A CATALOG OF THE COLEOPTERA OF AMERICA NORTH OF

A CATALOG OF THE COLEOPTERA OF AMERICA NORTH OF

A CATALOG
OF THE COLEOPTERA
OF AMERICA
NORTH OF MEXICO
FAMILY: SILPHIDAE
4~~
UNITED STATES
DEPARTMENT OF
~ AGRICULTURE
{fUJ))
AGRICULTURE
HANDBOOK
NUMBER 529-28
PREPARED 8Y
AGRICULTURAL
RESEARCH
SERVICE
FAMILIES OF COLEOPTERA IN AMERICA NORTH OF MEXICO
Fascicle' Family
Year issued
......... Cupedidae .................. ..... 1979
....... 1982
2 ......... Micromalthidae
3 ......... Carabidac ..
.. .... 1985
4 ......... Rhysodidae ".
...... 1984
5 ......... Amphizoidae
6 ......... Haliplidae .
8 ......... Noteridae .
9 ......... Dytiscidae
10. ....... Gyrinidae ..
13. ....... Sphaeriidae .
14. ....... Hydroscaphidae .
15. ....... H ydraenidae ..
16. ....... Hydrophilidae .
17.. ...... Georyssidae .
18. ....... Sphaeritidae .
20 ........ Histeridae ..
21 ....... Ptiliidae ..
22. ....... Limulodidae ..
23. ...... Dasyceridae ..................
.. .... 1984
24. ....... Micropeplidae .
25. ....... Leptinidae .
26. ....... Leiodidae .
27. ....... Scydmacnidae ..
.. .... 1993
28. ....... Silphidae .
29. ....... Scaphidiidae .
30. ....... Staphylinidae .
31 ....... Pselaphidae .
32. ....... Lucanidae .
33. ....... Passalidae .
.. .... 1984
34 ......... Scarabaeidae .
35. ....... Eucinelidae .
36. ...... Helodidae ..
37. ....... Clambidae ..
38. ...... Dascillidae .
39 ......... Rhipiceridae ..
.. Byrrhidae ..................
40
.. .... 1983
41 ....... Psephenidae .
42. ....... Brachypsectridae ....
43. ....... Artematopidae .
44 ......... Ptilodactylidae .
45 ........ Chelonariidae ..
Fascicle' Family
Year issued
46 ....... CaUirhipidae ..
47 ....... Heteroceridae .. ................. 1978
48 ....... Limnichidae .. .................. 1986
49 ....... Dryopidae
............ 1983
50 ....... Elmidae ..
...... 1983
51 ....... Buprestidae
52 ....... Cebrionidae .
53 ....... Elateridae "'
54 ....... Throscidae .
55. . Ccrophytidae ..
56 . ...... Perothopidae .
57 ....... Eucnemidae .
58 ....... Telegeusidae .
61 ....... Phengodidae .
62 . ...... Lampyridae .
63 ....... Cantharidae ..
64 ....... Lycidae.
65 ....... Derodontidae . .................. 1989
66 ....... Nusodendridae ..
67 ....... Dermestidae ....
69 ....... Ptinidae .
70 . ...... Anobiidae .
.................. 1982
71 . ...... Bostrichidae ...
72 . ...... Lyctidae.
74 ....... Trogositidae ..
76 ....... Cleridae .
78 ....... Melyridae ..
79 . ...... Lymexylidae .
81 . ...... Sphindidae .
82 ....... Nitidulidae .
83 ....... Rhizophagidae .
86 ....... Cucujidae ..
90 ....... Cryptophagidae .
92 . ...... Languriidae .. .................. 1983
93 ....... Ero{ylidae ...
94 ....... Phalacridae .
............. 1982
95 ....... Cerylonidae ..
96 ....... Corylophidae .
97 ....... Coccinellidae ..
98 ....... Endomychidae . ................. 1986
100 ....... Lathridiidae
Fascicle' Family
Year issued
102 ....... Biphylliuae .
103 ....... Byturidae .
104 ....... Mycetophagidae .
105 ....... Ciidae ....
107 ....... Prostomidae .
109 ....... Colydiidae
110 ....... Monommmidae ..
III ....... Cephaloidae .
112 ....... Zopheridae ..
115 . ...... Tenebrionidae .
116 ....... Alleculidac ..
117 ....... Lagriidae ..
118 ....... Salpingidae .
119 ....... Myc(eridae ...
120 ....... Pyrochroidae ..
121 ....... Othniidae .
122 ....... Inopeplidae .
123 ....... Oedemeridae ..
124 . ...... Melandryidae ..
125 ....... Mordellidae ..
126 ....... Rhipiphoridae .
127 ....... Meloidae .
128 ....... Anthicidae .
129 ....... Pedilidae .
130 ....... Euglenidae .
131 ....... Cerambycidae .
132 ....... Bruchidae ..
133 ....... Chrysomelidae ..
134 . ...... Nemonychidae ..
135 ....... Anthribidae ..
138 ....... Allocorynidae .
140 ....... Brentidae
141 ....... Platypodidae .
142 ...... Scolytidae .
143 ....... Curculionidac .
144 ....... Stylopidae.
145 ....... Fossil Coleoptera.
146 . ...... Ithyceridae .
-'
. ...... 1991
.. ..... 1982
....... 19H3
. ...... 1986
...... 1991
....... 1979
....... 1983
lMissing numbers are those assigned in the computer program to families not found in the United States and Canada.
Microfiche or printed copies of this publication may be purchased from the
National Technical Information Service, 5285 Port Royal Road, Springfield, VA 22161.
For additional information, contact NTIS at the address above.
All programs and seNices of the U.S. Department of Agriculture are offered on a nondiscriminatory
basis without regard to race, color, national origin, religion, sex, age, marital status, or handicap.
..,
A CATALOG
OF THE COLEOPTERA
OF AMERICA
NORTH OF MEXICO
FAMILY: SILPHIDAE
By
5. B. PECK
DEPT. OF BIOLOGY
CARLETON UNIVERSITY
OTTAWA, ON KIS 5B6
CANADA
UNITED STATES
DEPARTMENT OF
AGRICULTURE
S.E. MILLER
BISHOP MUSEUM
P.O. BOX 19000A
HONOLULU, HI 96817-0916
AGRICULTURE
HANDBOOK
NUMBER 529-28
PREPARED BY
AGRICULTURAL
RESEARCH
SERVICE
September 1993
FOREWORD
__
Many species of beetles are important pests of agricultural crops, stored food products, forests, wood
products and structures, and fabrics. Many other species, in contrast, are beneficial in the biological suppression of pest arthropods and weeds, as well as in the decomposition of plant detritus, animal carcasses,
and dung. Part of our national responsibility to American agriculture is to provide correct identification of
species of American beetles so that appropriate controls can be applied.
Most information about animal species, whether agricultural, biological, or experimental, is filed under
the species' scientific names. These names are therefore the keys to retrieval of such information. Because
some species have been known by several names, a complete listing of these names for each species is necessary.
For the user of scientific names, an up-to-date taxonomic catalog providing currently accepted names
and pertinent bibliographic and distributional data is an indispensable tool. Although taxonomic literature
is constantly changing to reflect current work, the traditional published taxonomic catalog remains static
with updating left to the individual user until it is revised. Production of catalogs in the past has been
laborious with long printing delays resulting in data that are obsolete before being published. However, the
computer now provides the capability of storing. updating, and retrieving taxonomic data; rapid publication through computer-driven typesetting machinery; and a greater degree of currentness and flexibility.
All the fascicles in this catalog of the beetles of America north of Mexico are produced by an original group of computer programs, designed and written during a pilot project by personnel of the
Systematic Entomology Laboratory and the Communication and Data Services Division, Agricultural
Research Service.
R. D. Plowman
Administrator
Agricultural Research Service
v
PREFACE
The Coleoptera, or beeties, are represented in the world by about 220,000 described species, of which
about 24,000 occur in the United States and Canada. A comprehensive taxonomic catalog of beetles for this area
has not been available except the series of world-based "Coleopterorum Catalogus" volumes (1909-present, Junk,
Berlin). The Leng "Catalogue of the Coleoptera of America North of Mexico" (J.D. Sherman, Jr., Mt. Vernon,
NY), which was published in 1920 with supplements to the end of 1947, is a checklist. However, it has served
professional and amateur alike for nearly 60 years as the principal source of scientific names of beetles. Since
1947, many new taxa have been described and many changes in status and nomenclature have appeared in
numerous ,,",cattered publications, but little effort has been made to summarize these changes.
This catalog will supplant the Leng catalog and supply additional essential information. It is produced by
an original suite of storage, retrieval, and printing programs written especially for automated taxonomic catalogs.
The catalog for each family is published as a separate fascicle with its introductory text, bibliography, and
sequence. The publishing of separate fascicles makes data available shortly after they are assembled. Computer
tapes for each fascicle are maintained for updating and necessary reprinting.
The information on each family is the responsibility of the respective author or authors. The editors modify
it only to correct obvious errors and to make it conform to the requirements of the computer programs.
No original proposal for a new name, taxon, status, or classification is given, such data having been
previously published, but new host and distributional data are often listed. The rules of "The International Code
of Zoological Nomenclature" are followed.
The geographic scope of this catalog includes the continental United States, Canada, Greenland, and the
associated continental islands. Names of taxa found only in other regions are excluded. If the range of a species
extends outside these geographic limits, this fact is indicated. On (or inside of) the back cover is a map of the 12
faunal regions based on historical and faunal criteria to simplify distribution recordings. Two-letter Postal Service
style abbreviations are used for States and Provinces, and faunal regions are indicated in each distribution record
by a diagonal line between groups of abbreviations.
It is not the purpose of this catalog to present a complete scheme of higher classification within the order.
The familial makeup is somewhat intermediate between that of R.H. Arnett in "The Beetles of the United States"
(1960-62, Catholic University Press, Washington, DC) and that of R.A. Crowson in "The Natural Classification
of the Families of Coleoptera" (1967, BiddIes Ltd., Guildford, England). Modifications of these two systems are
largely those advocated by J.F. Lawrence based in part on suggestions by taxonomic specialists for certain
families.
Generic groups and higher categories within the family are arranged phylogenetically as indicated by the
author of the particular fascicle, and species group names with their respective synonyms are arranged alphabetically.
Names referable to incertae sedis and nomen dubium are listed separately at the end of the nearest applicable taxon with notations as to their status.
Each available name is followed by its author, date proposed, and page number referring to the complete
bibliographic citation containing the original description. Following each generic name are the type-species and
method of its designation, necessary explanatory notes, and pertinent references on immature stages, taxonomy,
redescription, ecology, and keys. After the specific name entry are the original genus (if different from the
-vii
present placement), type-locality, geographical distribution by State, Province, and broad extralimital units,
explanatory notes, pertinent references to immature stages, taxonomy, redescription, and ecology, depository of
type-specimen and its sex, and hosts.
In addition to the list under the map (on or inside of the back cover) of faunal regions, the following
abbreviations are used in this catalog:
ABBREVIATIO'lS, GENERAL
Amer. Bor.-America Borealis
Amer. Sept.-America Septentrionalis
Automat.-Automatic
C. Amer.-Central America
Co.-County
Cosmop.-Cosmopolitan
Design.-Designated
F.-Female
Holarc.-Holarctic
Isl.-Island
M.-Male
Mex.-Mexico
Monot.-Monotypy
Mus.-Museum
N. Amer.-North America
Orig. des.-Original designation
Preocc.-Preoccupied
S. Amer.-South America
Sp.-Species
Subseq. monot.-Subsequent monotypy
Subsp.-Subspecies
Taut.-Tautonymy
Univ.-University
USA-United States of America
Var.-Variety
W. Ind.-West Indies
MUSEUMS IN THE UNITED STATES AND CANADA
AMNH-American Museum of Natural History,
New York
ANSP-Academy of Natural Sciences, Philadelphia, PA
BPBM-Bernice P. Bishop Museum, Honolulu
BYUC-Brigham Young University, Provo, UT
CASC-California Academy of Sciences, San
Francisco
CISC-University of California, Berkely
CNCI-Canadian National Collections, Ottawa
CUIC-Cornell University, Ithaca, NY
CWOB-C. W. O'Brien Collection, Tallahassee, FL
DHKC-D. H. Kistner Collection, Chico State
College, CA
ELSC-E. L. Sleeper Collection, Long Beach, CA
FMNH-Field Museum of Natural History, Chicago,
IL
FSCA-Florida State Collection, Gainesville
HAHC-H. & A. Howden Collection, Ottawa,
Canada
ICCM-Carnegie Museum, Pittsburgh, PA
INHS-Illinois Natural History Survey, Urbana
JGEC-J. G. Edwards Collection, San Jose, CA
KMFC-K. M. Fender Collection, McMinnville,
OR
KSUC-Kansas State University, Manhattan
LACM-Los Angeles County Museum, CA
LSUC-Louisiana State University, Baton Rouge
MCZC-Museum of Comparative Zoology,
Harvard University, Cambridge, MA
MSUC-Michigan State University, East Lansing
NCSM-North Carolina State University, Raleigh
NYSM-New York State Museum, Albany
OSEC-Oklahoma State University, Stillwater
OSUC-Ohio State University, Columbus
OSUO-Oregon State University, Corvallis
IAbbreviations for U. S. and Canadian museums abridged from Arnett. R. H .. Jr.. and Samuelson. G. A., 1969, "Directory of
Coleoptera Collections of North America (Canada Through Panama)," Cushing-Malloy, Ann Arbor, MI, 123 pp.
viii
PMNH-Peabody Museum, Yale University,
New Haven, CT
PSUC-Pennsylvania State Museum, University
Park
PURC-Purdue University, West Lafayette, IN
RUIC-Rutgers University, New Brunswick, NJ
SEMC-Snow Museum, University of Kansas,
Lawrence
SJSC-San Jose State College, CA
SLWC-S. L. Wood Collection, Provo, UT
SMSH-Stovall Collection, University of
Oklahoma, Norman
TAMU-Texas A. & M. University, College
Station
UCDC-University of California, Davis
UICM-University of Idaho, Moscow
UMMZ-University of Michigan, Ann Arbor
UMRM-University of Missouri, Columbia
USNM-U.S. National Museum of Natural
History, Washington, DC
WSUC-Washington State University, Pullman
MUSEUMS IN FOREIGN COUNTRIES
BMNH-British Museum (Natural History),
London
GUHC-Glasgow University, Hunterian
College, Scotland
HMO X-Hope Museum, Oxford, England
IPZE-Institut Pflanzenschutzforschung
Zweigstelle, Eberswald, East Germany
IRSB-Institut Royal Sciences Belgique,
Brussels
MFNB-Museum fUr Naturkunde (Humboldt),
Berlin
MGFT-Museum G. Frey, Tutzing, Munich,
West Germany
MHNL-Museum d'Histoire Naturelle, Lyon,
France
MNHP-Museum National d'Histoire Naturelle,
Paris
MNSL-Museum of Natural Sciences, Leipzig,
East Germany
MZBS-Museum Zoologia, Barcelona, Spain
NHRS-Naturhistoriske Riksmuseet, Stockholm
NMPC-Narodni Museum, Prague,
Czechoslovakia
SCUT-Spinola College, University of Turin,
Italy
SMTD-Staatliches Museum fUr Tierkunde,
Dresden, East Germany
UNAM-Universidad Nacional Autonorna,
Mexico City
UZMC-University Zoological Museum,
Copenhagen, Denmark
UZMH-U niversity Zoological Museum,
Helsinki, Finland
ZMAS-Zoological Museum, Academy of
Sciences, Leningrad
ZMPA-Zoological Museum, Polish Academy of
Sciences. Warsaw
ZMUL-Zoological Museum, University of
Lund, Sweden
ZMUM-Zoological Museum, University of
Moscow
ZSBS-Zoologische Sam m I u n g Bayerischen
Staates, Munich, West Gennany
ix
ACKNOWLEDGMENTS
We are indebted to many individuals who contributed to the planning and development of this catalog. We
are especially grateful to the foJlowing specialists who helped to make it as complete and accurate as possible:
Richard H. Foote, Systematic Entomology Laboratory (SEL), Agricultural Research Service (ARS), for his suggestions, guidance, and encouragement~ C. W. Sabrosky, SEL, for valuable counsel on nomenclatural problems:
1. F. Lawrence, Division of Entomology, Commonwealth Scientific and Industrial Research Organization. Canberra, Australia, for his recommendations on higher categories; and more than 50 coleopterists in Canada. the
United States, and Mexico for voluntarily contributing information about their specialty groups.
We wish to acknowledge the extensive computer-programming and editing-system support provided by
members of the former Communications and Data Services Division of ARS during the early years of this project.
We also thank Elaine Jamison for the data entry necessary for each fascicle.
R. D. Gordon, editor in chief
J. C. Barch, associate editor
F. C. Thompson, computer program advisor
Systematic Entomology Laboratory, Agricultural Research Service
Washington, DC
Editorial Board
J. M. Campbell, Biosystematics Research Institute, Agriculture Canada, Ottawa, ON
T. L. Erwin, Department of Entomology, Smithsonian Institution, Washington, DC
H. F. Howden, Department of Biology, Carleton University, Ottawa, ON
J. M. Kingsolver, Florida State Collection of Arthropods, Gainesville, FL
P. J. Spangler, Department of Entomology, Smithsonian Institution, Washington, DC
T. J. Spilman, Systematic Entomology Laboratory, ARS, Washington, DC
R. E. White, Systematic Entomology Laboratory, ARS, Washington, DC
CONTENTS
Family Silphidae ............................................................................................... I
Tribe Agyrtini .............................................................................................. 2
Tribe Silphini ............................................................................................... 3
Tribe Nicrophorini ...................................................................................... ,6
Bibliography .............................................................................................. 10
Index .......................................................................................................... 23
x
-
Family SILPHIDAE
By S. B. Peck and S. E. Miller
Silphidae, the so called carrion beetles, are from 5 to 35 mm long, are often brightly colored, and usually are found feeding on dead animals or sometimes fly larvae. Some, however,
seem more associated with decaying fungi or vegetable matter, and one genus is strictly phytophagous. The attractive red or orange and black Nicrophorus species are called burying beetles because they can rapidly excavate the soil from under a dead mouse or other small animal
until it sinks into the ground. Parental care of the larvae makes Nicrophorus one of the most
social of all beetles (work of Pukowski, abstracted in Balduf, 1935; Milne and Milne, 1944, 1976).
The Silphini do not show such parental care. Necrophilus are carrion feeders, Lyrosoma are
littoral, and Pteroloma and Apteroloma are montane and probably predatory. Little is known of
the biology of the Agyrtini.
Limited aspects of silphid biology in restricted areas have been studied, but some reports
are unreliable because of confused nomenclature. Olfactory sensitivity, of primary importance
in locating carrion, is acute, and in electrophysiological studies, the sensory receptors and specific decay compounds that activate them have been identified. Many species that live in one
area can "divide" the carrion resource by differing in seasonality, diel periodicity, Or habitat
preferences (Anderson, 1982c; Shubeck, 1971; Wilson and Fudge, 1984; Wilson and Knollenberg,
1984; Wilson, Knollenberg, and Fudge, 1984). Carrion-baited pitfall traps are best for collecting
silphids. They often can be used as experimental systems to answer specific ecological questions
(Scott, et al., 1987; Shubeck, 1976a, 1976b).
Silphids, biologically important in maintaining ecosystem health and productivity, aid in
the recycling of organic matter. While engaged in this process, they reduce the frequency and
suitability of carrion as a site of fly breeding and development. Their feeding may destroy p0tential infection foci of pathogenic bacteria or parasitic worms. Rabies virus, but not anthrax
bacteria, is inactivated in the gut of Nicrophorus. Silphids phoretically carry many species of
mites and nonpathogenic nematodes as commensals.
There has been much confusion in the use of generic and specific names for the silphids of
North America. For instance, the names Nicrophorus and Necrophorus have interchangeably
been used by authors, but Nicrophorus is considered correct. For the 15 species of Nicrophorus
of the United States and Canada, over 110 names have been applied, inclUding about 65 used as
variations and aberrations with no taxonomic status. Reviews and revisions (Madge, 1980; and
Miller and Peck, 1979), in combination with this catalog, should bring the fauna into evolutionary perspective with the world fauna and should help establish nomenclatorial stability. A detailed study of adult and larval characters of Agyrtini, with about 45 world species (Lawrence,
1982; Lawrence and Newton, 1982; and Newton, unpublished), shows that this tribe should be
elevated to family status and placed near the Leiodidae. Lawrence and Newton (1982) also indicate that the remaining two tribes-Silphini with about 90 world species and Nicrophorini with
about 68 world species-could be united as a subfamily of the Staphylinidae. Madge (1980) uses
a different system (Silphinae, Nicrophorinae, Trigonurinae, and Agyrtinae) for the Silphidae,
which he regards as a valid family. Other important general works on North American silphids
are by Anderson (1982a), Anderson and Peck (1985), Arnett (1944), Hatch (1927 a, 1927b, 1928,
1957), Madge (1980), Miller and Peck (1979), Peck and Anderson (1985), Peck and Kaulbars
(1988), and Peck and Miller (1982). Keys to identify the genera and species are in Anderson and
Peck (1985) and Peck (1989).
Several species recorded in the literature as occurring in North America represent unestablished introductions of palaearctic species or mislabelings. These species, which are not included in the catalog, are Dendroxena quadrimaculara (Scopoli, 1772), Nicrophorus germanicus
(Linnaeus, 1758), N. humator (Gleditsch, 1767), N. vespillo (Linnaeus, 1758), Silpha tristis.(Illiger, 1798), and S. tyrolensis (Laicharting, 1781). Additional information may be found in
Arnett (1944), Hatch (1927b), and Miller and Peck (1979). Nicrophorus grandior Angell (1912)
was proposed for a specimen of N. germanicus (lectotype by Peck and Miller, 1982) and for a
specimen of N. humator, both mistakenly labeled "California."
Eurasian synonyms of holarctic genera and species (not used in the North American literature and not included here) may be found in Hatch (1928). We have not included aberrations
and varieties named by Hatch (1957) because he stated that these "do not refer to discrete pop-
1
2
COLEOPl'ERA CATALOG
ulations but to segregates of the same." Varietal names proposed by Portevin are included
'
however, as they may have been meant in a subspecific sense.
R. S. Anderson, R. B. Madge, and A. F. Newton contributed to the preparation of this catalog.
This manuscript was received October 1981, modified September 1986, and amended November 1992.
Tribe AGYRTINI
Genus NECROPHILUS Latreille
Necrophilus Latreille, 1829: 500. Type-species: Silpha subterranea Dahl (monot.).
Necrobius Gistel, 1834: 147. Type-species: Silpha subterranea Dahl (monot.).
hydrophiloides Guerin-Meneville, 1835: pI. 17, fig. 12. ''I'ile de Norfolk, dans I'Amerique
boreale"; AKI BC WA OR IDI CA. First published as a named figure in the text
(Guerin-Meneville, 1844: 61).
ater Motschulsky, 1845: 363. AK.
IMMATURE STAGES: Blaisdell, 1901: 263.
pettiti Horn, 1880: 243. ON: Grimsby; MI ONI MO IL IN OH KY I NY VAl LA AL TN GA SC
NC FL. The lectotype was designated in Peck, 1981: 181.
MeZC.
F.
ECOLOGY: Peck, 1981: 182.
TYPE DEPOSITORY:
SEX OF TYPE:
Genus IPELATES Reitter
Ipelates Reitter, 1885: 89, 90. Type-species: Hadrambe latissimus Reitter (monot.).
Pelates Horn, 1880: 244 (preoccupied by Cuvier and Valenciennes, 1829). Type-species:
Necrophilus latus Mannerheim (monot.).
Sphaeroloma Portevin, 1905: 422. Type-species: Sphaeroloma sikkimensis Portevin (orig.
des.).
Pelatines Cockerell, 1906: 240 (replacement name for Pelates Horn). Type-species:
Necrophilus latus Mannerheim (automat.).
Brachyloma Portevin, 1914: 214. Type-species: Brachyloma curtum Portevin (design. by
Hatch, 1928: 70) = striatipennis (Lewis).
Necrophilodes Champion, 1923: 49. Type-species: Necrophilus castaneicolor Champion
(orig. des.). Misspelled Necrophiloides by Hatch, 1927b, p. 333 and Hlisnikovsky,
1963, p. 314.
TAXONOMY: SchawaIler, 1983: 101.
latus Mannerheim, 1852: 331 (Necrophilus). AK: Sitkha; AK NTI BC WA ORI ABI CA.
Genus AGYRTES I"roelich
Agyrtes Froelich, 1799: 15 (nominate subgenus extra-limital). Type-species: Mycetophagus
castaneus Fabricius (design. by Latreille, 1810: 427).
TAXONOMY: Hlisnikovsky, 1964b: 275; Peck, 1974: 501.
Subgenus AGYRTECANUS Reitter
Agyrtecanus Reitter, 1901: 102. Type-species: Agyrtes bicolor Laporte (monot.).
Lendomus Casey, 1924: 184. Type-species: Lendomus politus Casey (monot.)
(LeConte).
=
longulus
longulus LeConte, 1859a: 282 (Necrophilus). CA: San Francisco, Table Mt.; AKI BC WA OR JDI
CA. The lectotype was designated by Peck, 1974: 501.
TYPE DEPOSITORY: Mezc.
politus Casey, 1924: 185 (Lendomus). PQ: Anticosti lsI. Error of nomenclature and
distribution from erroneous specimen data (Peck, 1974: 504),
TYPE DEPOSITORY: USNM.
TAXONOMY:
Peck, 1974: 501.
SILPlllDAE
-
3
similis Fall, 1937: 29. CA: Pasadena; CA.
TYPE DEPOSITORY: MCZC.
TAXONOMY: Peck, 1974: 504.
Genus LYROSOMA Mannerheim
Lyrosoma Mannerheim, 1853: 174. Type-species: Pteraloma pallidum Eschscholtz (design. by
Hatch, 1928: 71).
TAXONOMY:
Hlisnikovsky, 1964a: 39.
opacum Mannerheim, 1853: 175. AK: Atkha, Aleutian Isl.; St. Paul, Pribilof Isl.; AK; Other:
Siberia.
HOST: Supposedly living in rotting kelp, at times completely submerged by tides (Van Dyke, 1921:
161). Also recorded from nests of maritime birds (Ferris, 1918: 125).
Genus PTEROLOMA Gyllenhal
Pteroloma Gyllenhal, 1827: 418. Type-species: Harpalus forsstramii Gyllenhal (monot.).
Adolus Fischer von Waldheim, 1828: 242. Type-species: Adolus brunneus Fischer von
Waldheim (monot.) = forsstromii (Gyllenhal).
Holocnemis Schilling, 1829: 93. Type-species: Holocnemis gravenhorstii Schilling (monot.)
= forsstramii (Gyllenhal).
TAXONOMY:
Madge, 1980: 354.
nebrioides Brown, 1933: 213. AB: Waterton; BC ID/ AB MT.
TYPE DEPOSITORY: eNCL
SEX OF TYPE:
F.
EcOLOGY: Johnson. 1987: 381.
Genus APTEROLOMA Hatch
---.
Apteroloma Hatch, 1927a: 12 (as subgenus). Type-species: Necrophilus tenuicorne LeConte (orig.
des.).
TAXONOMY: Van Dyke, 1928: 19; Bolivar y Pieltain and Hendrichs, 1972: 207; Miller and Peck, 1979:
89.
KEYS: Van Dyke, 1928: 21; Bolivar y Pieltain and Hendrichs, 1972: 210; Miller and Peck, 1979: 98.
arizonicum Van Dyke, 1928: 24 (Pteraloma). AZ: near Prescott; AZ NM TX.
TYPE DEPOSITORY:
CASCo
caraboides Fall, 1907: 235 (Pteroloma). W A: Wenatchee; CA: Mt. San Antonio.
TYPE DEPOSITORY: MCZC.
tahoecum Fall, 1927: 136 (Pteraloma). CA: Lake Tahoe; OR/ CA NV.
TYPE DEPOSITORY: MCZC.
TYPE: F.
SEX OF
tenuicornis LeConte, 1859b: 84 (Necraphilus). WA: Puget Sound.
TYPE DEPOSITORY: MCZC.
SEX OF TYPE:
TAXONOMY:
F.
Peck and Miller, 1982: 154.
Tribe SILPHINI
Genus NECRODES Leach
Necrodes Leach, 1815: 88. Type-species: Silpha littoralis Linnaeus (design. by Curtis, 1830: pI.
334).
Asbolus Bergroth, 1884: 229 (unnecessary replacement name for Necrodes Leach). Typespecies: Silpha littoralis Linnaeus (automat.).
Protonecrades Portevin, 1922: 508. Type-species: Silpha surinamensis Fabricius (design. by
Hatch, 1928: 123).
TAXONOMY:
Madge, 1980: 354.
surinamensis Fabricius, 1775: 72 (Silpha). southern (North) America; BC WA OR IDI AB SK
MB MT SDI MN WI MI ON PQ/ NK PE NS NFl UTI NE KS IA MO IL IN OH
4
COLEOPl'ERA CATALOG
~mumoo~oc~~~~n~ru~n~~u~
AL TN GA SC NC FL.
TYPE DEPOSITORY: GUHC.
IMMATURE STAGES: Dorsey, 1940: 130; Ratcliffe, 1972: 366.
TAXONOMY: Ratcliffe, 1972: 364.
EcoLOGY: Ratcliffe, 1972: 380; Eisner and Meinwald. 1983: 358.
Genus OXELYTRUM Gistel
Oxelytrum Gistel, 1848: 190. Type-species: Oxelytrum aequinoctiale Gistel (orig. des.) =
cayennense(SturrrU.
Oxelytrium (error). (In Gistel, 1857: 545, 566).
Oxelytron (error). (In Gistel, 1857: 546).
Hyponecrodes Kraatz, 1876: 357. Type-species: Silpha cayennensis Sturm (design. by
Madge, 1980: 355).
Katanecrodes Schouteden, 1905: 195. Type-species: Silpha apicalis BruHe (design. by
Hatch, 1928: 113).
Paranecrodes Portevin, 1921b: 81. Type-species: Necrodes biguttatus Philippi (monot.).
TAXONOMY: Madge. 1980: 357.
discicolle Brulle, 1836: 75, pI. 5, fig. 4 (Silpha). Bolivia: near Cochabamba; TX; Mex., C. Amer.,
S. Amer.
analis Chevrolat, 1843: 26 (Necrodes). Mex.: "Orixaba".
discretus Portevin, 1921b: 84 (Hyponecrodes) (as variety). "Guatemala, Colombie,
Venezuela, Brezil, Paraguay". The stated type-locality also includes "Argentine et
Perou" (Portevin, 1921b: 84).
TAXONOMY: Davis, 1980: 249: Peck and Anderson, 1985: 263.
Genus THANATOPHILUS Leach
Thanatophilus Leach, 1815: 89. Type-species: Silpha sinuata Fabricius (monot.).
Pseudopelta Bergroth, 1884: 229 (unnecessary nomen nudum). Type-species: Silpha
sinuata Fabricius (monot.).
Philas Portevin, 1903: 331. Type-species: Silpha truncata Say (monot.).
Silphosoma Portevin, 1903: 333. Type-species: Silpha metallescens Fairmaire (monot.).
Chalcosilpha Portevin, 1926: 31. Type-species: Silpha micans Fabricius (design. by Hatch,
1928: 79).
TAXONOMY: Schawaller. 1981: 3.
coloradensls Wickham, 1902: 180 (Si/pha). co: Argentine Pass, nr. Georgetown, over 12,000 ft;
AKI BCI MTI WY UT COl NM.
TYPE DEPOSITORY:
USNM.
SEX OF TYPE: M.
obalskii Portevin, 1920b: 507 (Silpha). BC: Lake Teslin.
EcOLOGY:
Bjorkman and Hatch, 1939: 96; Peck and Anderson. 1982: 112.
lapponicus Herbst, 1793: 209, pI. 52, fig. 4 (Silpha). Lappland; AK YT NT GLI BC WA OR IDI
~gmMT~~~MWOO~~~~wym~~~u
ILl PA DCI NH RII AZ NM; Mex., Holarc. See Hatch (1928: 80) for Eurasian
synonyms.
caudata Say, 1823: 192 (Silpha). "Missouri Territory". The neotype was not designated
(Peck and Miller, 1982: 154).
tuberculata Germar, 1824: 81 (Silpha). "North America".
californica Mannerheim, 1843: 253 (Si/pha). CA.
IMMATURE STAGES: Dorsey, 1940: 126.
EcoLOGY: Korobeinikov and Esyunin, 1984: 40.
sagax Mannerheim, 1853: 173 (Silpha). AK: Kaktnu River, Kenai Peninsula; AK YT NTI BC
WA OR IDI AB SK MB ND SDI ON PQI WY CO. The species was long considered
a junior synonym of Thanatophilus trituberculatus (Kirby, 1830).
TAXONOMY:
Miller and Peck, 1979: 91.
-
-
SILPHIDAE
5
trituberculatus Kirby, 1837: 101 (Oiceoptoma). "Boreal America"; AK NT/ AB SK MB/ ON.
The specific type-locality is between New York and Cumberland House (SK).
IMMATURE STAGES: Anderson, 1987: 34.
TAXONOMY: Miller and Peck, 1979: 91.
truncatus Say, 1823: 193 (Silpha). "eastern Colorado"; COl NE KSI AZ NM TX; Mex. The
neotype was designated by Peck and Miller, 1982: 154.
TYPE DEPOSITORY:
MeZC.
SEX OF TYPE: M.
Genus OICEOPTOMA Leach
Oiceoptoma Leach, 1815: 89. Type-species: Silpha thoracica Linnaeus (monot.).
Oeceoptoma Agassiz, 1847: 256 (unjustified emendation). Type-species: Silpha thoracica
Linnaeus (monot.).
Isosilpha Portevin, 1920a: 398. Type-species: Eusilpha hypocrita Portevin (monot.).
IMMATURE STAGES: Dorsey, 1940: 123.
inaequale Fabricius, 1781: 87 (Silpha). "North America"; ND SDI MN WI MI ON PQI COl KS
MO IL IN OH KY I NY PA NJ MDI MA RI CTI TX OKI AR LA AL TN GA NC
FL.
noveboracense Forster, 1771: 17 (Silpha). "Northern North America"; AB SK MB MT ND SDI
MN WI MI ON PQI NS/ WYI IA MO IL IN OH KY/ NY PA NJ DC WV VAl ME
NH VT MA RI CT/ TX OKI AR MS TN GA NC.
marginalis Fabricius, 1777: 215 (Silpha). "Boreal America".
EcOLOGY:
-
Anderson, 1989: 18.
rugulosum Portevin, 1903: 333 (Silpha). GA: Savannah; IL INI NY I TX OK/ AL GA SC NC FL.
This name was erroneously put into synonymy under O. inaequale by Hatch,
1927b: 343. The lectotype was designated by Peck and Kaulbars, 1988: 50.
TYPE DEPOSITORY:
MNHP.
Genus NECROPHILA Kirby and Spence
Necrophila Kirby and Spence, 1828: 509. Type-species: Silpha americana Linnaeus (orig. des.).
Necrobora Hope, 1840: 151 (unnecessary name). Type-species: Silpha americana Linnaeus
(orig. des.).
Necrotropha Gistel, 1848: 121. Type-species: Silpha americana Linnaeus (monot.).
Eusilpha Semenov-Tian-Shanskij, 1891: 299. Type-species: Silpha japonica Motschulsky
(design. by Hatch, 1928: 111).
Calosilpha Portevin, 1920a: 396. Type-species: Silpha ioptera Kollar and Redtenbacher
(design. by Hatch, 1928: 113).
Deutosilpha Portevin, 1920a: 396. Type-species: Silpha rufithorax Wiedemann (monot.).
Chrysosilpha Portevin, 1921a: 538. Type-species: Silpha formosa Laporte (design. by
Hatch, 1928: 112).
Deuterosilpha (error). (In Hatch, 1928: 112).
americana Linnaeus, 1758: 360 (Silpha). "America" (north); MB SDI MN WI MI ON PQI NSI
IA MO IL IN OH KY/ NY PA NJ DE MD WV VAl NH MA RI CTI TX OK/ AR
AL TN GA SC FL.
peltata LeConte, 1853: 279 (Silpha). PA. Name first published as Scarabaeus peltatus by
Catesby (1748: pI. 10, fig. 7) and repeated by Edwards (1771), but suppressed by Int.
Comm. Zool. Nomen. Opinion 259. Name first validated by LeConte (1853: 279).
affine Kirby, 1837: 103 (Oiceoptoma). NS.
caruui£nse Kirby, 1837: 104 (Oiceoptoma). ON: Lake Huron.
terminatum Kirby, 1837: 103 (Oiceoptoma). NS.
IMMATURE STAGES: Dorsey, 1940: 126.
EcoLOGY:
Fisher and Tuckerman. 1986: 21; Anderson. 1989: 18.
Genus HETEROSILPHA Portevin
Heterosilpha Portevin, 1926: 83. Type-species: Silpha ramosa Say (design. by Hatch, 1928: 100).
6
COLEOPI'ERA CATALOG
TAXONOMY:
Miller and Peck, 1979: 92.
aenescens Casey, 1886: 171 (Silpha). CA: San Francisco; OR/ CA; Mex. The lectotype was
designated by Miller and Peck, 1979: 93.
TYPE DEPOSITORY: USNM.
SEX OF TYPE: M.
REDESCRIPTION: Miller and Peck, 1979: 93.
ramosa Say, 1823: 193 (Silpha). "the upper Missouri" (River); NT/ BC WA OR ID/ AB SK MB
MT ND SD/ MN WI MI ON/ CA NV/ WY UT COl NE/ AZ NM; Mex. The
neotype was designated by Miller and Peck, 1979: 93.
TYPE DEPOSITORY: Meze.
SEX OF TypE:
M.
cervaria Mannerheim, 1843: 252 (Silpha). CA.
IMMATURE STAGES: Dorsey, 1940: 129; Brewer and Bacon, 1975: 787.
REDESCRIPTION: Miller and Peck, 1979: 93.
EcOLOGY: Brewer and Bacon. 1975: 786.
Genus ACL YPEA Reitter
Aclypea Reitter, 1885: 81. Type-o;pecies: Pettis undata Mueller (design. by Hatch, 1928: 93). This
name is used in reference to Blitophaga following Seidlitz (1888: 311), who is
regarded as first reviser in accordance with Article 24 (a) (i) of the International
Code of Zoological Nomenclature (Miller and Peck, 1979: 90).
Blitophaga Reitter, 1885: 82. Type-species: Silpha opaca Linnaeus (design. by Hatch,
1928: 95).
TAXONOMY: Anderson and Peck, 1984: 248.
bituberosa LeConte, 1859c: 6 (Silpha). WY: near Ft. Bridger; NT/ BC WA OR ID/ AB SK MB
MT ND/ CAl WY UT COl NE KS.
TYPE DEPOSITORY: MCZe.
SEX OF TYPE:
F.
IMMATURE STAGES: Cooley, 1917: 97; Anderson and Peck, 1984: 254.
TAXONOMY: Peck and Miller. 1982: 154.
EcoLOGY: Cooley, 1917: 95; Anderson and Peck, 1984: 251.
HOST: Adults and larvae feed on Monolepis nuttalliana (Chenopodiaceae) and related native and
introduced plants, such as beet, sugarbeet, spinach, pumpkin, squash. etc. (Hatch, 1957: 10).
opaca Linnaeus, 1758: 361 (Silpha). Europe; AK YK NT; Other: Eurasia. Older records of A.
bituberosa are often erroneously under this name (Miller and Peck, 1979: 90).
ECOLOGY: Anderson, 1989.
Tribe NICROPHORINI
Gl!nus NICROPHORUS Fabricius
Nicrophorus Fabricius, 1775: 71. Type-species; Silpha vespillo Linnaeus (design. by Latreille,
1810: 427). Many species of Nicrophorus were described under the unjustified
emendation Necrophorus (Herman, 1964; 5; Madge, 1980: 357). Designation of the
lectotype by Latrielle was for the emendation Necrophorus.
Necrophorus Thunberg, 1789: 7 (unjustified emendation). Type-species: Silpha vespillo
Linnaeus (design. by Latreille, 1810: 427).
Necrophagus (error) Leach. 1815: 88.
Cyrtoscelis Hope, 1840: 149. Type-species: Silpha vespillo Linnaeus (orig. des.).
Acanthopsilus Portevin, 1914: 223. Type-o;pecies: Necrophorus concolor Kraatz (monot.).
Necrocharis Portevin, 1923: 68. Type-species: Silpha carolina Linnaeus (monot.).
Eunecrophorus Semenov-Tian-Shanskij, 1933: 152. Type-species: Nicrophorus americanus
Olivier (orig. des.).
Necrocleptes Semenov-Tian-Shanskij, 1933: 153. Type-species: Silpha humator Gleditsch
(orig. des.).
Necropter Semenov-Tian-Shanskij, 1933: 154. Type-species: Necrophorus investigator
Zetterstedt (orig. des.).
-
SILPHIDAE
7
Stictonecropter Semenov-Tian-Shanskij, 1933: 154. Type-species: Necrophorns pustulatus
Herschel (orig. des.).
Neonicrophorus Hatch, 1946: 99. Type-species: Silpha germanica Linnaeus (orig. des.).
IMMATURE STAGES: Anderson, 1982a: 256.
TAXONOMY: Portevin, 1926; Hatch, 1927a; Herman, 1964: 6; Madge, 1980: 357.
EcoLOGY: Milne and Milne. 1944: 312, and 1976: 87; Wilson, 1983: 57; Anderson, 1982a: 249, and
1982c: 1315.
americanus Olivier, 1790: 6, pI. 1, fig. 3. "North America"; MN WI MI ON PQI NSI NE KS IA
MO IL IN OH KYI NY PA NJ MD DC VAl ME NH MA RI CTI AR MS TN GA
SCNC.
grandis Fabricius, 1792: 247. "North America".
virginicus Froelich, 1792: 123. VA.
ECOLOGY: Davis, 1980: 248; Anderson, 1982b: 363; Hecht, 1988: 39618; Korol et aI., 1989: 167.
carolinus Linnaeus, 1771: 530 (Bilpha). "Carolina"; ABI WY COl NE KS ILl NY MD VAl AZ
NM TX OKI LA MS AL TN GA SC NC FL.
mediatus Fabricius, 1801: 334 (Nicrophorus). "Carolina".
mysticallis Angell, 1912: 307. AZ: Tuba.
TYPE DEPOSITORY:
USNM.
defodiens Mannerheim, 1846: 513. AK: Sitka; AK YK NTI BC WA OR IDI AB SK MB MT SDI
MN WI MI ON QBI NK PE NS NFl CA NVI WY COl NY NJ WVI ME NH VT
MAl TNNC.
pollinctor LeConte, 1854: 19 (junior homonym of Nicrophorns pollinctor Mannerheim,
1853). OR, WA. The lectotype was designated by Peck and Miller, 1982: 152.
--
TYPE DEPOSITORY:
SEX OF TYPE: M.
MCZC.
conversator Walker, 1866: 320. BC: "Slecy Station".
plagiatus Motschulsky, 1869: 352. CA.
lateralis Portevin, 1903: 330 (as variety of vespilloides). CA.
IMMATURE STAGES: Leech, 1934: 37.
TAXONOMY: Leech, 1937: 156; Miller and Peck, 1979: 95; Peck and Miller, 1982: 152.
EcoLOGY: Leech, 1934: 36.
guttula Motschulsky, 1845: 53. AK: Sitka; BC WA OR IDI AB SK MT ND SDI MNI CA NVI
WY UT COl NEI AZ NM; Mex.
hecate Bland, 1865: 382. "Colorado Territory". The lectotype was designated by Miller
and Peck, 1979: 97.
TYPE DEPOSITORY:
ANSP.
punctostriatus Pierce, 1949: 66 (as subspecies of guttula). CA: Los Angeles Co., Rancho La
Brea. The Rancho La Brea specimens are of the Pleistocene Age (Pierce, 1949: 55).
TYPE DEPOSITORY:
LACM.
immaculosis Hatch, 1957: 15. CA. This name was used as an aberration by Hatch, 1927b:
362 but as a subspecies by Hatch, 1957: 15; it takes the authorship and date of use
as a subspecies (International Code of Zoological Nomenclature, Art. lOb: p. 10).
IPZE.
TAXONOMY: Miller and Peck, 1979: 97; Peck and Miller, 1982: 153; Anderson and Peck, 1986: 286.
TYPE DEPOSITORY:
hybridus Hatch and Angell, 1925: 216. MB: Aweme; BC WA OR IDI AB MB MT NDI MNI WY
UT COl NE IAI AZ NM.
TYPE DEPOSITORY;
USNM.
investigator Zetterstedt, 1824: 151. Sweden; AK YTI BC WA OR IDI AB MTI MN ON PQI NS
NFl WY UT COl MEl AZ NM; Holarc. See Hatch (1928: 139) for Eurasian
synonyms.
maritimus Guerin-Meneville, 1835:, pI. 17, fig. 8. AK: Sitcha lsI. This was first published
by Guerin-Meneville in 1835 as a named figure; text (p. 60) published in 1844.
melsheimeri Kirby, 1837: 97. "from New York to Cumberland-House" (SK).
TYPE DEPOSITORY:
BMNH.
8
COLEOPl'ERA CATALOG
particeps Fischer von Waldheim, 1844: 139 (as variety of Nicrophorus lunatus). AK: Sitka.
Heller (1896: 23) shows Turcoman, the locality cited by Fischer von Waldheim
(1844: 139), to be in error.
aleuticus Gistel, 1848: 190. AK: Sitka.
infodiens Mannerheim, 1853: 170. AK: Unalaschka lsI.
pollinctor Mannerheim, 1853: 169. AK: Atkha lsI. Peck and Miller (1982:152) designated
the lectotype.
confossor LeConte, 1854: 20. WA: Prairie Paso.
TYPE DEPOSITORY:
SEX OF TYPE: M.
MeZC.
TAXONOMY: Arnett, 1947: 110; Peck and Miller, 1982: 152.
marginatus Fabricius, 1801: 334. "America"; BC WA OR IDI AB MT ND SDI MN WI MI ON
PQI CAl WY UT COl NEKS IA MO IL IN KYI NY PA NJ MD VAl ME MA RI
CTI AZ NM TX OKI AR LA TN NC FL; Mex.
requiescator Gistel, 1848: 190 (synonymy following Hatch, 1928: 147). "North America".
montezumae Matthews, 1888: 92. Mex.
TYPE DEPOSITORY: BMNH.
labreae Pierce, 1949: 63, figs. 4-10 (as subspecies of guttula). CA: Los Angeles Co., Rancho
La Brea. The Rancho La Brea specimens are of the Pleistocene Age (Pierce, 1949:
55). Miller and Peck (1979: 96) designated the lectotype.
TYPE DEPOSITORY: LACM.
latifrons Pierce, 1949: 67, fig. 14 (as subspecies of investigator). CA: Los Angeles Co.,
Rancho La Brea. The Rancho La Brea specimens are of the Pleistocene Age
(Pierce, 1949: 55).
TYPE DEPOSITORY:
LACM.
mckittricki Pierce, 1949: 66, fig. 11. CA: Kern Co., McKittrick. The McKittrick specimens
are of the Pleistocene Age (Pierce, 1949: 55).
TYPE DEPOSITORY: LACM.
obtusiscutellum Pierce, 1949: 67, fig. 12. CA: Los Angeles Co., Rancho La Brea. The
Rancho La Brea specimens are of the Pleistocene Age (Pierce, 1949: 55).
TYPE DEPOSITORY: LACM.
TAXONOMY: Miller and Peck, 1979: 96.
mexicanus Matthews, 1888: 91. Mex.; UT COl KS JAI AZ NM; Mex.
EcoLOGY: HaHner et al., 1982: 51, and 1983: 649.
nigrita Mannerheim, 1843: 251. CA; BC WA ORI CA. The name is used as a noun in apposition;
it is masculine singular of Nigritae, an African people living along the Niger
River.
ruficornis Motschulsky, 1869: 352 (Necrophorus). CA.
alpha Pierce, 1949: 67, fig. 13 (as subspecies of investigator). CA: Los Angeles Co., Rancho
La Brea. The Rancho La Brea specimens are of the Pleistocene Age (Pierce, 1949:
55). Miller and Peck (1979: 95) designated the lectotype.
TYPE DEPOSITORY: LACM.
TAXONOMY: Miller and Peck. 1979: 95-6.
obscurus Kirby, 1837: 97. "from New York to Cumberland-House" (SK); IDI AB SK MB MT
ND SDI MNI UTI NE IA MO.
melsheimeri LeConte, 1853: 275. NB, UT.
orbicollis Say, 1825: 177. "United States"; SDI MN WI MI ONI NSI NE KS IA MO IL IN OH
KYI NY PA NJ MD DC WV VAl ME NH MA RI CTI OKI AR AL TN GA SC NC
FL. Peck and Miller (1982: 153) designated a neotype.
TYPE DEPOSITORY:
SEX OF TYPE: M.
MCZC.
hallii Kirby, 1837: 98. NS, MA.
-
9
SILPHIDAE
quadrisignatus Laporte, 1840: 1. "Amerique du Nord".
TAXONOMY: Peck and Miller. 1982: 153.
EooLOGY:
Scott, 1989: 133; Trumbo. 1991: 82.
pustulatus Herschel, 1807: 271. "North America"; MN WI MI ON PQI KS IA MO IL IN OHI
m~m~oc~~m~ru~n~o~~n.
bicolon Newman, 1838: 385. "North America".
tardus Mannerheim, 1853: 170. AK: Sitka lsI.
sayi Laporte, 1840: 2. "North America"; AB SK MBI MN WI MI ON PQI NFl IA IL INI NY
PA NJ MD ~I ME ~ MA CTI OKlO AL TN NC.
lunulatus Gistel, 1848: 189 (synonymy following Hatch, 1928: 129). "North America".
lunatus LeConte, 1853: 277 (preoccupied Fischer von Waldheim, 1842). NY, GA. Peck and
Miller (1982: 152) designated the lectotype.
TYPE DEPosITORY:
MeZC.
SEX OF TYPE: M.
luniger Harold, 1868: lO4 (replacement name for lunatus LeConte).
TAXONOMY:
Peck and Miller, 1982:
m.
152.
tomentosus Weber, 1801: 47. "America"; MB MT SDI MN WI MI ON PQI NB NSI COl NE KS
IA MO IL IN OHI NY PA NJ MD 00 VAl ME NH MA CTI NM n OKI AR AL
TN GA SC NC FL.
velutinus Fabricius, 1801: 334. "Carolina".
aurigaster Portevin, 1925: 170 (as variety of velutinus). "United States".
EcOLOGY:
-
Scott and Traniello, 1988: 693; Trumbo, 1990: 49l.
vespilloides Herbst, 1783: 32. Germany: Berlin; AK YT NTI BCI AB SK MB NDI MN MI ON
PQI NB NS NFl IL INI NY PAl ME NH MA; Holarc. See Hatch (1928: 136) for
Eurasian synonyms.
hebes Kirby, 1837: 96. NS.
TYPE DEPOSITORY:
BMNH.
pygmaeus Kirby, 1837: 98, pI. II, fig. 3. "Rocky Mountains".
BMNH.
Leech, 1937: 158.
Bartlett, 1987a: 471, 1987b: 179, and 1988: 297.
TvPE DEPosITORY:
TAXONOMY:
EcOLOGY:
Unplaced Species
pUlsator Gistel, 1848: 190 (Necrophorus) (incertae sedis). Canada.
NOMEN NUDUM
brasiliensis Dejean, 1833: 118 (Necrodes) (not available). Brasilia. nomen nudum; this
name was published without a diagnosis or indication.
10
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1847
Nomenclatoris zoologici index universalis, continens nomina systematica classium,
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1982b On the decreasing abundance of Nicrophorus american us Olivier (Coleoptera: Silphidae) in Eastern North America. The Coleopterists Bulletin, vol. 36, no. 2, pp.
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The larva of Thanatophilus trituberculatus (Kirby) (Coleoptera: Silphidae). The Coleopterists Bulletin, vol. 41, no. I, p. 34.
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Two new North American species of Necrophorus (Coleop.). Entomological News,
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A revision of the Nearctic Silphini and Nicrophorini based upon the female genitalia (Coleoptera, Silphidae). Journal of the New York Entomological Society, vol.
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-
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1884
Bemerkungen zur dritten Auflage des Catalogus Coleopterorum Europae auctoribus L. v. Heyden, E. Reitter et J. Weise. Berliner Entomologische Zeitschrift,
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Ratcliffe, B. C.
1972
The natural history of Necrodes surinarnensis (Fabr.) (Coleoptera: Silphidae).
Transactions of the American Entomological Society, vol. 98, no. 4, pp. 359-410.
20
Reitter, E,
1885
Reitter, E,
1901
Say,T,
1823
COLEOPI'ERA CATALOG
Bestimmungs-Tabellen der europaeischen Coleopteren. XII. Necrophaga. (Platypsyllidae, Leptinidae, Silphidae, Anisotomidae und Clambidae). Verhandlungen
Naturforschen Vereines, W. Burkart, Bruenn [= Br'no], vol. 23, pp. 3-122.
Uebersicht der bekannten Agyrtes-Arten. Wiener Entomologische Zeitung, vol. 20,
p.l02.
Descriptions of coleopterous insects collected in the late expedition to the Rocky
Mountains, performed by order of Mr. Calhoun, Secretary of War, under the
command of Major Long. Journal of the Academy of Natural Sciences of Philadelphia, vol. 3, pt. I, pp. 139-216.
Reprinted in LeConte, 1859d.
Say,T,
1825
Descriptions of new species of coleopterous insects inhabiting the United States.
Journal of the Academy of Natural Sciences of Philadelphia, vol. 5, pt. I, pp.
160-204.
Reprinted in LeConte, 1859d.
Schawaller, W.
1981
Taxonomie und Faunistik der Gattung Thanatophilus (Coleoptera: Silphidae).
Stuttgarter Beitrage zur Naturkunde, Serie A (Biologie), no. 351, pp. 1-2l.
Schawaller, W.
1983
Die Arten der Gattung [pelates (sensu lato) (Coleoptera, Silphidae, Agyrtinae).
Revue Suisse de Zoologie, vol. 90, fasc. 1, pp. 101-110.
Schilling, P. S.
1829
Holocnemis. Beitraege zur Entomologie, besonders in bezug auf die schlesische
Fauna, vol. I, pp. 93-94.
Schouteden, H.
1905
Les Hyponecrodes sud-americains. Annales de Societe Entomologique de Belgique,
vol. 49, pp. 195-199.
Scopoli, J. A.
1772
Annus historico naturalis Annus, Lipsiae [= Leipzig], vol. 5, 128 pp.
Scott, M. P.
1989
Male parental care and reproductive success in the burying beetle Nicrophorus orbicollis. Journal of Insect Behavior, vol. 2, no. I, pp. 133-137.
Scott, M. P. and J. F. A. Traniello
1988
Behavioural cues trigger ovarian development in the burying beetle Nicrophorus
tomentosus. Journal of Insect Physiology, vol. 33, no. 10, pp. 693-696.
Scott, M. P., J. F. A. Traniello, and I. A. Fetherston
1987
Competition for prey between ants and burying beetles (Nicrophorus spp.): differences between northern and southern temperature sites. Psyche, vol. 94, nos. 34, pp. 325-332.
Seidlitz, G.
See Seidlitz, 1891 .
1888
Seidlitz, G.
Fauna Baltica. Die Kaefer (Coleoptera) der deutschen Ostseeprovinzen Russlands.
1891
ed. 2 (1887-1891). Koenigsberg, Familien LVI, Gattungen 192 and Arten 818 pp.
Fauna Baltica was initially issued in 6 fascicles that were combined in 1891 as a complete work:
fase. 1, 1887, Familien pp. I-XL, Gattungen pp. 1-16, Arlen pp. 1-96; fase. 2, 1888 (spring), Gattungen pp. 17-48, Arlen pp. 97-224; fase. 3, 1888 (autumn), Familien pp. XU-XLVII, Gattungen pp. 4980, Arten pp. 225-336; fase. 4, 1889, Gattungen pp. 81-128, Arlen pp. 337-512; fase. 5, 1890, Gattun·
gen pp. 129-160, Arlen pp. 513.£08; fase. 6, 1891, Familien pp. XLIX-LVI, Gattungen pp. 161-192,
Arlen pp. 609-818.
-
SILPHIDAE
21
Semenov-Tian-Shanskij, A. P.
1891
Diagnoses Coleopterorum novorum ex Asia Centrali et Orientali. Horae Societatis
Entomologicae Rossicae, (1890) vol. 25, pp. 262-382.
On the table of contents page the author's name is cited as Andreas Semenow.
Semenov-Tian-Shanskij, A. P.
1933
De tribu Necrophorini (Coleoptera, Silphidae) classificanda et de ejua distributione
geographica. Travaux de l'Institut Zoologique de l' Academie des Sciences de
1'URSS (Trudy Zoologicheskogo Institua Leningrad), vol. I, pp. 149-160.
Title in French and Russian; text in Latin and Russian.
Sherborn, D. D. and F. J. Griffin
1934
On the dates of publication of the natural history portions of Alcide d'Orbigny's
"Voyage Amerique Meridionale." Annals and Magazine of Natural History, ser.
10, vol. 13, no. 73, pp. 130-134.
Shubeck, P. P.
1971
Diel periodicities of certain carrion beetles (Coleoptera: Silphidae). The Coleopterists Bulletin, vol. 25, no. 2, pp. 41-46.
Shubeck, P. P.
1976a An alternative to pitfall traps in carrion beetle studies (Coleoptera). Entomological
News, vol. 87, nos. 5 & 6, pp. 176-178.
Shubeck, P. P.
1976b Carrion beetle reponses to poikilotherm and homoiotherm carrion (Coleoptera: Silphidae). Entomological News, vol. 87, nos. 9 & 10, pp. 265-269.
-
Sturm, J.
1826
Catalog meiner Insecten-Sammlung. Nuernberg [= Nuremberg], pt. 1, Kaefer, viii
and 207 pp., illus.
Thomson, C. G.
Skandinaviens Coleoptera, synoptiskt bearbetade. Lund, vol. 4, 269 pp.
1862
Thunberg, K. P.
1789
Periculum entomologicum quo characteres generum insectorum. J. Edman, Upsaliae [= Uppsala], 16 pp.
Trumbo, S. T.
1990
Regulation of brood size in a burying beetle, Nicrophorus tomentosus (Silphidae).
Journal of Insect Behavior, vol. 3, pp. 491-500.
Trumbo, S. T.
1991
Reproductive benefits and the duration of paternal care in a biparental burying
beetle, Necrophorus [sic] oribicollis. Behaviour, vol. 117, pp. 82-105.
Van Dyke, E. C.
1921
Coleoptera from the Pribilof Islands, Alaska. Proceedings of the California Academy of Sciences, ser. 4, vol. 11, no. 4, pp. 156-166.
Van Dyke, E. C.
1928
The American species of Pteroloma (Coleoptera-Silphidae) and a new Japanese species. Bulletin of the Brooklyn Entomological Society, vol. 23, no. I, pp. 19-26,
illus.
-
Walker, F.
List of Coleoptera. In J. K. Lord, The naturalist in Vancouver Island and British
1866
Columbia. R. Bentley, London, vol. 2, pp. 309-334.
Weber, F.
1801
Observationes entomologicae, continentes novorum quae condidit generum characteres, et nuper dectectarum specierum descriptiones. Bibliopolii Academici novi,
Kiliae [= Kiel], pp. iv-xii and 4-116.
22
COLEOPTERA CATALOG
Wickham, H. F.
1902
Two new Silphidae from Colorado. The Canadian Entomologist, vol. 34, no. 7, pp.
180-181.
Wilson, D. S.
1983
The effect of population structure on the evolution of mutualism: A field test involving burying beetles and their phoretic mites. The American Naturalist, vol.
121, no. 6, pp. 851·870.
Wilson, D. S. and J. Fudge
1984
Burying beetles: Intraspecific interactions and reproductive success in the field.
Ecological Entomology, vol. 9, no. 2, pp. 195-203.
Wilson, D. S. and W. G. Knollenberg
1984
Food discrimination and ovarian development in burying beetles (Coleoptera: Silphidae: Nicrophorus). Annals of the Entomological Society of America, vol. 77,
no. 2, pp. 165-170.
Wilson, D. S., W. G. Knollenberg, and J. Fudge
1984
Species packing and temperature dependent competition among burying beetles
(Silphidae, Nicrophorus). Ecological Entomology, vol. 9, no. 2, pp. 205-216.
Zetterstedt, J. W.
1824
Nya Svenska Insect-Arter. Konglifa Vetenskaps Akademiens Handlingar, 1824, pp.
149-159.
-
INDEX
Names are indexed as follows:
CAPITALS: All names for taxa above the generic
level;
Boldface: Valid generic and subgeneric names;
Roman: Valid specific and subspecific names;
Italics: All invalid names such as synonyms, nomina
nuda, and extra-limital taxa even though valid.
The generic name following the author's name indicates the present placement of the species. Synonyms of
species-group names are listed with the original spelling.
Acanthopsilus Portevin ..................................
6
Aclypea Reitter .................................................................. . 6
Adolus Fischer von Waldheim ................. ,...................... . 3
aenescens Casey, Heterosilpha ............. ,"', .. ,................... . 6
affine Kirby, Necrophila ........................... ,........ ,... ,.... ,.... , 5
Agyrtecanus Reitter .......... ,.... ,............................. ,............ . 2
Agyrtes Froelich ... ,.......................... ,... ,........ , .................... . 2
AGYRTINl... ....................................................................... .
2
aleuticus Gistel, Nicrophorus ....................................... .. 8
alpha Pierce, Nicrophorus ............................................... . 8
americana Linnaeus, Necrophila .................................... . 5
americanus Olivier, Nicrophorus ................................... ..
7
analis Chevrolat. Oxelytrum .......................................... ..
4
Apteroioma Hatch ............................................................. . 3
arizonicum Van Dyke. Apteroloma ................................ . 3
Asbolus Bergroth ............................................................... . 3
ater Motschulsky, Necrophilus ........................................ . 2
aurigaster Portevin, Nicrophorus ................................... . 9
bicolon Newman. Nicrophorus ........................................ . 9
bituberosa LeConte, Aclypea ........................................... . 6
6
n
••••••••••••••••••
~~~h;fu~a~~~~~·i·~::::::::::::::::::::::::::::::::::::::::::::::::::::::::::
--
brasiliensis Dejean ............................................................ .
californica Mannerheim, Thanatophilus ...................... ..
Calosilpha Portevin ............................................ .
canadense Kirby. Necrophila ........................................... .
caraboides Fall, Apteroioma ............................................ .
carolinus Linnaeus. Nicrophorus ................................. .
caudata Say, Thanatophilus ............................................ .
cervaria Mannerheim, Heterosilpha .............................. .
g~~~:;~~~ ~~~:~t~::::::::::::::::::::::::::::::::::::::::::::::::::::::::
2
9
4
5
5
3
7
4
6
4
5
coloradensis Wickham, Thanatophilus ......................... ..
confossor LeConte, Nicrophorus ..................................... ..
conversator Walker, Nicrophorus ................................... .
Cyrtoscelis Hope ................................................................. .
defodiens Mannerheim, Nicrophorus ............................. .
8
7
~~~~~i;)fahp~~;~in·:::::::::::::::::::::::::::::::::::::::::::::::::::::::::
5
discicolle Brulle, Oxelytrum .......................................... ..
discretus Portevin, Oxelytrum ........................................ .
Eunecrophorus Semenov-Tian-Shanskij ......................... .
Eusilpha Semenov-Tian-Shanskij ................................... .
grandis Fabricius, Nicrophorus ....................................... .
guttula Motschuisky, Nicrophorus ................................. .
hallii Kirby, Nicrophorus ................................................ .
hebes Kirby, Nicrophorus ................................................. .
hecate Bland, Nicrophorus ............................................... .
Heterosilpha Portevin ...................................... ..
Holocnemis Schilling ....................................................... .
hybridu~ ~atch and .Angell. ~icropP.orus ...: ................ ..
hydrophtiOldes Guerm-Menevllle, Necrophllus ........... ..
Hyponecrodes Kraatz ......................................................... .
immaculosis Hatch, Nicrophorus ................................... .
inaequale Fabricius. Oiceoptoma ......... ..
infodiens Mannerheim, Nicrophorus ........................... ..
investigator Zetterstedt, Nicrophorus .................... ..
[pelates Reitter .................................................................. .
Isosilpha Portevin ............................................................ .
Katanecrodes Schouteden ................................................ .
labreae Pierce, Nicrophorus ............................................. .
lapponicus Herbst, Thanatophilus .................................. .
latera lis Portevin, Nicrophorus ..................................... ..
latifrons Pierce. Nicrophorus .................................. .
latus Mannerheim, Ipelates ................................ .
Lendomus Casey ................................ .
4
6
7
5
4
4
6
5
7
7
8
9
7
5
3
7
2
4
7
5
8
7
2
5
4
8
4
7
8
2
2
longulus LeConte. Agyrtes ...
lunatus LeConte, Nicrophorus
luniger Harold, Nicrophorus ........ .
lunulatus Gistel, Nicrophorus ..
Lyrosoma Mannerheim ....................... ..
marginalis Fabricius, Oiceoptoma .............................. ..
marginatus Fabricius. Nicrophorus ... .
maritimus Guerin-Meneville, Nicrophorus
mckittricki Pierce, Nicrophorus.
mediatus Fabricius, Nicrophorus .... .
melsheimeri Kirby, Nicrophorus ................................ ..
melsheimeri LeConte, Nicrophorus .............................. ..
mexicanus Matthews, Nicrophorus ............................ ..
montezumae Matthews, Nicrophorus ............. ..
mysticallis Angell, Nicrophorus ......................... ..
nebrioides Brown. Pteroloma .... .. ................. .
Necrobius Gistel ....
.. ............. .
Necrobora Hope .............. ,............................................... .
Necrocharis Portevin .................................. ..
Necrocleptes Semenov-Tian-Shanskij ............ ..
Necrodes Leach .............
.. .................... ..
Necrophagus. error ................ .
Necrophila Kirby and Spence .............................. .
Necrophilodes Champion .............. ..
Necrophilus Latreille.
.. ......................... .
Necrophorus Thunberg ......................... .
Necropter Semenov-Tian-Shanskij ......
............... .
Necrotropha Gistel ......................................... .
Neonicrophorus Hatch ............ .
NICROPHORINI............
.................................... .
Nicrophorus Fabricius ............. .
nigrita Mannerheim, Nicrophorus ,.... .
NOMEN NUDUM ....................................... .
noveboracense Forster, Oiceoptoma ...... .
obalskii Portevin, Thanatophilus
obscurus Kirby, Nicrophorus ............. ..
obtusiscutellum Pierce, Nicrophorus .. .
Oeceoptoma Agassiz .............................. ..
Oiceoptoma Leach ............................. ..
opaca Linnaeus. Aclypea ................................ .
opacum Mannerheim, Lyrosoma ..
orbicollis Say, Nicrophorus ......... ..
Oxelytrium, error ........ .. .............. ..
Oxelytron, error ....... ,..
Oxelytrum Gistel.......
.. ............... ..
Paranecrodes Portevin .....
.. ..... ..
particeps Fischer von Waldheim, Nicrophorus.
Pelates Horn ....... .................................... .
Pelatines Cockerell ...........
..................... .
peltata LeConte, Necrophila.
.. ............ ..
pettiti Horn, Necrophilus ........................ ..
Philas Portevin ............... ........
.......... ............. .
plagiatus Motschulsky, Nicrophorus ...... .
poWus Casey. Agyrtes .............. ..
pollinctor LeConte. Nicrophorus ........... .
pollinctor Mannerheim, Nicrophorus
Protonecrodes Portevin........ ............... ..
Pseudopelta Bergroth
Pteroloma GylienhaL ....
pulsator Gistel, Nicrophorus .....
punctostriatus Pierce, Nicrophorus ", .. .
pustulatus Herschel, Nicrophorus ....... .
pygmaeus Kirby. Nicrophorus ............... ..
quadrisignatus Laporte. Nicrophorus.
ramosa Say, Heterosilpha .................... .
requiescator Gistel, Nicrophorus ...... ..
ruficornis Motschulsky, Nicrophorus
rugulosum Portevin, Oiceoptoma ..
sagax Mannerheim, Thanatophilus ...
sayi Laporte, Nicrophorus .....
SILPHIDAE ...... .
SILPHINI .................. .
Silphosoma Portevin ....... .
similis Fall, Agyrtes ......... .
Sphaeroloma Portevin.............
.......... .
Stictonecropter Semenov-Tian-Shanskij .. .
23
2
9
9
9
3
5
8
7
8
7
7
8
8
8
7
3
2
5
6
6
3
6
5
2
2
6
6
5
7
6
6
8
9
5
4
8
8
5
5
6
3
8
4
4
4
4
8
2
2
5
2
4
7
2
7
8
3
4
3
9
7
9
9
9
6
8
8
5
4
9
1
3
4
3
2
7
24
COLEOPTERA CATALOG
surinamensis Fabricius, Necrodes ...................................
tahoecum Fall, Apteroioma ..............................................
tardus Mannerheim, Nicrophorus ...................................
tenuicornis LeConte, Apteroloma ....................................
terminatum Kirby, Necrophila.........................................
Thanatophilus !..each .........................................................
3
3
9
3
5
4
tomentosus Weber, Nicrophorus.................. ...................
trituberculatus Kirby, Thanatophilus ............................
truncatus 8(1' Thanatophilus ..........................................
tuberculata ermar, Thanatophilus................................
velutinus Fabricius, Nicrophorus.....................................
vespilloides Herbst, Nicrophorus .....................................
virginicus Froelich, Nicrophorus
9
5
5
4
9
9
7
-
NO
MT
SO
WY
-
UT
Co
NM
SOUTHWEST
AB
AK
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AR
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DE
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10
-
IL
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