Overview of Fossil Hominids Geological Time Scale
Transcription
Overview of Fossil Hominids Geological Time Scale
Geological Time Scale Overview of Fossil Hominids Period Quarternary Alan R. Rogers Tertiary March 14, 2011 Epoch Recent Pleistocene Pliocene Miocene Age (myr) 0.01–0 1.7–0.01 5–1.7 23–5 Events agriculture, complex societies genus Homo genus Australopithecus genus Sahelanthropus Hominin sites of the Pliocene and early Pleistocene Early middle Pleistocene. Archaic H. sapiens colonized Europe. Early upper Pleistocene 137–10 kya Miocene hominins Species Sahelanthropus tchadensis The Toumai skull Age 6–7 mya EA SA CA * CA: Central Af; mya: millions of years ago. Gracile hominins of the Pliocene Species Ardipithecus ramidus Australopithecus anamensis Australopithecus afarensis Australopithecus africanus Age 4.4 mya 4.2–3.9 mya 3.9–2.8 mya 3.5–2.3 mya I 6–7 mya I Central Africa I Sahelanthropus tchadensis I brain: 320–380 cc I Ancestral to humans? to chimps? I Ardipithecus ramidus I 4.4 mya I East Africa I Small brain I Intermediate between hominins and earlier apes Ardipithecus ramidus EA * * * * SA CA * EA: East Af; SA: South Af; CA: Central Af; mya: millions of years ago Ardipithecus ramidus Ardipithecus ramidus I Ardipithecus ramidus I Small brain I Small canines I Ardipithecus ramidus I Opposable big toe, flexible wrist I Walked upright; also climbed on branches The “Lucy” skeleton I 3.2 mya I East Africa I I Mrs. Ples Australopithecus afarensis most complete australopithicine skeleton Robust hominins of the Pliocene Species Australopithecus aethiopicus Australopithecus boisei Australopithecus robustus I Australopithecus africanus I 2.3–2.8 mya I Sterkfontein, S Africa E African robust australopithicine Age 2.7–2.3 mya 2.2–1.3 mya 1.8–1 mya EA * * SA CA * EA: East Af; SA: South Af; CA: Central Af; mya: millions of years ago I A. afarensis and H. sapiens I afarensis walked like we do (bipedal locomotion) I had a chimpanzee-sized brain I Australopithecus boisei I 1.5 mya I E Turkana, E Africa I massive jaw muscles I huge molars Africanus had big teeth. Boisei had even bigger teeth. I Australopithecus africanus I Australopithecus boisei I 3.0–3.3 mya I 1.5 mya I S Africa I Lake Natron, Tanzania I Thick molars I huge molars I Thick enamel I extremely thick enamel I Short canines I massive cheekbones I enormous jaw muscles I especially on robust forms On robust forms, jaw muscles reached to top of head. The genus Homo Species Homo habilis Homo erectus archaic Homo sapiens (incl Neandertal) a.m. Homo sapiens Age 1.9–1.6 mya 1.8–0.4 mya A f * * E u A s * * 400–40 kya 90–0 kya * * * * * * A m O z * * Af: Africa; Eu: Europe; As: Asia; Am: Americas; Oz: Australia; mya: millions of years ago; kya: thousands of years ago. I big molars I thick enamel I small diastema means small canine Homo habilis (sm brain & teeth) I the old Homo habilis is now 2 species I H. rudolphensis (on left): large brain; large teeth I H. habilis (on right): small brain; smaller teeth Homo rudolfensis (lg brain & teeth) Homo ergaster (↑ brain, ↓ teeth) I early African Homo erectus I restricted to Africa; 1.8–0.6 mya I also called Homo ergaster I larger brain (900 cc) I small gut I smaller cheek teeth The Nariokotome skeleton I early African H. erectus I 12 years old I 5’4” tall (like a modern human) I body proportions like ours I small teeth I powerful muscles I I Between 800 and 500 kya there appeared hominins with higher, more rounded crania and larger brains I archaic Homo sapiens I includes Neanderthals I archaic H. sapiens or early modern I 90–190 kya Asian Homo erectus Shanidar, Iraq Jebel Irhoud I Neandertal Tiny hominins of Flores Island, Indonesia early modern Homo sapiens I Homo floresiensis I Stood about 1m high. I Brain 380 cc. I 18 kya I Genetic deformity? I Island dwarfism? Summary I Hominins are the bipedal apes in that portion of the evolutionary tree that includes humans but not chimpanzees. I Sahelanthropus was a Miocene ape that may have been ancestral to humans, to chimps, or to both. Not clear whether it walked upright. I In the Pliocene, there was a variety hominin species. They walked upright but had chimp-sized brains. I Larger brains began to appear at the end of the Pliocene, with the advent of Home habilis and Homo erectus. I In the early Pleistocene, Homo erectus expands out of Africa and throughout Europe and Asia. Summary (cont) I The species called archaic Homo sapiens appears about 400 kya. It retains the strong brow ridges and receding chin of H. erectus, but has a larger brain. I Anatomically modern Homo sapiens appears about 100 kya and spreads throughout the world about 50 kya. I erectus/ergaster’s ribcage less conical I implies a smaller gut I implies a high-quality diet