie& Gelletich, 1933); 55-(74) p,m CR. Potonie, 1934); 55 p,m (R. Potonie, 1951);40-80 p,m (Delcourt & Sprumont, 1955);
30-(40)-54 p,m in equatorial diameter (Pocock, 1962); 30-43 p,m in diameter
(Chlonova, 1969); 50-55 p,m in equatorial diameter (Miki, 1972); 30-59 p,m in
equatorial diameter (Srivastava, 1972); 50-90 p,m (Epionis) and 45-80 p,m
(Loksbergen) (Krutzsch & Vanhoorne, 1977); 47 X 45 p,m diameter (Takahashi
& Jux, 1982); present specimen: 66 X 62 p,m in equatorial diameter, exine 2. 2
p,m thick.
Previous records: Late Palaeocene-Eocene, Dorog (Hungary) (R. Potonie &
Gelletich, 1933); Eocene, Geiseltal (R. Potonie, 1934); Late Palaeocene-Oligocene, West Germany (Thomson & Pflug, 1953); Wealden, Hainaut (Delcourt &
Sprumont, 1955); Neocomian-Senonian, western Canada (Pocock, 1962); EoceneMiddle Oligocene, Middle Europe (Krutzsch, 1967); Cretaceous, Siberia and the
Far East (Chlonova, 1969); Maastrichtian, Zeisko-Buleinsk basin (Bratzeva,
179
1969); Coniacian, Futaba (Mild, 1972); Santonian-Campanian, Kuji (Miki, 1972);
Palaeocene, Alabama (U. S. A.) (Srivastava, 1972); Eocene, Northern and Southern Bakony, Mor Graben (Hungary) (Kedves,1973); Upper Landonian, Loksbergen & Epionis (Belgium) (Krutzsch & Vanhoorne, 1977); middle Oligocene,
Bergisch land (West Germany) (Takahashi & Jux, 1982).
Remarks: Cicatricosisporites dorogensis R. Potonie & Gelletich (1933) appears
in the Palaeogene of Europe and in the Cretaceous of Siberia, the Far East
(USSR), and Japan.
Botanical affinity: R. Potonie & Gelletich (1933) compared C. dorogensis with
the recent spores of Lygodium japonicum (Schizaeaceae) and Ceratopteris thalictroides (Parkeriaceae). Thomson & Pflug (1953) pointed out a similarity
to Anemia or Mohria and Kedves (1973) indicated a close relationship to the
family Schizaeaceae (Anemia).
Cicatricosisporites ef.hallei Delcourt & Sprumont
PI. 45, fig. 4.
1955 Cicatricosisporites hallei Delcourt & Sprumont, Mem. Soc. BeIge CeoI. Paleont.
d'HydroI.,N.S.in4°, no. 5. pp. 17-19. pI. 1. figs. la-b.
1963 Cicatricosisporites hallei Deicourt & Sprumont 1955. Delcourt. Dettman & Hughes,
Palaeontology, 6, pt. 2. p. 287, pI. 43, figs. 6-7.
1967 Cicatricosisporites hallei Delcourt & Sprumont 1955, Hedlund, Pollen et spores, 9,
no. 3, pp. 579-580, figs. la-b.
1967 Cicatricosisporites hallei Delcourt & Sprumont, Norris, Palaeontographica, B, 120,
Lfg. 1-4, pp. 92-93, pI. 11, figs. 15-20.
1968 Cicatricosisporites hallei Delcourt & Sprumont 1955, Hedlund & Norris, Pollen et
spores, 10, no. 1, pI. 2. fig. 4.
1971 Cicatricosisporites hallei Deicourt & Sprumont 1955. Playford, Palaeontology, 14,
pt. 4, pI. 103, fig. 19.
1975 Cicatricosisporites hallei Delcourt & Sprumont 1955. Srivastava, Paleobioi. Contin., 6, no. 2, p. 27, pI. 11, figs. 5-6.
1985 Cicatricosisporites hallei Delcourt & Sprumont. Yu et aI., p. 78, pI .8. figs. 9-10.
Description: See Delcourt & Sprumont (1955).
Measurements: 35-(40)-57 IJ,m in equatorial diameter (Delcourt & Sprumont,
1955); 43. 2 X 45. 6 IJ,m and 63. 7 IJ,m in diameter (Hedlund, 1967); 32 - 55 IJ,m in
diameter (Singh, 1971); 39 IJ,m in equatorial diameter (Srivastava, 1975); 42-45
IJ,m in diameter (Yu et aI., 1985); present specimen: 65 X 51 IJ,m in equatorial
diameter, ribs 2-3.5 IJ,m wide.
180
K.
TAKAHASHI
& R.
SUGIYAMA
Previous records: Wealden, Hainaut (Belgium) (Delcourt & Sprumont, 1955;
Delcourt, Dettmann & Hughes, 1963); Albian, Alberta (Singh, 1964); Albian,
Alberta (Norris, 1967); Cenomanian, Oklahoma (Hedlund, 1967); Albian, Oklahoma (Hedlund & Norris, 1968); Albian, Saskatchewan and Manitoba (Playford, 1971); Albian, southern U. S. A.(Srivastava, 1975); Middle Cretaceous
(Aptian-Albian), Jiangxi (China) (Yu et a1., 1985).
Remarks: Cicatricosisporites hallei Delcourt & Sprumont is similar to C.
dorogensis Potonie & Gelletich, but differs in possessing larger and higher
mun.
Botanical affinity: Schizaeaceae.
Cicatricosisporites minuticanaliculatus n. sp.
PI. 47, figs. 3-4; pI. 48, fig. 1; pI. 49, figs. 2-3.
Description: Trilete microspores. Outline triangular with convex or rarely
concave sides and rounded corners in polar or somewhat oblique view and
subcircular in lateral view. Trilete laesurae with ca. 4 f..lm wide lip extend
nearly to the equatorial periphery. Proximal face pyramidal; distal face
convex, occupying the most part of the spore body. Exine about 2. 5-5. 5
f..lm thick; sculpture canaliculate; 10-13 proximal ribs of each inter-radial
region parallel to the side and one laesura and oblique to the other laesura
and sides; proximal area around the laesurae smooth; distal pattern consists
of 3 or 4 sets of ribs, the ribs are parallel to each other within each set but oblique to the ribs of the other two sets and to their sides, ribs about 3 - 4 f..lm
wide and with somewhat sinuous margins separated by 1-2 f..lm wide canals,
sometimes ribs within a set branched. No trianqular area on the distal pole.
Measurements: 72.5-91. 3 f..lm in equatorial diameter, 84-85 f..lm in height.
Holotype: PI. 47, fig. 4; 91. 3 X 72. 5 /l-m in equatorial diameter; exine 2.5-4.3
f..lm thick, canaliculate; laesurae with 4 f..lm wide lip; distal ribs consists of 4
sets, each set oblique, 8-10 ribs; slide C 31- b.
Name derivation: minutusOat.) = small, minute; canaliculatus Oat.)= canaliculate.
Comparison: The new species is alike both Cicatricosisporites augustus Singh
(1971) and C. delicatus Phillips & Felix (1972) in rib pattern on the distal face,
but the former is much larger than the two latters.
181
Cicatricosisporites pseudotertiarius Krutzsch
PI. 42, fig. 3.
1959 Cicatricosisporites pseudotertiarius Krutzsch, Geologie, Jrg. 8. Beih. 21/22, p. 170,
pI. 32, figs. 346 - 348.
Description: See Krutzch (1959).
Measurements: Ca. 60-70 J1,m in size (Krutzsch, 1959); present specimen: 61
J1,m in height.
Previous record: Middle Eocene, Geiseltal (Germany) (Krutzsch, 1959).
Remarks: The single specimen observed here is identified with Cicatricosisporites pseudotertiarius Krutzsch from the Middle Eocene of Geiseltal (Germany). This specimen possesses eight remarkable ribs with sinuous margins
on each side.
Cicatricosisporites senonicus n. sp.
PI. 47, figs. 2,5; pI. 48, figs. 2-3; pI .49, fig. 1.
Description: Trilete microspores. Amb subtriangular or subcircular with
convex sides and rounded corners in oblique and lateral views. Y mark slender, with ca. 3-4 J1,m wide lips, extending nearly to the equatorial corners.
Proximal face pyramidal, psilate; distal face convex, occupying the most part
of the spore body.
Exine 2 - 3
J1,m
thick; sculpture cica tricose or canalicu-
late; distal sculptural pattern consists of 3 or 4 sets of 10 -13 ribs in number
with a very weak contrast.
The ribs are parallel to each other within each
set but oblique to the ribs of the other two or three sets and to their sides.
They are about 2-3 J1,m wide separated by 1. 5-3 JJ,m wide canals and develop
very weakly with strongly sinuate margins. Sometimes ribs within a set are
branched or anastomosed.
Measurements: 86-108 J1,m in diameter, 97 J1,m in height (5 specimens).
Holotype: PI. 47, fig. 2; 86 X 86 J1,m in diameter; exine 2. 5 J1,m thick, cicatricose or canaliculate; ribs 2-3 J1,m wide separated by 1. 5 - 2 J1,m wide canals,
developing very weakly; slide C 31 - b.
Name derivation: senonicus= from the stratigraphic term Senonian.
Remarks: The new species, Cicatricosisporites senonicus, is similar to C. minu-
182
K.
TAKAHASHI
& R.
SUGIYAMA
ticanliculatus n. sp., but differs in possessing very weakly developed ribs with
strongly ragged margins.
Cicatricosisporites subrotundus Brenner
PI. 47, figs. 1a - b.
1963 Cicatricosisporites' subroturu:h.Ls Brenner,
Bull., 27, p. 51, pi. 10, figs. 1-2.
Dept. CeoI., Mines and Water Resources,
Description: See Brenner (1963).
Measurements: 37- (46) - 55 J.lm in maximum diameter, muri 6 - 8 J.lm wide
(Brenner, 1963); present specimen: 54 X 48 /.lm in equatorial diameter, muri
(ribs) 7 J.lm wide.
Previous record: Lower Cretaceous (Albian), Maryland (U. S. A.) (Brenner,
1963).
Remarks: The single specimen encountered here is identified well enough with
Cicatricosisporites subrotundus Brenner (1963) from the Lower Cretaceous
(Albian) Patapsco Formation of the Potomac Group, Maryland (U. S. A.) in
all diagnostic features and can be distinguished from Cicatricosisporites patapscoensis Brenner (1963) by its narrower and more numerous muri.
Genus Cingulatisporites Thomson 1953 emend. Potonie 1956.
Type species: Cingulatisporites levispeciosus Pflug 1953.
Remarks: The genus Cingulatisporites was established by Thomson in Thomson & Pflug (1953) for trilete spore having an uniformly developed zona (cingulum). However, Krutzsch (1963) asserted that the type species, C. levispeciosus, must belong to Leiotriletes, while besides he wrote by mistake Cingulatisporites levis Thomson & Pflug 1953.
Hiltman (1967) gave the emended diagosis of Cingulatisporites and excluded verrucate to corrugate forms in agreement with Potonie (1955, p. 58).
The genus Cingutriletes Pierce (1961) with equatorial flange or cingulum
was instituted by Pierce (1969, p. 25). However, Dettmann (1963) and Krutzsch (1963) pointed out that this includes the genera Gleicheniidites, Stereisporites etc. and Krutzsch (1963) transferred the type species Cingutriletes
183
conguens Pierce (1961) to Stereisporites subgenus Cingutriletes congurens
(Pierce) Krutzsch as a subgenotype. Accordingly, the genus Cingutriletes
becomes null.
Cingulatisporites iwatensis n. sp.
PI. 23, figs. 2, 3a - b.
Description: Trilete microspores. Amb triangular with convex sides and rounded corners in polar view. A narrow zona (cingulum) is ca. 2-3 f.J.,m wide
and surrounds equatorially the spore body. The trilete mark is straight,
with narrow lips (ca. 1-4 IJ,m wide), and reachs the inner edge of the zona.
Exine very finely punctate.
Measurements: 46 X 35-38 IJ,m in equatorial diameter (two specimens).
Holotype: PI. 23, fig. 2; 46 X 35 IJ,m in equatorial diameter; zona (cingulum)
2 IJ,m wide; endexine O. 5 IJ,m thick; exine finely punctate; Y mark framed with
ca. 2 f.J.,m wide lips; slide C 31-c.
Name derivation: iwatensis=from Iwate Prefecture.
Comparison: The new species, Cingulatisporites iwatensis, is comparable with
Cingulatisporites levispeciosus Pflug in Thomson & Pflug (1953, p. 58, pI. 1,
fig. 16) from Palaeocene brown coal of Wehmingen near Sarstedt, Hannover
(West Germany), but differs from the latter in having Y mark always framed
with lips and finely punctate exine.
Genus Clavatisporites Kedves & Simoncsics 1964.
Type species: Clavatisporites clarus Kedves & Simoncsics 1964.
Remarks: The genus Clavatisporites Kedves & Simoncsics (1964) provided with
dense ornamentation of clavate elements is a senior synonym of Clavatriletes
Herbst (1965) with coarse clavae.
Clavatisporites sp.
PI. 23, figs. 7a - b.
Description: Azonotrilete microspore. Outline triangular with convex sides
and rounded corners in polar view. Exine thin, with dense ornamentation of
clavate elements; clavae 2-3 IJ,m high, heads 1- 5 IJ,m wide. Trilete laesurae
K.
184
TAKAHASHI
& R.
SUGIYAMA
reach the equatorial corners and are framed with ca. 3-4 lim wide lips.
Measurements: 76 X 74 J,lm in equatorial diameter (one specimen).
Remarks: The single specimen was observed. This resembles Clavatisporites
clarus Kedves & Simoncsics (1964), C. pulcher Kedves & Simoncsics (1964), and
C. (al. Clavatriletes) hammenii Herbst n. comb. (Herbst, 1965, vol. 4, no. 5.
p. 144. pI. 2, figs. 14-15), but differs from C. clarus in its larger size, from
C. pulcher in its larger size and triangular form, and from C. hammenii in its
larger size and shorter clavae. After all, due to only one specimen, the authors cannot determine a specific epithet.
Genus Cyathidites Couper 1953.
Type' species: Cyathidites australis Couper 1953.
Cyathidites australis Couper
PI. 8. figs. 6-7; pI. 9, figs. 1-2. 5.
1953 Cyathidites australis Couper, N. Z. G. S., Paleont. Bull., vol. 22, p. 'l7, pI. 2, figs.
11-12.
1953 Cyathidites australis Couper, 1953. Delcourt & Sprumont, Mem. Soc. Beige GeoI,
Paleont. d'HydroI., N. S., In 4°, no. 5. pp. 27-28.
1958 Cyathidites australis Couper, R. Potoni~, Beih. Geol. Jb., 23. p. 13, pI. 1, fig. 2.
1966 Cyathidites australis Couper, Burger, J. J. Groen &Zoon, p. 237, pI. 2, fig. 2; pI.
5. fig. 2.
1970 Cyathidites australis Couper, Kemp, Palaeontographica, B, 131, p. 84, pl. 10, fig.
1.
1972 Cyathidites australis Couper, Miki, Jour. Fac. Sci., Hokkaido Univ., Ser. IV, Geoi.
& MineraI., 15, nos. 3-4, p. 543, pI. 4, fig. 9.
1972 Cyathidites australis Couper, Miki, Jour. Geol. Soc. Japan, 78, no. 5, pl. I, fig.
5.
1973 Cyathidites australis Couper, Miki, Jour. Geoi. Soc. Japan, 79, no. 3.
1973 Cyathidites australis Couper, Takahashi, Palynol. Cenophyta, pI. I, fig. 1.
1976 Cyathidites australis Couper, Norvick & Burger, BMR Bull., 151, p. 116, pI. 18,
figs. 3-5.
1978 Cyathidites australis Couper, Lei, Acta Bot. Sinica, 20, no. 3, pI. 1, fig. 2.
1980 Cyathidites australis Couper, Burger, BMR Bull., 189. P. 48, pI. I, figs. 1, 2, 6.
1981 Cyathidites australis Couper, Lei, Acta Bot. Sinica, 23. no. 3, pI. 1, fig. 3.
1988 Cyathidites australis Couper, Takahashi, Bull. Fac. Liberal Arts, Nagasaki Univ.,
Nat. Sci., 28, no. 2, pp. 80-81, pI. 1, figs. 7.
185
Description: See Couper (1953).
Measurements: 54-(60)-77 j1,m in equatorial diameter (Couper, 1953); 5477 j1,m in equatorial diameter (Delcourt & Sprumont, 1955); 55-70 j1,m in size
(Burger, 1966); 55-(63)-88 j1,m in equatorial diameter (Kemp, 1970); 50-78
j1,m in equatorial diameter (Miki,1972); 78.3 j1,m in equatorial diameter
(Takahashi, 1988); present specimens: 50-69 j1,m X 45-68 j1,m in equatorial
diameter.
Occurrence: Uge Member, south of Kanuka (C 31) and Uge (C 34).
Previous records: Jurassic to Lower Cretaceous, New Zealand (Couper, 1953);
Wealden, Hainaut (Belgium) (Delcourt & Sprumont, 1955); Jurassic to Berriasian, the eastern Netherlands (Burger, 1966); Upper Aptian to Albian, southern
England (Kemp, 1970); Santonian, Kuji (Miki, 1972); Coniacian, Futaba (Miki,
1972); Turonian, Saku (Hokkaido) (Miki, 1973); Cenomanian, Bathurst Basin
(Australia) (Norvick & Burger, 1976); Early and Middle Albian, Surat Basin
(Australia) (Burger, 1980); Coniacian to Santonian, Futaba (Takahashi, 1988).
Remarks: Cyathidites australis Couper is notably less abundant and much
larger than C. minor Couper. This species occurs in the Turonian to Santonian
in Japan.
Botanical affinity: Cyatheaceae, Cyathea.
Cyathidites minor Couper
PI. 8, figs. 8-10; pI. 9, figs. 3-4, 6.
1953 Cyathidites minor Couper, N. Z. G. S., Paleont. BulL, 22, p. 28, pI. 2, fig. 13.
1962 Cyathidites minor Couper, Pocock, Palaeontographica, B, 111, Lfg. 1-3, p. 43,
pI. 4, figs. 57 - 58.
1966 Cyathidites minor Couper, Burger, J. J. Groen & 2oon, p. 237, pI. 4, fig. 1.
1967 Cyathidites minor Couper, Norris, Palaeontographica, B, 120 Lfg. 1-4, p. 86, pI.
10, fig. 2,
1970 Deltoidospora minor Couper, Pocock, Palaeontographica, B, 130, Lig. 1-2, p. 28,
pI. 5, fig. 3.
1970 Cyathidites minor Couper, Dutta & Sah, Palaeontographica, B, 131. Lfg. 1-4, p.
11, pL 2, figs. 2, 5, 6.
1970 Cyathidites minor Couper, Kemp, Palaeontographica, B, 131, Lfg. 1-4, p. 84, pI.
10. figs. 2-3.
1972 Cyathidites minor Couper, Miki, Jour. Geol. Soc. Japan, 78. no. 5, pI. I, fig. 6.
1973 Cyathidites minor Couper, Miki, Jour. Geol. Soc. Japan, 73, no. 3, pI. 1, fig. 8.
1973 Cyathidites minor Couper, Takahashi, Palynol. Cenophyta, pI. I, fig. 2.
1975 Deltoidospora minor (Couper)Pocock, Srivastava, Paleobiol. Continent., 6, no, 2,
p. 37, pI. 17. figs. 4-5.
186
1980
1981
1985
1985
K.
Cyathidites minor Couper,
11, pI. 1, fig. 2.
1988 Cyathidites minor Couper,
Sic.,28, no. 2, pp. 81-82,
TAKAHASHI
& R.
SUGIYAMA
Burger, BMR Bull., 189, p. 48, pI. 1, figs. 4-5.
Lei, Acta Bot. Sinica, 23, no. 3, pI. 1, fig. 2.
Yu et a1., p. 87, pI. 12, figs. 10, 12-15.
Sun et aI., Bull. Inst. GeoI., Chinese Acad. Geol, Sci.,no.
Takahashi, Bull. Fac. Liberal Arts, Nagasaki Univ., Nat.
pI. 1, figs. 8-10.
Measurements: 31-(35)-45 Jim in equatorial diameter (Couper, 1953); 24(35)-45 Jim in equatorial diameter (Pocock, 1962); 25-55 /.lm in size (Burger,
1966); 30.0-(36.0)-45.0 J1m in equatorial diameter (Pocock, 1970); 39-50
/.lm in size (Dutta & Sah, 1970); 26-(37)-49 f../:ffi in equatorial diameter (Kemp,
1970); 25-45 J1m in equatorial diameter (Singh, 1964, 1971); 25-56 J1m in size
(Srivastava, 1975); 31-42 Jim in diameter (Yu et aI., 1985); 31-39 /lJIl in equatorial diameter (Takahashi, 1988); present specimens: 32-46 J1m X 30-46 J1m
in equatorial diameter.
Occurrence: Uge Member, south of Kanuka (C 31), north of Uge station (C33),
and Uge harbor (A)(C 39).
Previous records: Jurassic to Lower Cretaceous, New Zealand (Couper, 1953);
Jurassic to Upper Cretaceous, western Canada (Pocock, 1962, 1970); Jurassic
to Middle Valanginian, the eastern Netherlands (Burger, 1966); Albian, Alberta
(Norris, 1967); Tertiary (Lower Eocene), Assam (Dutta & Sch, 1970); AptianAlbian, southern England (Kemp, 1970); Coniacian,Futaba (Japan) (Miki,
1972); Cenomanian to Turonian, Saku (Hokkaido) (Miki, 1973); Coniacian
and Santonian, Futaba (Takahashi, 1973, 1988); Albian, southern U. S. A. (Srivastava, 1975); Cenomanian, Bathurst Island (Australia) (Norvick & Burger,
1976); Lower Cretaceous, Surat Basin (Australia) (Burger, 1980); Upper Triassic to Jurassic, Lancang (China) (Lei, 1981); Lower Cretaceous (BerriasianBarremian), Jiangxi (China) (Yu et aI., 1985); Paleocene-Early Eocene, Lingbao
Basin (China) (Sun et aI., 1985).
Remarks: Up to this time this species is reported from the Cenemanian to
Santonian in Japan.
Botanical affinity: Cyatheaceae, Cyathea.
Cyathidites splendens Harris
PI. 2, fig. 1.
1965 Cyathidites splendens Harris, Palaeontographica, B, 115, Lfg. 4-6, p. 79, pI. 24,
187
figs. 13-15.
Description: See Harris (1965).
Measurements: 88- (96) -103 /lm in equatorial diameter (Harris, 1965); present specimen: 113 X 111 /lm in equatorial diameter, exine 4-6 /lm thick.
Occurrence: Uge Member, Uge harbor (B)(C 40).
Previous record: Basal Tertiary, Victoria (Australia) (Harris, 1965).
Comparison: The present specimen possesses somewhat larger size and thicker exine than the original specimens described and illustrated by Harris
(1965). Cyathidites splendens Harris is evidently distinguished from Cyathidites australis Couper by its larger size and thicker exine and from Cyathidites punctatus (Delcourt & Sprumont) Delcourt, Dettmann & Hughes by its
larger size and less concave sides.
Botanical affinity: Cyatheaceae (?).
Genus Deltoidospora Miner 1935 ex R. Potonie 1956.
Type species: Deltoidospora hallii Miner 1935.
Remarks: Regarding a nomenclatural validity of the genus Deltoidospora, see
Srivastava's discussion (1975, pp. 36-37) as well as Takahashi's one (1988,
p. 78).
Deltoidospora cascadensis Miner
PI. 4, fig. 2.
1953 Deltoidospora cascadensis Miner,
24, figs. 9-12.
GeoI. & Mineral., 15. nos. 3- 4. p.
Am. Midland Naturalist, 16. no. 4. p. 618. pI.
Miki, Jour. Fac. Sci., Hokkaido Univ., Ser. IV,
549, pI. 6, figs. 2- 3.
Miki, Jour. Geoi. Soc. ,Japan, 78. no. 5. p. 243.
Miki, Jour. GeoI. Soc., Japan, 79, no. 3, p. ID7.
Description: See Miner (1935).
Measurements: 30-41 IJ-m in diameter (Miner, 1935); 28-35 /lm in equatorial
diameter (Miki, 1972); present specimen: 42 X 38 /lm in equatorial diameter,
exine 1. 5 /lm thick.
Previous records: Lower Cretaceous, Montana (U. S. A.)(Miner, 1935); Santon-
188
K.
TAKAHASHI
& R.
SUGIYAMA
ian to (Campanian), Kuji (Miki, 1972): Coniacian, Futaba (Miki, 1972); Cenomanian to Turonian, Saku (Hokkaido) (Miki, 1973).
Remarks: The single specimen wasfound. This is identified with Deltoidospora
cascadensis Miner in size and form.
Deltoidospora diaphana Wilson & Webster
PI. 4, fig. 5 (?): pI. 7, fig. 3; pI. 10, figs. 2-3.
1946 Deltoidospora diaphana Wilson & Webster,
Am. Jour. Bot., 33, no. 4, p. 273, fig.
a.
1957 Deltoidospora diaphana Wilson & Webster, Rouse, Canad. Jour. Bot., 35, p. 364,
pI. 2, figs. 42-43.
1959 Deltoidospora diaphana Wilson & Webster, 1946, Rouse, Micropaleont., 5, no. 3,
p. 311, pI. 1, figs. 31-32.
Description: See Wilson & Webster (1946).
Measurements: 42-46 J.1.m in diameter (Wilson & Webster, 1946); 42-44 J.1.m in
diameter (Rouse, 1957); 40-60 J.1.m in diameter (Rouse, 1959); present specimens: 41-48.5 J.1.m X 25-41 J.1.m in equatorial diameter, exine 3 j..tm thick.
Previous records: Paleocene, Montana (U. S. A.) (Wilson & Webster, 1946);
Santonian, southern Alberta (Canada) (Rouse, 1957); Upper Jurassic, British
Columbia (Canada) (Rouse, 1959).
Remarks: The Uge specimens are identical with those from the Paleocene Fort
Union Formation of Montana (Wilson & Webster, 1946) and with those from
the Upper Jurassic and Upper Cretaceous of western Canada (Rouse, 1957,
1959).
Deltoidospora psilostoma Rouse
PI. 4, figs. 3 - 4.
1959 Deltoidospora psilostoma Rouse, Micropaleont., 5, no. 3, pI. 311, pI.
~
figs. 7-
8.
1972 Deltoidospora cf. psilostomaRouse, Miki, Jour. Fac. Sci., Hokkaido Univ., Ser. IV,
Geo!. & Mineral., 15, nos. 3-4, p. 549, pI. 6, fig. 8.
1973 Deltoidospora psilostoma Rouse, Miki, Jour. Soc. Japan, 79, no. 3, p. 207.
189
1988 Deltoidospora psilostoma Rouse, Takahashi, Bull. Fac. Liberal Arts, Nagasaki
Univ., Nat. Sci., 28, no. 2, p.78, pI. 1, figs. la-b.
Description: See Rouse (1959).
Measurements: 50-70 /.lm in size (Rouse, 1959); 60-70 /.lm in equatorial diameter (Miki, 1972); 46.7-72 /.lm in equatorial diameter (Takahashi, 1988);
present specimens: 70-72 /Jm X 58-68 /Jm in equatorial diameter, exine 3.55 /Jm thick.
Previous records: Upper Jurassic, British Columbia (Canada) (Rouse, 1959);
Santonian, Kuji (Miki, 1972); Turonian, Saku (Hokkaido) (Miki, 1973); Coniacian
to Santonian, Futaba (Takahashi, 1988).
Remarks: Two specimens observed here are identical with Deltoidospora psilostoma Rouse (1959) from the Upper Jurassic Kootenay coal-bearing formation of southeastern British Columbia, Canada in its size and form.
Botanical affinity: Rouse (1959) stated that the botanical affiliations of this
spore are obscure.
Deltoidospora nodaensis Miki
PI. 10, fig. 4.
1972 Deltoidospora nodaense Miki, Jour. Fac. Sci., Hokkaido Univ.,Ser. IV, GeoL & MineraL, 15, nos. 3-4, pp. 549-550, pI. 6, figs. 4-7.
1972 Deltoidospora nodaense Miki, Jour. GeoL Soc. Japan, 78, no. 5, p. 243.
1973 Deltoidospora nodaense Miki, Jour. GeoL Soc. Japan, 79, no. 3, p. 207.
Description: See Miki (1972).
Meacurements: 25-(29)-36 /Jm in equatorial diameter (Miki, 1972); present
specimen: 28.8 X 27 /.lm in equatorial diameter.
Occurrence: Uge Member, north of Uge station (C 33).
Previous records: Santonian to (Campanian), Kuji (Miki, 1972); Coniacian,
Futaba (Miki, 1972); Cenomanian to Turonian, Saku(Hokkaido) (Miki, 1973).,
Remarks: Miki (1972) used Deltoidospora nodaense as a specific eqithet.
However, according to the ICBN Articles 32. 5 and 73.10, the termination-ense
must be corrected to -ensis. This spore is similar to Deltoidospora taenia
Rouse and D. microferma Rou3e from the Upper Cretaceous (Late Santonian
to Early Campanian) and Eocene of western British Columbia, Canada, but
differs from D. taenia in having straight laesurae and from D. microforma in
190
K.
TAKAHASHI
& R.
SUGIYAMA
possessing larger size.
Genus Densoisporites Weyland & Krieger 1953 emend. Dettmann 1963
emend. Bharadwaj & Kumar 1972.
Type species: Densoisporites velatus Weyland & Krieger 1953.
Densoisporites cf. microrugulatus Brenner
Pl. 25, fig. 6.
1963 Densoisporites microrugulatus Brenner, Dept. GeoI. Mines, Water Resour., Bull.,
27, p. 61, pI. 15, figs. 6a - b; pI. 16, fig. 1.
1967 Densoisporites microrugulatus Brenner, Norris, Palaeontogaphica, B, 120, Lfg. 14, p. 99, pI. 14, fig. 11.
1975 Densoisporites microrugulatus Brenner, Williams & Brideaux, GeoI. Surv. Canada,
Bull. 236, pI. 40, fig. 2.
1985 Densoisporites microrugulatus Brenner, Yu et aI., p. 62, pI. 2, figs. 4-5, 10.
Description: See Brenner (1963).
Measurements: 36 - (63) - 77 /.tm in maximan diameter (Brenner, 1963); 4059 J1,m in diameter (Yu et a1., 1985); present specimen: 39 J1,m in equatorial
diameter.
Occurrence: Uge Member, south of Kanaka (C 31).
Previous records: Upper Barremian to Albian, Maryland (U. S. A.) (Brenner,
1963); Albian, Alberta (Canada) (Norris, 1967); Lower and Middle Cretaceous,
Jiangxi (China) (Yu et al., 1985).
Remarks: The single specimen encountered here is comparable with Densoisporites microrugulatus Brenner (1963) from the Potomac Group of Maryland
(U. S. A.). This is alike D. perinatus Couper (1958) from the Jurassic of British
Isles and D. mesozoicus Srivastava & Roy (1964) emend. Bharadwaj & Kumar
(1972) from the Lower Cretaceous of Kutch, India, but differs from two latters
by narrower cingulum.
Botanical affinity: According to Potonie (1956), D. velatus is compared with
the recent spores of Selaginella scandens Spring.
Genus Dictyophyllidites Couper 1958 emend. Dettmann 1963.
Type species: Dictyophyllidites harrisii Couper 1958.
191
Dictyophyllidites harrisii Couper
PI. 9, figs. 7-9; pI. 10, figs. 7-8.
1958 Dictyophyllidites harrisii Couper, Palaeontographica, B, 103, Lfg. 4-6, p. 140,
pI. 21, figs. 5-6.
1970 Deltoidospora harrisii (Couper) Pocock, Palaeontographica, B, 130, Lfg. 1- 2, p.
29, pI. 5, fig. 16.
1972 Dictyophyllidites harrisii Couper 1958, Srivastava, Palaeontographica, B, 139, p.
12, pI. 7, figs. 4-5.
1975 Dictyophyllidites harrisii Couper 1958, Srivastava, Paleobioi. Continent., 6, no. 2,
p. 38, pI. 17, figs. 6-7.
Measurements: 36-56 J,J.m in equatorial diameter (Couper, 1958); 46.3 - 57. 2
J,J.m in equatorial diameter (Pocock, 1970); slightly smaller than the size range
given by Couper (1958) (Srivastava, 1975); present specimens: 42.3 -61 /-tm X
36.2-59 J,J.m in equatorial diameter.
(A)(C 33).
Previous records: Jurassic (Bajocian), British Isles (Couper, 1958); Jurassic,
western Canada (Pocock, 1970); Maastrichtian, Alberta (Canada) (Srivastava,
1972); Albian, southern U. S. A.(Srivastava, 1975).
Botanical affinity: Couper (1958) compared D. harrisii with the spores of the
Jurassic Dictyophyllum rugosum L. & H., and D. equiexinus (Couper) Dettmann
with Phlebopteris angustiloba (PresL) and Matonidium goepperti (Ett.).
Genus Endoculeospora Staplin 1960.
Type spocies: Endoculeospora rarigranulata var. rarigranulata Staplin 1960.
Endoculeospora cf. delicata Burger
PI. 24, fig. 1.
1976 Endoculeospora delicata Burger. BMR, Bull., 160, p. 10, pI. 6, figs. 2-6.
1980 Endoculeospora delicataBurger. BMR, Bull., 189, p. 60, pI. 15, fig. 9.
Description: See Burger (1976).
Measurements: Spore body, equatorial 26 - 34 /-tm; entire spore, equatorial
34-44 j.1m (Burger, 1976); present specimen: spore body 43 X 27 /-tm in diameter,
entire spore 47 X 41 J,J.m in diameter.
192
K.
TAKAHASHI
& R.
SUGIYAMA
Occurrence: Uge Member, south of Kanaka (C 31).
Previous records: Neocomian to Aptian, Queensland (Australia) (Burger,
1976); Albian, Surat Basin (Australia) (Burger, 1980).
Remarks: The specimen found here is somewhat larger than the Burger's
original specimens.
Botonical affinity: Unknown.
Genus Extrapunctatosporis Krutzsch 1959.
Type species: Extrapunctatosporis extrapunctoides Krutzch 1959.
Extrapunctatosporis fayumensis Takahashi & Jux
PI. 53, fig. 2.
1989 Extrapunctatosporis/ayumensis Takahashi & Jux, Bull. Fac. Liberal Arts, Nagasaki
Univ., Nat. Sci., 29, no. 2. pp. 385-386. pI. 6. figs. 13-15; pI. 7. figs. 2-3 (cf.).
Description: See Takahashi & Jux (1989).
Measurements: 33-48 ;.tID in length and 26-39 /.lm in width (Takahashi & Jux,
1989); present specimen: 41 X 35 j.lm.
Previous record: Late Eocene to Early Oligocene, Fayum Oasis (Egypt) (Takahashi & Jux, 1989).
Remarks: Only one specimen was found. This is comparable with Extrapunctatosporis fayumensis Takahashi & Jux which is a monolete spore with
subcircular to broad-elliptical contour, slender and narrow monolete mark,
and weakly granulate ornamentation.
Botanical affinity: Athyriaceae, Athyrium.
Extrapunctatosporis microalveolatus Krutzsch
PI. 49, fig. 4.
1967 Extrapunctatosporis microalveolatus Krutzsch, Atlas, Lfg. IV & V, p. 163, pI. 59.
figs. 9-12.
1986 Extrapunctatosporis microalveolatus Krutzsch, Takahashi & Jux, Bull. Fac. Liberal
Arts, Nagasaki Univ., Nat. Sci., 26. no .2, p .74.
Measurements: 27-35 ;.tID in length (Krutzsch, 1967); 30
j.lm
in length (Taka-
193
hashi & Jux , 1986); present specimen: 35 X 20 /.lm.
Previous records: Late Oligocene, Germany (Krutzsch, 1976); Late Oligocene,
St. Augustin (W-Germany) (Takahashi & Jux, 1986).
Remarks: The single specimen observed here is assigned to Extrapunctatosporis microalveolatus Krutzsch which was found only in the Late Oligocene
of Germany.
Botanical affinity: Athyriaceae, Athyrium.
Extrapunctatosporis sp.
PI. 51, fig. 7.
Description: Azonomonolete microspore. Outline bean-shaped in lateral view.
Monolete furrow simple, curved. Exospore thin, 0.5 /.lm thick, very finely
punctate, with secondary folds due to fossilization.
Remarks: The large specimen encountered here is similar to Extrapunctatosporis extrapunctoides Krutzsch (1959) from Middle Eocene coal seam of Geiseltal (Germany), but can be distinguished by its larger size and thinner exospore.
Botanical affinity:Athyriaceae.
Genus Foveosporites Balme 1957.
Type spocies: Foveosporites canalis Balme 1957.
Remarks: The genus Foveosporites was established by Balme (1957) for trilete
spores ornamented with pits or short channels irregularly distributed.
Foveosporites perfossus n. sp.
PI. 28, figs. 2-3.
DescriptioQ: Trilete spores.
Amb subcircular in polar view.
Trilete laesurae
straight, extending nearly to the periphery of the spore. Exine ornamented
with small, rounded or slightly elongated channels (0. 5 - 5 /.lm long) irregularly distributed, somewhat crumpled by secondary folds; ectexine ca. 3 - 4
/.lm thick; endexine ca. 0.5 /.lm thick; muri 2-3 /.lm high.
Measurements: 54-66 p.m X 48-52 p.rn in diameter.
194
K.
TAKAHASHI
& R.
SUGIYAMA
Holotype: PI. 28, fig. 3; 66 X 52 fJ,m in diameter, exine foveae-like channels
irregularly distributed; ectexine ca. 3 - 4 fJ,m thick; endexine O. 5 fJ,m thick;
muri ca. 3 fJ,m high; slide C 31-d.
Name derivation: per (lat.)=through; fossa (lat.)=ditch or trench.
Comparison: The new species resembles Foveosporites canalis Balme from
the Lower Cretaceous of Western Australia, but differs from the latter in
possessing larger size and thicker exine.
Bortanical affinity: Lycopodiaceae, Lycopodium.
Foveosporites sp.
PI. 27, figs. 4a - b.
Description: Trilete spore.
Outline circular in polar VIew. Trilete rays
straight, relatively short, always with 3 fJ,m wide lips. Exine 7-10 fJ,m thick,
ornamented with small and rounded channels (0. 5-1 fJ,m in diameter) irregu1arly distributed, and large and rounded verrucae (6-7 J,Jm in diameter) sporadically distributed on the proximal surface.
Measurements: 105 X 98 p.m in equatorial diameter.
Remarks: A single large form of the trilete spore was discovered, which could
not be identified specifically.
Genus Foveotriletes van der Hammen 1956 ex Potonie 1956.
Type species: Foveotriletes scrobiculatus (Ross 1949) Potonie 1956.
Remarks: Van der Hammen (1956, Bol. Geol., vol. 4, nos. 2-3, p. 36, 76, pI.
3) used the generic name Foveotriletes. However, this is a nomen nudum,
because he did not give its diagnosis (ICBN (1983) Article 41. 2J. Potonie
(1956, p. 43) established validly the genus Foveotriletes for trilete spore with
small round foveae that form a dense shallow reticulum. The genus Foveotriletes Pierce (1961, p. 26) is a junior homonym of Foveotriletes Potonie
(1956).
Foveotriletes
cr. scrobiculatus (Ross) Potonie
PI. 28, fig. 1.
1949 "Trilites scrobiculatus"in Ross, Bull. Geol. lnst. Upsala Univ., 34, pl. I, fig. 5.
195
1953 Microreticulatisporites (Trilites) scrobiculatus (Ross)Weyland & Krieger, Palaeon
tographica, B, 95, Lfg. 1-3, p. 11, pI. 4, fig. 23.
1956 Foveotriletes scrobiculatus (Ross 1949) Potoni~. Beih. Oeol. Jb., 23, p. 43, pl. 5,
fig. 49.
Description: See Potonie (1956).
Measurements: 40-50 Ji,m in diameter (Weyland & Krieger, 1953); ca. 38 Ji,m
(Potonie, 1956); 20-36 Ji,rn X 31-48 fJ.m in size (Jansonius & Hills, 1976); present specimen: 55 X 52 Ji,m in equatorial diameter. exine 2 Ji,m thick.
Previous records: Late Santonian to Early Campanian, southern Sweden
(Ross, 1949); Middle Senonian, Aachen (W-Germany) (Weyland & Krieger,
1953).
Remarks: The specimen found here is somewhat larger than the specimens
previously described of Foveotriletes scrobiculatus.
Genus Gleicheniidites Ross 1949.
Type species: Gleicheniidites senonicus Ross 1949.
Gleicheniidites cf. confossus Hedlund
PI. 14, fig. 3.
1966 Gleicheniidites confossus Hedlund, Oklahoma Geo!.
figs. 8a-c.
SUfV.
Bull., 112, p. 17. pl. 1,
Description: See Hedlund (1966).
Measurements: 25.0-42.0 Ji,m in diameter (Hedlund, 1966); present specimen:
39 X 39 Ji,m in equatorial diameter, exine 2-3 Ji,m (side) and 1. 5 Ji,m (corner).
Precious record: Cenomanian, Oklahoma (U. S. A.) (Hedlund, 1966).
Remarks: The present specimen is identified with Gleicheniidites confossus
Hedlund (1966) from the Cenomanian Woodbine Formation of Oklahoma (U.
S. A.), but possesses somewhat thicker exine.
Botanical affinity: Gleicheniaceae, Gleichenia.
Gleicheniidites senonicus Ross
PI. 12, figs. 8-10; pI. 13, figs. 1-10;
pI. 14, figs. 1- 2.
196
K.
TAKAHASHI
& R.
SUGIYAMA
1949 Gleicheniidites senonicus Ross, Bull. GeoI. Inst. Upsala, 34, pp. 31-32, pI. 1, fig. 3.
1953 Gleicheniidites circinidites Cookson, Austr. Jour. Bot., 1, no. 3, p. 464, pI. 1, figs.
5-6.
1953 Gleicheniidites obtusangulus (R. Pot.) Thomson & Pflug f. minor Pflug, Thomson
& Pflug, Palaeontographica, B, 94, P. 51, pI. 1, figs. 35-40.
1955 Gleicheniidites senonicusRoss, Delcourt & Sprumont, Mem. Soc. BeIge GeoI. Paleont.
d' HydroI., N. S., in 4° ,no. 5, pp. 26-27, pI. 1, fig. 5.
1957 Gleichenia concavisporites Rouse, Canada. Jour. Bot., 35, p. 363, pI. 2, figs. 36,
48; pI. 3, figs. 49.
1958 Gleicheniidites senonicus Ross, Couper, Palaeontographica, B, 103, p. 138, pI. 19,
figs. 13-15.
1959 Gleicheniidites senonicus (Ross 1949), De1court & Sprumont, Ann. Soc. GeoI. Nord,
79, p. 33, pI. 3, fig. 5; pI. 12, fig. 37.
1959 Gleicheniidites (Toridistalisporis) toriconcavus Kruzsch, Geologie, Jrg. 8, Beih. 21
/22, p. 112, pI. 12, figs. 110-111.
1961 Cingutriletes interruptus Pierce, Univ. Minnesota, Minnesota GeoI. Surv., Bull. 42,
p. 26, pI. 1, fig. 5.
1961 Psilatriletes vulgaris Pierce, Univ. Minnesota, Minnesota GeoI. Surv., Bull. 42, pp.
27-28, pI. 1, fig. 9.
1961 Punctatriletes parvimundu.s Pierce, Univ. Minnesota, Minnesota GeoI. Surv., Bull.,
42, pI. 28, pI. 1, fig. 13.
1961 Gleicheniidites corcinidites Cookson, Chlonova, Trudy Inst. GeoI. Geophy., Acad.
Sci. USSR, Siberian Br., 7, p. 44, pI. 3, figs. 19, 21.
1964 Gleicheniidites senonicus Ross, 1949,
Skarby, Acta Univ. Stockholm, Stochholm
Contrib. GeoI. 11, no. 3, pp. 65-73, pI. 1, figs. 1-3; pI. 2, figs. 1-8; pI. 3, figs.
1-11; text-fig. 1:1-11.
1965 Gleichenia circinidites Cookson, Stanley, Bull. Amer. Paleont., 49, no. 222, pp.
246-247, pI. 28, figs. 15-16.
1965 Gleicheniidites circinidites (Cookson) Dettmann, Harris, Palaeontographica, B, 115,
Lfg. 4-6, p. 82, pI. 25, fig. 17.
1966 Gleicheniidites circinidites (Cookson) Brenner, Burger, p. 238, pI. 3, figs. la -c.
1966 Gleicheniidites simplex Burger, p. 239, pI 3, figs. 3a- b.
1966 Gleicheniidites senonicus Ross 1949, Burger, p. 239, pI. 3, figs. 5a - c.
1967 Gleicheniidites circinidites (Cookson) Dettmann, Drugg, Palaeontographica, B, 120,
Lfg. 1-4, p .41, pI. 7, fig. 7.
1967 Gleicheniidites senonicus Ross, Norris Palaeontographica, B, 120, Lfg. 1-4, pp.
95-96, pI. 13, figs. 6-7.
1969 Gleicheniidites senonicus Ross, Chlonova, Trudy, Inst. GeoI. Geophys., Acad. Sci.
USSR, Siberrian Br., 91, p, 47, pI. 3, figs. 7-8.
1969 Gleichenia circinidites Cookson, Chlonova, Trudy Inst. GeoI. Geophys., Acad. Sci.
USSR, Siberian Br., 91, p. 48, pI. 3, fig. 9.
1970 Gleicheniidites senonicus Ross, Kemp, Palaeontographica, B, 131, Lfg. 1- 4, p.
103, pI. 18, figs. 3-7.
197
1972 Gleicheniidites senonicus Ross, Miki, Jour. Fac. Sci., Hokkaido Univ., Ser. IV, Geoi.
& MineraL, 15, nos. 3-4, p. 535, pI. 2, fig. 1.
1972 Gleicheniidites circinidites (Cookson) Krutzsch, Miki, Jour. Fac. Sci., Hokkaido Univ., Ser. IV, Geoi. & Mineral., 15, nos. 3-4, pp. 535-536, pI. 2, fig. 3.
1973 Gleicheniidites senonicus Ross, Miki, Jour. Geoi. Soc. Japan, 79, no. 3, pI. 1, fig.
7.
1975 Gleicheniidites senonicus Ross, 1949, Srivastava, PaleobioI. , Continent., 6, no. 2,
p. 41, pI. 18, figs. 7-15.
1976 Gleicheniidites circinidites (Cookson) Dettmann, Norvick & Burger, BMR, BulL,
151, p. 125, pI. 22, figs. 3-7.
1981 Gleicheniidites senonicus Ross, Sung et al., p. 56, pl. 3, fig. 14.
1986 Gleicheniidites senonicus Ross, Farabee & Canright, Palaeontographica, B, 199, Lfg.
1-3, p. 19, pI. 3, figs. 5-7.
Description: See Ross (1949) and Skarby (1964).
Measurements: 16 X 26 (13-19 X 24-29) /.lrn and 17 X 37 (15-19 X 34-39) /.lm
in diameter, exine 1. 5-2 /.lm thick (Ross, 1949); 27-33 /.lm X 37-60 /.lm (2740 /.lm ) in equatorial diameter, exine 1. 5-2. 5 /.lm thick (Cookson, 1953); 2040 /.lm in diameter (Thomson & Pflug, 1953); 10-30 /.lm in diameter (Delcourt
& Sprumont, 1955); 31-40 /.lm in diameter (Rouse, 1957); 30 /.lm in diameter,
exine 1 /.lm (Krutzch, 1959); 33 /.lm in maximum diameter, exine 1 /.lm thick,
peripheral flange 3. 5 /.lm wide (Pierce, 1961); 30 /.lm in maximum diameter,
exine 1. 5 /.lm thick (Pierce, 1961); 30- (32) - 36 /..lm in diameter, exine 1. 5 /..lm
thick (side 3-4 /.lID thick) (Chlonova, 1961); 24-(33.2)-41 /..lm, 25-(31. 9)41 /.lID, and 25-(31. 3)-38 /..lm in length of side in polar view, 1. 5-(2.6)-4.5
/.lm in width of equatorial thickening (Skarby, 1964); 25-40 /..lm in equatorial
diameter, exine up to 2 /.lm thick between apexes (Stanley, 1965); 33-43 /..lm in
diameter, exine 4. 5- 6 /.lm at the sides (Burger, 1966); 27 - 33 /..lm in diameter,
exine 2.5-3 /.lm at the sides (Burger, 1966); 27-35 /..lm in diameter, exine 2. 53.5 /.lm at the sides (Burger, 1966); 29 - 37 /..lm in diameter, exine 2 - 3 /..lm
thick in the interradial equatorial area (Drugg, 1967); 21-30 /..lm in diameter,
(Chlonova, 1969); 27-37 /..lm in diameter, (Chlonova, 1969); 26-(35)-41 /..lffi in
equatorial diameter(Miki, 1972); 23 /.lm in equatorial diameter(Miki, 1972);
20-30 /.lm in diameter (Srivastava, 1975); 20 /.lm in diameter (Sung et a1., 1981);
22- (28.1)-40 /.lm in equatorial diameter (Farabee & Canright 1986); present
specimens: 22.4-43 /.lm X 22. 9-43 /..lm in equatorial diameter, exine 2. 5-5
/..lm thick at the sides.
(A)(C 33).
198
K.
TAKAHASHI
& R.
SUGIYAMA
Previous records: Upper Cretaceous (Late Santonian), Lake Ivosjoo. (Sweden)
(Ross, 1949); Early Tertiary, south Australia (Cookson, 1953); Early and Middle Tertiary, Germany (Thomson & Pflug, 1953); Wealden, Hainaut (Belgium)
(Delcourt & Sprumont, 1955); Santonian, western Canada (Rouse, 1957); Middle Eocene, Geiseltal (Germany) (Krutzsch, 1959); Lower Upper Cretaceous,
Minnesota (U. S. A.) (Pierce, 1961); Upper Upper Cretaceous, western Siberian
lowland (Chlonova, 1961); Late Upper Cretaceous and Paleocene, South Dakota (U. S. A.) (Stanley, 1965); basal Tertiary, Victoria (Australia) (Harris,
1965); Uppermost Jurassic to Middle Valanginian, the eastern Netherlands
(Burger, 1966); Albian, central Alberta (Canada) (Norris, 1967); Maastrichtian
and Danian, California (U. S. A.) (Drugg, 1967); Santonian to (Campanian),
Kuji (Miki, 1972); Cenomanian to Turonian, Saku (Hokkaido) (Miki, 1973);
Albian, southern United States (Srivastava, 1975); Cenomanian, Bathurst Island
(Australia) (Norvick & Burger, 1976); Lower Cretaceous, Jiangsu (China) (Sung
et al., 1981); Maastrichtian, Wyoming (U. S. A.) (Farabee & Canright, 1986).
Remarks: Skarby (1964) noted the wide variety in the morphology of this
specIes.
Botanical affinity: Gleicheniaceae, Gleichenia and Dicranopteris (Skarby, 1964).
Gleicheniidites verrucatus n. sp.
PI. 23, figs. 1a - b, 5a - b.
Description: Trilete tetrahedral spores. Outline triangular with convex and
concave sides and rounded corners in polar view. Trilete laesurae slender,
narrow, slightly curved, extending to the equatorial corners. Proximal face
rather flat, smooth; distal face verrucate to granulate. A wall thickening
which is developed along the sides between the corners is 2-4 J.lm wide.
Measurements: 30-36 J.lm in equatorial diameter.
Occurrence: Uge Member, north of Uge station (A)(C 31) and Uge harbor (C
17).
Holotype: PI. 23, figs. 1a - b; 30 X 31 J.lm in equatorial diameter; exine 2 J.lm
thick; thickening between the corners 2-3 J.lm wide; ornamentation verrucate
to granulate; slide C 33-a.
Name derivation: From the ornamentation of the distal face; verruca (lat.) =
a wart.
Remarks: The authors cannot find a species comparable with the present specImens. The new species is different from Gleicheniidites peregrinus (Bolkh.)
Krutzsch, G. echinatus (Bolkh.), and G. conspiciendus (Bolkh.) in having dif-
199
ferent ornamentation.
Botanical affinity: Gleicheniaceae.
Genus Ischyosporites Balme 1957.
Type species: Ischyosporites crateris Balme 1957.
Ischyosporites sp.
PI. 27, fig. 3.
Descriptionp: Trilete spore. Amb triangular with convex sides and rounded
apIces. Trilete laesurae slender, straight, almost reaching the proximal periphery. Exine ornamented by heavy ridges anastomosing to from an irregular reticulum with rounded or elliptical lacunae; lumina 4-10 /.lm in diameter, larger on the proximal face and smaller on the distal face; muri 8 /.lffi high
and 5-6 /.lm wide.
Measurements: 86 X 82 /.lm in equatorial diameter (one specimen).
Occurrence: Uge Member, north of Kanuka (C 31).
Remarks: The single specimen was found. Although this has a charateristic
ornamentation, the authors cannot decide its specific name.
Botanical affinity: Schizaeacea, Lygodium.
Genus Jimboisporites Sohma 1969.
Type species: Jimboisporites kujiensis Sohma 1969.
Jimboisporites senonicus Miki
Plo 21, figs. 3a -c; pI. 22, figs. 1-2.
1972 Jimboisporites senonicus Miki, Jour, Fac. Sci., Hokkaido Univ., Ser. IV, Geol. & Mineral.,15, nos. 3-4, pp. 552-553, pI. 7, figs. 1-4.
1972 Jimboisporites senonicus Miki, Jour. Geol. Soc. Japan, 78, no, 5, pI. 1, fig. 11.
Description: See Miki (1972).
Measurements: 35 -50 /.lm in equatorial diameter (Miki, 1972); present specimens: 51-59 /.lm X 45-57 /.lm in equatorial diameter, exine thin or up to 5
/.lm thick, verrucae O. 8-3 /.lm high.
Occurrence: Uge Member, south of Kanuka (C 30, north of Uge station (A)
(C 33), and north of Uge station (C 15).
Previous records: Santonian to Campanian, Kuji (Miki, 1972); Coniacian, Fu-
200
K.
TAKAHASHI
& R.
SUGIYAMA
taba (Miki, 1972).
Remarks: The specimens are identified with Jimboisporites senonicus Miki
(1972) from the Tamayama (Santonian) and Sawayama (Campanian) Formations of Kuji, northeastern Honshu, Japan, although they are somewhat larger In SIze.
Genus Laevigatosporites Ibrahim 1933.
Type species: Laevigatosporites vulgaris (Ibrahim 1932) Ibrahim 1933.
Laevigatosporites dehiscens Takahashi
PI. 49, figs. 5-11; pI. 51, figs. 8-9; pi. 52, figs. 1-4.
1961 Laevigatosporites dehiscens Takahashi, Mem. Fac. Sci., Kyushu Univ., SeT. D,
GeoI., 11, no. 3, p. 290, pI. 16, figs. 4-8.
GeoI., 12, no. 1, pI. 1, figs. 9-16.
GeoI. ,14, no. 3, p. 215, pI. 29, figs. 9-13; pI. 40, figs. 13-15.
1979 Laevigatosporites dehiscens Takahashi, Takahashi & Kim, Palaeontographica, B,
170, Lfg. 1-3. pp. 23-24, pI. 1, figs. 10-11.
1982 Laevigatosporites dehiscens Takahashi, Takahashi & Shimono, Bull. Fac. Liberal
Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, p. 30, pI. 5, figs. 6-7.
1988 Laevigatosporites dehiscens Takahashi, Bull. Fac. LiberalArts, Nagasaki Univ .•
Nat. Sci., 28, no. 2, pp. 100-10l,pl. 10, figs. 6,9,13.
1989 Laevigatosporites dehiscens Takahashi, Takahashi & Jux, Bull. Fac. Liberal Arts,
Nat. Sci., 29, no. 2, p. 381, pI. 5. figs. 13-15.
Description: See Takahashi (1961).
Measurements: Ca. 27.5-52 pm in size (Takahashi, 1961); 31-32 pm X 17.419 pm in size (Takahashi, 1979); 38.7-42.5 pm X 22.5-27.6 pm, exine less
than 1 /.lm thick (Takahashi & Shimono, 1982); 29.2-43.5 /.lm X 16. 1- 25 pm,
exine 1 pm + thick (Takahashi, 1988); 30 - 36 /.lm X 21 - 27 JJ,m in size, exine O. 5
pm or less than O. 5 pm thick (Takahashi & Jux, 1989); present specimens:
28.5-47 pm X 16.5-31 pm in size, exine O. 5-1 pm thick.
(A)(C 33).
Previous records: Palaeogene and Miocene, Hokkaido, Jooban, west J~pan,
Shikoku (Takahashi, 1961, 1962, 1963): Campanian and Maastrichtian (Taka-
201
hashi, 1964); Early and Middle Miocene, Yeoungill Bay (Koera) (Takahashi &
Kim, 1979); Maastrichtian, Hida (Takahashi & Shimono, 1982); Coniacian to
Santonian, Futaba (Takahashi, 1988); Late Eocene to Early Oligocene, Fayum
Oasis(Egypt) (Takahashi & Jux, 1989).
Remarks: This species occurs widely in the Tertiary and Upper Cretaceous
of Japan. This is different from Laeuigatosporites gracilis Wilson & Webster
(1946) from the Palaeocene Fort Union Formation of Montana (U. S. A.) and
L. haaerdti (R. Pot. & Van.) Thomson & Pflug haardti from the Tertiary of
middle Europe by its thinner exine.
Botanical affinity: Polypodiaceae.
Laeuigatosporites longus n. sp.
PI. 50, figs. 8-9.
Description: Monolete spores which has the shape of a long bean in equatorial
view. Spore walliaevigate, two-layered, 1 Ji,m thick; ectexine double as thick
as endexine. Dehiscence furrow slightly concave, moderately long (16-35 /lID).
Measurements: 61-71 j.1,m in length and 28-34 j.1,m in width.
Width/length ratio: 0.46-0.48.
Occurrence: Uge Member, north of Uge station (A) (C 33) and north of Uge
station (C 15).
Holotype: PI. 50, fig. 9; 71 j.1,m in length and 34 j.1,m in wisth; exine smooth,
two-layered, 1 j.1,m thick; dehiscence furrow moderately long, slightly curved;
width/length ratio: 0.48; slide C 33-a.
Name derivation: longus (lat.)=long.
Remarks: Laeuigatosporites longus n. sp. is similar to the Palaeozoic (Namurian to Westphalian) species Laeuigatosporites minor Loose, but the former is
much narrower than the latter and possesses a slightly curved furrow.
Botanical affnity: Polypodiaceae.
Laeuigatosporites nitidulus n. sp.
PI. 50, figs. 4-7; pI. 51, figs. 1-3.
Description: Monolete spore with an elliptical or oval contour in lateral and
polar views. Dehiscence furrow straight, moderately long (23-34 j.1,m), slender but conspicuous when sometimes gaping. Exospore thin, 1. 5
smooth or laevigate, with secondary folds due to fossilization.
Measurements: 55-70 j.1,m in length and 38-50 j.1,m in width.
j.1,m
thick,
202
K.
TAKAHASHI
& R.
SUGIYAMA
Width/1ength ratio: O. 67 - O. 76.
(C 15).
Holotype: Pl. 50, fig. 4; 57 X 38 IJ,m in size; exine smooth, 1. 5 Jim thick; dehiscence furrow straight, gaping, 2.5 fJ,m long; width/length ratio: 0.67; slide
C15-d.
Derivation of name: nitidus (lat.) = bright, brilliant.
Remarks: The present specimens are comparable with Laevigatosporites nitidus (Mamczar) Krutzsh nitidus from the Tertiary in Europe and L. javanicus
Takahashi from the Eocene in Java and the Middle Tertiary in Nigeria, but
differ from L. nitidus nitidus in having larger form and thinner exospore, and
from L. javanicus in possessing larger form.
Laevigatosporites ovoideus Takahashi
Pl. 48, Fig. 7; pI. 52, fig. 7.
1961 Laevigatosporites ovoideus Takahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, Geol.,
11, no. 3, pp. 288-289, pI. 16, figs. 9-14.
1979 Laevigatosporites ovoideus Takahashi, Takahashi & Kim, Palaeantagraphica, B,
170, Lfg. 1-3, p. 24, pI. 1, figs. 12-16; pI. 2, fig. 2.
1982 Laevigatosporites ovoideus Takahashi, Takahashi & Shimana, Bull. Fac. Liberal
Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, pp. 30-31, pI. 5, figs. 14-16.
1988 Laevigatosporites ovoideus Takahashi, Bull. Fac. Liberal Arts, Nagasaki Univ.,
Nat. Sci., 28, no. 2, pp. 101-102, pI. 10, fig. 11.
1989 Laevigatosporites ovoideus Takahashi, Takahashi & Jux, Bull. Fac. Liberal Arts,
Nagasaki Univ., Nat. Sci., 29, no. 2, p. 382, pI. 5, fig. 16; pI. 6, fig. 2.
Measurements: 35-54 pm in size, thin (one-layered) (Takahashi, 1961); 24-40.8
fJ,m in length and 18. 7 - 29. 5 fJ,m in width, exine O. 7 -1 Jim thick (Takahashi
& Kim, 1979); 37-48.4 IJ,m in length and 26-31. 5 IJ,m in width, exine 0.8 Jim
or less (Takahashi & Shimono, 1982); 36.3 X 27.8 Jim in size, exine 1 Jim thick
(Takahashi, 1988); 30 X 27. 5 fJ,m in size, exine O. 5 IJ,m thick (Takahashi & Jux,
1989); present specimens: 29 X 26 Jim (SEM: 37. 7 X 30 Jim) in size, exine O. 5
Jim thick, furrow 16 Jim long.
Previous records: Miocene, Sasebo (Takahashi, 1961); Early and Middle Miocene, Changgi and Yonil, Korea (Takahashi & Kim, 1979); Maastrichtian, Hida
203
(Takahashi & Shimono, 1982); Coniacian, Futaba (Takahashi, 1988); Late Eocene to Early Oligocene, Fayum Oasis, Eqypt (Takahashi & Jux, 1989).
Remarks: The present specimen (pI. 48, fig. 7) is identified rather with Laeuigatosporites ouoideus Takahashi than L. ouatus Wilson & Webster, L. qracilis
Wilson & Webster, and L. oui/ormis Takahashi & Jux because of its elliptical
form and thinner exospore.
Laeuigatosporites probatus Takahashi
PI. 50, fig. 3; pI. 51, fig. 4.
1964 LaeuigatosporitesprobatusTakahashi,
Mem. Fac. Sci., KyushuUniv., Ser. D, Geol.,
14, no. 3, pp. 214-215, pI. 29, fig. 8.
1982 Laeuigatosporites probatus Takahashi, Takahashi & Shimono, Bull. Fac. Liberal
Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, pI. 31, pI. 6. figs. 1-2.
Measurements: 41-52.5 j.1,m in size, exospore thin (Takahashi, 1964); 50-55.5
fJ-m in length and 37 -39 fJ-m in width, exine thin, less than 1 /..trrl thick (Takahashi & Shimono, 1982); present specimens: 44-48 /lm in length and 31-38 /lm
in width, exine thin, O. 5 fJ-ffi thick, furrow 21- 23 fJ-m long.
(A)(C 33).
Previous records: Campanian-Maastrichtian, Yubari coal-field, Hokkaido
(Takahashi, 1964); Maastrichtian, Hida (Takahashi & Shimono, 1982).
Remarks: Laeuigatosporites probatus is a middle-sized monolete spore and
broadly elliptical in lateral view. Its dehiscence furrow with no lip is slender
and narrow. Its exospore is very thin and always secondarily folded.
Laeuigatosporites prominens Takahashi
PI. 50. figs. 1-2; pI. 52. figs. 5-6.
1964 Laeuigatosporites prominens Takahashi,
01.,14, no. 3, pp. 213-214, pI. 29, figs.
Arts, Nagasaki Univ., Nat. Sci., 22, no. 2,
Mem. Fac. Sci., Kyushu Univ., Ser. D, Ge1-7; pI. 40, figs. 12,18.
Takahashi & Shimono, Bull. Fac. Liberal
p. 31, pI. 5, fig. 17; pI. 6, figs. Sa-b.
Bull. Fac. Liberal Arts, Nagasaki Univ.,
204
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TAKAHASHI
& R.
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Nat. Sci., 28. no. 2. p. 102. pI. 10. fig. 14.
Measurements: 37.5-50 Jim in size, exospore 1-2 /l-m thick (Takahashi, 1964);
48.4-54 /l-m in length and 31-33. 5 /l-m in width, exine O. 8 /l-m thick (Takahashi & Shimono, 1982); 56. 7 X 38. 3 /l-m in size, exine 1 /l-m thick (Takahashi,
1988); present specimens: 47-48 /l-m in length and 26-29 /l-m in width (SEM:
44-46 /l-m in length and 26. 5-32. 3 /l-m in width), exine 1 /l-m thick.
(A)(C 33).
Previous records: Campanian-Maastrichtian, Yubari coal-field, Hokkaido
(Takahashi, 1964); Maastrichtian. Hida (Takahashi & Shimono, 1982); Coniacian, Futaba (Takahashi, 1988).
Remarks: Laeuigatosporites prominens Takahashi is a late Upper Cretaceous
species together with L. probatus Takahashi.
Laeuigatosporites senonicus Takahashi
PI. 48, figs. 4, 6; pI. 51, fig. 6.
1964 LaeuigatosporitessenonicusTakahashi, Mem. Fac. Sci., Kyushu Univ., Ser. D, Ceo I. ,
14. no. 3. P. 215. pI. 29. figs. 14-19; pI. 40. figs. 16-17.
1982 Laeuigatosporites senonicus Takahashi, Bull. Fac. Liberal Arts, Nagasaki Univ.,
Nat. Sci., 22. no. 2. p. 30. pI. 5. figs. 8-13.
1988 Laeuigatosporites senonicus Takahashi, Bull. Fac. Liberal Arts, Nagasaki Univ.,
Nat. Sci., 28, no. 2, p. 101, pI. 10. figs. 7. 8. 10, 12.
Measurements: 24-42 /l-m in length and 18-31. 3 /l-m in width, exospore up to
1. 5 j.lm thick (Takahashi, 1964); 30-39 /l-m in length and 24-32.5 /1ITl in width,
exine 0.5-1 /l-m thick (Takahashi & Shimono, 1982); 29.3-38.3 /l-m in length
and 20. 6-28. 4 /l-m in width, exine l/l-m + thick (Takahashi, 1988); present specimens: 33-41 /l-m in length and 28-39 /l-m in width (SEM: 41. 5 X 31. 5 /l-m in
size), exine O. 5 -1. 5 /l-m thick.
Occurrence: Uge Member, south of Kanuka (C 31), Uge (C 34), and Uge harbor (C 17).
Previous records: Campanian-Maastrichtian, Yubari coal-field, Hokkaido (Takahashi, 1964); Maastrichtian, Hida (Takahashi&Shimono, 1982); Coniacian,
205
Futaba (Takahashi, 1988).
Remarks: Laevigatosporites senonicus Takahashi appears restrictedly in late
Upper Cretaceous of Japan.
Laeuigatosporites sp.
PI. 53, fig. 1.
Description: Monolete spore. Outline broadly elliptical in lateral view. Dehiscence side concave; monolete furrow slightly concave, slender, narrow and
moderately long; distal side konvex semicircularly. Exospore smooth, 1 fJ,m
thick, crumpling due to fossilization; exine around the furrow finely punctate.
Measurements: 93 X 69 fJ,m in size.
Width/length ratio: O. 74.
Remarks: The present specimen is a large-sized monolete spore and differs
from the following large-sized monolete spores in having finely punctate ornamentation around the monolete furrow: Laevigatosporites gigantiformis
Takahashi (1961, 73.6-90.7 fJ,m), L. eogigantiformis Takahashi (1962, 58.6-
78.2 fJ,m), L. josensis Takahashi & Jux (1989, 76-80 fJ,m), L. discordatus Pflug
(1953, 50-90 fJ,m), and L. pseudodiscordatus Krutzsch (1959, 50-80 fJ,m).
Genus Latosporites Potonie & Kremp 1954.
Type species: Latosporites latus (Kosanke 1950) Potonie & Kremp 1954.
Latosporites rotundus Takahashi & Jux
PI. 48, fig. 5; pI. 51, fig. 5.
1989 Latosporites rotundus Takahashi & Jux, Bull. Fac. Liberal Arts, Nagasaki Univ.,
Nat. Sci., 29, no. 2, pp. 382-383, pi. 4, figs. 1-2; pi. 5, figs. 1-2.
Description: See Takahashi & Jux (1989).
Mesasurements: 51- 55 J.lm in equatorial axis and 39 - 51 J.lm in polar axis,
exine O. 5 - O. 7 J.lm thick (Takahashi & Jux, 1989); present specimens: 54 fJ,m
in length and 50-55 fJ,m in width, exine up to 1 fJ,m thick.
Occurrence: Uge Member, nouth of Uge station (A) (C 33) and north of Uge
station (C 6).
206
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Previous record: Late Eocene to Early Oligocene, Fayum Oasis, Egypt (Takahashi & Jux, 1989).
Remarks: The genus Latosporites was introduced by Potonie & Kremp (1954)
for monolete spores with broadly oval to circular outlines and distal distenSlOns. The specimens from the Uge Member are identical with Latosporites
rotundus Takahashi & Jux (1989) from the Late Eocene to Early Oligocene
Qasr EI Sagha Formation in Fayum Oasis, Egypt.
Genus Leiotriletes Naumova 1939 ex Ishchenko 1952 emend. Potonie
& Kremp 1954.
Type species: Leiotriletes sphaerotriangulus (Loose 1932) Potonie & Kremp
1954.
Leiotriletes cf. conuexiformis Chlonova
PI. 1, figs. 6a - b; pI. 3, fig. 6.
1960 Leiotriletes conuexiformis Chlonova, Trudy Inst. Geol. Geophys., Acad. Sci. USSR,
Siberian Br., 3, p. 31, pI. 4, figs. 1- 2.
1982 Leiotriletes cf. conuexi/ormis Chlonova, Takahashi & Shimono, Bull. Fac. Liberal
Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, p. 22, pI. 2, fig. 6.
Description: See Chlonova (1960).
Measurements: 30-(36)-39.5 pm in size, exine thin (Chlonova, 1960); 30.535.5 pm in equatorial diameter, exospore thin (Takahashi & Shimono, 1982);
present specimens: 28-30 pm in equatorial diameter, exospore 1 JJ-m thick.
Occurrence: Uge Member, north of Uge station (A)(C 33) and Uge harbor (C
17).
Previous records: Cenomanian-Turonian, Simonobo-Suchkobo, Chulym; Danian-Early Palaeogene, Teuriches (Chlonova, 1960); Maastrichtian, Hida (Takahashi & Shimono, 1982).
Remarks: The present specimens are almost identical with Leiotriletes convexiformis Chlonova from the Upper Cretaceous and Early Palaeogene in the
Chulym-Eniseisk basin, Siberia, but they are more circular and more or less
smaller than the Siberian form.
Leiotriletes giganticus n. sp.
207
PI. 1, figs. 1- 3; pI. 2, figs. 2- 4.
Description: Trilete spores. Amb rounded-triangular with convex or concave
sides and rounded corners in polar view. Exospore smooth, two-layered, 1.53 /l-m thick. Ectexine is thicker than endexine. Trilete laesurae straight or
slightly curved, mostly with lips, extending almost or three fourths the distance to the periphery.
Measurements: 82-140 /l-m X 67-105 J1.m in equatorial diameter.
(A)(C 33).
Holotype: PI. 1, fig. 1; 96 X 89 J1.m in equatorial diameter; exine laevigate,
two-layered, 3 /l-m thick; slide C 31- b.
Name derivation: giganticus Oat.) = gigantic.
Remarks: The trilete large specimens resemble Leiotriletes maxoides Krutzsch
maximus (Pflug) Krutzsch from the Oligocene and Miocene of Europe, L.
grandiosus Krutzsch and L. paramaximus Krutzsch from the Middle Eocene,
Geiselte1 (Germany), but differ from L. maxoides maximus in having a simple
form at end of the Y-mark, from L. grandiosus by their thicker exospore and
larger form, and from L. paramaximus in their larger form.
Botanical affinity: Schizaeaceae, Lygodium.
Leiotriletes maxoides Krutzsch maximus (Pflug) Krutzsch
PI. 4, fig. 1.
1953 Divisisporites maximus Pflug, Thomson & Pflug, Palaeontographica, B, 94, p. 52,
pI. 1, figs. 57 - 58.
1953 Laevigatisporites pseudomaximus Pflug & Thomson, Thomson & Pflug, Palaeontographica, B, 94, p. 54, pI. 2, figs. 20- 22.
1962 Leiotriletes maxoides Krutzsch maximus (Pflug) Krutzsch, Atlas, Lfg. I, p. 20,
pI. 3, figs. 1- 4.
1984 Leiotriletes maxoides Krutzsch 1962a maximus (Pflug 1953) Krutzsch 1959b, Kirchner, Palaeontographica, B, 192, p. 91, pI. 1, fig. 1.
1986 Leiotriletes maxoides Krutzsch maximus (Pflug) Krutzsch, Takahashi & Jux, Bull.
Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 26, no. 2, pp. 37-38, text-fig. 3.
Description: See Krutzsch (1959, 1962).
Measurements: More than 60 Jim, ectexospore more than 2 Jim thick, endexospore less than O. 5 /l-m thick (Pflug in Thomson & Pflug, 1953); more than 80
/l-m in size (Krutzsch, 1962); 80-110 Jim in size (Kirchner, 1984); 80-98 /l-m in
208
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equatorial diameter, exine 2-3 j1,m thick (Takahashi & Jux, 1986); present
specimen: 86 X 84 j1,m in equatorial diameter, exine 3 fJ-m thick.
Previous records: Middle Tertiary, Germany (Pflug in Thomson & Pflug, 1953);
Upper Oligocene, southern Bavaria (W. Germany) (Kirchner,1984): Upper
Oligocene, St. Augustin, (W. Germany) (Takahashu & Jux, 1986).
Remarks: Leiotriletes maxoides maximus differs from L. maxoides maxoides
in having larger size and no pseudotoroidal claw at end of the trilete rays
and from L. maxoides minor in size and thickness of exospore.
Leiotriletes rotundiformis (Maljavkina) Chlonova
PI. 1, figs. 4-5; pI. 2, fig. 5 (cf.).
1949 Cardiolina trisecta Maljavkina var. rotundi/ormis Maljavkina, Trudy VNIGRI, N.
S,33, pp. 37-38, pI. I, figs. 18, 19, 22.
1960 Leiotriletes rotundiformis CMaljavkina) Chlonova, Trudy Inst. Geol. Geophys.,
Acad. Sci. USSR, Siberian Br., 3, p. 32, pI. 4, fig. 6.
1982 Leiotriletes rotundiformis CMaljavkina) Chlonova, Takahashi & Shimono, Bull.
Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 22, no. 2, p. 22, pI. 1, figs. 5-8;
pI. 2, figs. 1-2.
Description: See Maljavkina (1949).
Measurements: 40-60 fJ-m in diameter (Maljavkina, 1949); 52-(56)-60 j1,m in
diameter (Chlonova, 1960); 46-57 fJ-m X 38-52 j1,m in diameter, exospore thin,
less than 1 j1,m thick (Takahashi & Shimono, 1982); present specimens: 61-66
fJ-m X 51-64 j1,m in diameter, exospore 1. 5-2. 5 j1,m thick.
Occurrence: Uge Member, south of Kanuka (C 31) and Uge harbor (C 17).
Previous records: Middle Jurassic, Emba, Sagiz (Maljavkina, 1949): Cenomanian- Turonian, Simonobo-Suchkobo, Chulym (Chlonova, 1960) Maastrichtian,
Hida (Takahashi & Shimono, 1982).
Remarks: The present specimens are referable to Leiotriletes rotundiformis
(Maljavkina) Chlonova from the Cenomanian to Turonian, Chulym-Eniseisk
basin, Siberia, but they are more or less larger than the original specimens
described by Maljavkina.
Leiotriletes wolffi Krutzsch wolffi
PI. 3, fig. 4 ( cf.); pI. 10, fig. 1.
209
1962 Leiotriletes wol//i Krutzsch wol//i, Atlas, Lfg. I, p. 26. pI. 6, figs. 1-14.
1984 Leiotriletes wol//i Krutzsch 1962 wol//i Krutzsch 1962, Mohr, Palaeontographica,
B,191, p. 38, pI. 1. fig. 5.
1986 Leiotriletes wol//i Krutzsch wolffi, Takahashi & Jux, Bull. Fac Liberal Arts, Nagasaki Univ., Nat. Sci., 26. no. 2, pp. 40-41, pI. 2, fig. 7.
Measurements: Ca. 35-45 pm in size, exospore up to 1 p,m thick (Krutzsch,
1962); 35 pm in size, exine thin (ca. 1 pm thick) (Mohr, 1984); 32-44 pm in
size, exine 1. 5-2 pm thick (Takahashi & Jux, 1986); present specimens: 4046 pm X 36.2-44 pm in equatorial diameter, exospore 1. 5 JJ,m thick.
(A)(C 33).
Previous records:Oligocene-Pliocene, Middle Europe (Krutzsch, 1962); Late
Miocene, Rhine land (W. Germany) (Mohr, 1984); Late Oligocene, St. Augustin
(W. Germany) (Takahashi & Jux, 1986).
Remarks: Leiotriletes wol//i Kr. wol//i is superficially similar to L. trianguloides Kr. from the Oligocene to Pliocene of Germany, L. wol//i Kr. brevis Kr.
from the Mio-Pliocene of Germany, L. seidewitzensis Kr. from the Early Miocene of Germany, and L. balowensis Doring from the Late MaIm and Wealden
of NW Germany (DDR), but differs from 1. trianguloides by its spore form,
from L. wol/li brevis by its spore size and length of the Y -rays, from L. seidewitzensis by its spore size and form, and from L. balowensis by its length of
the Y-rays.
? Leiotriletes sp.
PI. 11. fig. 2.
Description: Trilete spore. Outline rounded-triangular with convex sides and
rounded corners in polar view. Trilete laesurae straight, relatively slender,
extending to two-thirds of distance to the equatorial corners. Exospore 35 pm thick, chagrenate.
Measurements: 50 X 44 pm in equatorial diameter.
Remarks: The single specimen which was observed, is doubtful to identify
with the genus Leiotriletes and besides not specifically identifiable.
210
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Genus Leptolepidites Couper 1953.
Type species: Leptolepidites verrucatus Couper 1953.
Leptolepidites sp.
PI. 23, fig. 4.
Description: Trilete spore. Outline triangular with concave sides and rounded
corners in polar view. Exine thin; proximal face mostly smooth and peri
phery of the proximal face verrucate; distal face verrucate; verrucae 2 IJ.m
high. Y -mark conspicuous, straight or slightly sinuous, reaching the equatorial corners.
Remarks: A single specimen was found, but its specific identification IS not
yet possible.
Botanical affinity: ? Leptolepia.
Genus Lycopodiacidites Couper 1953.
Type species: Lycopodiacidites bullerensis Couper 1953.
Lycopodiacidites circuZaris n. sp.
PL 26, figs. 6a - b; pI. 30, figs. 6a - b.
Description: Trilete spores. Amb circular to sub-circular in polar view.
Trilete laesureae conspicuous, sinuous, almost reaching the equatorial periphery. Exine 3-3.5 /.lm thick; proximal face smooth, distal face clearly and/or
heavily sculptured as small and large verrucae to irregular rugulae; verrucae
2-12 /.lm in diameter.
Measurements: 47-49 /.lm X43-45 IJ.m in equatorial diameter.
Holotype: PI. 26, figs. 6a - b; 47 X 45 IJ.m in equatorial dia.meter; exine 3. 5
IJ.m thick; proximal face smooth and distal face sculptured as various verrucae-like warts, 2 -12 IJ.m in width; Y-mark sinuous reaching the equatorial
margin; slide C 31-d.
Name derivation: circularis (lat.) = circular.
Remarks: The present specimens are superficially similar to Lycopodiacidites
bullerensis Couper and L. cirstatus Couper from the Jurassic in New Zealand,
but differ from both species of New Zealand in ornamentation. Klaus (1960)
211
described Lycopodiacidites kuepperi from the Upper Triassic Cardia Formation
and Halobia shale near Salzburg (Austria). However, this differs from the
present specimens in size, ornamentation, and trilete laesurae.
Botanical affinity: Lycopodiaceae.
Genus Lygodiidites Pocock 1964.
Type species: Lygodiidites laevigatus Pocock 1964.
Lygodiidites laevigatus Pocock
PI. 32, fig. 4.
1964 Lygodiidites laevigatus Pocock,
Grana Palynologica, 5. p. 180. pI. 5. fig. 2.
Description: See Pocock (1964).
Measurements: Ca. 62.4 Jlm in equatorial diameter, sexine ca. 6.0 Jlm thick
(Pocock, 1964); present specimen: 57 Jlm in diamter of spore body (50 Jlm in
diameter except sexine), sexine 5 Jlm thick.
Previous record: Middle-Uppe Albian, Saskatchewan (Canada) (Pocock, 1964).
Remarks: This specimen is identical with Lygodiidites laevigatus Pocock from
Upper Mannville strata (Middle-Upper Albian) of Saskatchewan (Canada).
Lygodiidites tohokuensis n. sp.
PI. 31, figs. 1-2; pI. 32, figs. 1-3.
Discription: Trilete spores. Amb rounded-triangular with convex sides and
rounded apices. Exine two-layered, chagrenate; ectexine 5 -16 fJ,m thick,
with sponge-like sculpture, especially remarkable on the corners and near the
margin; endexine thin, smooth. Trilete laesurae conspicuous, slightly undulate, mostly with lips (max. 4-5 Jlm wide).
Measurements: 76-97 Jlm X 65 - 92 IJ,m in equatorial diameter of the whole
body, 61-72 Jlm X 50-69 Jl,m in diameter of central body.
Occurrence: Uge Member, south of Kanuka (C 31), Uge (C 34), and Uge harbor (C 17).
Holotype: PI. 32, fig. 1; 89 X 85 Jl,m in equatorial diameter of the whole body,
69 X 69 Jl,m in diameter of central body; exine 7 -12 Jl,m in width, with spongelike sculpture on marginal zone and near corners; trilete rays with lips (4 - 5
K.
212
TAKAHASHI
& R.
SUGIYAMA
Jlm wide); slide C 34-C.
Name derivation: From Tohoku (northeast) district.
Remarks: The large specimens which were found from three localities of the
Uge Member, resemble Lygodiidites laevigatus Pocock and L. balmei Pocock
from the Middle-Upper Albian Upper Mannville Group of Saskatchwan (Canada), but differ from L. laevigatus in having larger size of spore body and
sponge-like or vacuolate sculpture of exine, and from L. balmei in possessing
larger size of spore body and thicker exine.
Botanical affinity: Schizaeaceae, Lygodium,
Genus Monoleiotriletes Krutzsch 1959.
Type species: Monoleiotriletes angustus Krutzsch 1959.
Monoleiotriletes gracilis Krutzsch.
PI. 2, fig. 6a-b; pI. 3, fig. 7; pI. 5,
figs. 3-5; pI. 10, fig. 5.
1959 Monoleiotriletes gracilis Krutzsch, Geologie, Jg. 8, Beih., 21/22, p. 65, pl. 4,
fig. 24.
1982 Monoleiotriletes gracilis Krutzsch, Takahashi & Jux, Bull. Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 23. no. I, p. 30, pI. 1, fig. 3.
1984 Monoleiotriletes gracilis Krutzsch 1959, Kirchner, Palaeontographica, B, 192, pI.
1, fig. 3.
1986 Monoleiotriletes gracilis Krutzsch, Takahashi & Jux, Bull. Fac. Liberal Arts, Nagasaki Univ., Nat. Sci., 26, no. 2, p. 47, pI. 2, figs. 11-12.
1988 Monoleiotriletescf.gracilis Krutzsch, Takahashi, Bull. Fac. Liberal Arts. Nagasaki
Univ., Nat. Sci., 28. no. 2, p. 86, pI. 3, figs. 12-13.
Measurements: Less than 50 Jlm in size, exine ca. 1/2 - 3/4 Jlm thick (Krutzsch, 1959); 33-50 Jlm diameter, exine up to 1 Jlm thick (Krutzsch, 1962); 36.3
j1,m X 28. 6 j1,m in equatorial diameter, exine O. 5 j1,m thick (Takahashi &
Jux, 1982); 36 Jlm in size (Krutzsch, 1984); 30-40 Jlm in equatorial diameter,
exine less than 1 Jlm thick (Takahashi & Jux, 1986); 30-37. 3 Jlm in equatorial
diameter, exine less than 1 Jlm tick (Takahashi, 1988); present specimens: 3446 Jlm X 34-43 Jlm in equatorial diameter, exine thin.
(A)(C 33).
Previous records: Middle Ecocene, Geiseltal (Germany) (Krutzsch, 1959);

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