(Anisoptera: Gomphidae) engaged in the study

Odonaiologica 17(4):
December
429-434
I,
1988
Descriptions of three new taxa in the genus Davidius
from Shaanxi, China
Selys
Zhu¹, Z.-d.
H.-q.
of
'Department
(Anisoptera:
Yan
²
Biology, University
People’s Republic
department
of
Biology, Hanzhong
D.
miaotaiziensis
allotype $:
same
Yuanba.
Shaanxi:
Miaotaizi,
and
the
figured.
The
—
1987
(holotype
Vl-1983),
D.
and
1X-I984;
and
China,
13-VI-I987),
D.
College, Hanzhong, Shaanxi,
of China
Revised
/
3 distal
types
are
abdominal
deposited
Shaanxi:
yuanbaensis
same
segments and
in
Accepted April
davidi shaanxiensis
allotype 2:
the
Li
²
of China
locality,
The latter is similar to D. d. assamensis
unmarked
7-9.
n.
sp.
locality,
Shaanxi:
16,
S.-s.
Shanxi, Taiyuan, Shanxi,
Teachers
People’s Republic
Received December
and
of
Gomphidae)
sp.
ssp.
Miaotaizi.
I2-VI-I986;
n.
Department
of
China,
(holotype $:
are
described
from which it differs
the sternal processes
by
China,
(holotype
n.
I5-VI-I987)
Laidlaw,
1988
6,
on
Biology, Hanzhong
by
segments
Teachers
College.
INTRODUCTION
Since
1981, the second and the third authors
are
in the
engaged
study
odonate fauna of the southern Shaanxi. The present paper is based
collections
of the
the rich
gathered.
The type
Teachers
on
specimens
College,
are
deposited
Hanzhong,
in the
of
Department
Biology,
Hanzhong
Shaanxi, China.
DAVIDIUS MIAOTAIZIENSIS SPEC. NOV.
Figures
Material
same
25-V1-I985,
male.
mm.
examined.
locality,
one
—
—
I3-V1-I987.
Holotype $: China, Shaanxi, Miaotaizi,
—
of which has
Total
1-7
length
Paratype $\
no
45
cross
mm,
vein
same
in
locality,
discoidal
5-VI-I987.
cell of hind
abdominal length 32
I2-V1-1986.
—
2
$\
—
Allotype 2:
Shaanxi,
Yuanba,
wing.
mm,
hind
wing length
27.5
430
Figs
H.-q. Zhu,
1-7.
hamule,
left
dorsal view;
view;
Colors.
Head.
black
at
lapping
miaotaiziensis
Davidius
appendages,
—
—
—
ventral
of
—
—
a
9); (0
7:
Anterior
Vulvar scale,
Pterothorax;
hamule left
view;
—
the front
—
A
stripe;
black
a
stripe
on
stripe
and
antefrons
are
yellow.
bordered with
yellow
and
over-
yellow.
mesothorax collar stripe confluent with
the antehumeral
present
Anal
occiput yellow.
of hind lobe
complete
(2)
Posterior
ventral view.
transverse
dirty yellow;
—
(5)
and
the
stripe slightly converging above;
laterally
broad
two
postero-lateral
yellow stripes
suture.
Dorsum and sides of segment
1, auricula and baso-lateral spot
small
spot
a narrow
margin
(7)
Fore-area and sides
(Fig. 1).
narrow
segment 2,
segment 3,
—
Fig.
(4)
and hind part of
upper humeral spots
Abdomen.
distal
on
epifrons,
shorter median dorsal
a
view;
margin anteclypeus
the front of
separated by
1-6:
(Figs
Li
Black, the pale parts of head, thorax and abdomen
Pterothorax
isolated
n.
Penis, left view;
Labrum with
Prothorax.
a
(6)
sp.
Same, left
(3)
Z.-d. Yan & S.-s.
of 7
triangular
basal
and
baso-dorsal
ring
of 4
to
8, yellow;
and
baso-lateral spot
of
6, three basal spots of segment 7, and each
remaining segments
and
anal
appendages
unmarked.
Anal
appendages (Figs 2, 3).
slightly diverging backwards,
outer
a
tooth
basally;
each
—
The
ending
superiors
in
a
the inferior shorter, round
small dorsal process
on
each side.
about
curved
as
long
upward tip,
as
segment 10,
and
bearing
apices broadly separated
an
and with
New taxa
Genitalia of second segment
fore branch
FEMALE.
29
mm;
3
—
to
as
Total
42
length
6
pale
mm,
area on
broader and
the sternite and
examined.
locality
MALE.
—
and
date,
Total
—
slightly
with
bifid
a
at
has
length
no
China:
45
cross
2
on
apex
wing length
more
baso-lateral spots
lateral
spot;
on
7
only
8; the vulvar scale cordiform,
as
(Fig. 7).
SPEC. NOV.
8-13
Shaanxi.
vein in discoidal
Yuanba,
cell
abdominal length
mm,
hind
of abdomen
confluent;
longitudinal
YUANBAENSIS
Holotype $:
one
its
hamule
small, apex of its
mm,
yellow marking
segment
and it is very small
Figures
Material
penis
divaricated,
posterior
one;
abdominallength 30.6
sides of
marked
spot present,
Anterior hamule
than the rounded scale lobe.
DAVIDIUS
same
—
curved tooth;
backwardly
a
longer
follows;
as
baso-lateral
long
in
(Figs 4-6).
coloration similar to the male, but the
extensive
on
ending
blunt and
post-lobe
431
Davidius from China
and recurved toward the hind
slightly longer
robuster and
in
Vl-1983;
of hind
33
—
Paratypes (3 $):
wing.
mm,
hind
wing length
27
mm.
Figs
—
8-13.
(10)
(13)
Davidius
Same, left
Penis,
yuanbaensis
view;
left view.
—
(II)
sp.
n.
(<3): (8)
Anterior
Pterothorax;
hamule,
left
view;
—
—
(9)
(12)
Anal
appendages,
Posterior
dorsal
hamule. left
view;
view; —
432
Zhu,
H.-q.
Colors. —
Head.
the
Black,
pale parts
Labrum has
—
ventral
margin; anteclypeus dirty;
ventral
margin;
antefrons and
Prothorax.
with
two
laterally
postero-lateral
Abdomen.
—
yellow;
A
a
stripe;
pair
of twin
Anal
spots
the antehumeral
Dorsum and sides of
to
on
median
yellow.
area,
stripe
confluent
stripe slightly converging
a
broader black
stripe
2,
segment I,
dorsal
a
on
(Figs 9,
on
—
10).
5
7 which
to
The
and the inferiors each
Genitalia of second segment
to 9
are
becoming
to
4,
progres-
yellow; remaining segments
(Figs
simple
ending
11-13).
in
—
superiors
upturned
a
slightly
recurved
diver-
apex.
Similar to those of D. miaotai-
but fore-branch of anterior hamule broader and
into
auricula and
stripe,
and baso-lateral spot and lateral spot of 3
7; each distal margin of 6
appendages
penis tapering
at
unmarked.
appendages
ging backwards,
at
bordered with black
mesothoracic collar
stripes separated by
smaller baso-lateral spots present
and anal
yellow.
are
unmarked.
complete
very broad
abdomen
front, bordered with black
on
suture.
smaller from 5
ziensis,
stripe
yellow postclypeus
occiput
—
baso-lateral spot of segment
sively
the
and
area
(Fig. 8).
shorter median dorsal
a
above and
the
Fore
—
Pterothorax
Li
of head, thorax and
transverse
a
Yan & S.-s.
Z.-d.
the
posterior
lobe of
prominent tip.
a
DAVIDIUS DAVIDI SHAANXIENSIS SSPEC. NOV.
Figures
Material
same
MALE.
—
examined.
locality,
—
Holotype $:
14-22
China:
Shaanxi.
1X-I984;
Miaotaizi.
Allotype
$:
wing length
29
I5-VI-I987.
Total
length
44
mm,
abdominal
length
34
mm,
hind
mm.
Colors.
Head.
the
Black,
—
—
Labrum
bordered with black
at
pale
parts of head, thorax and abdomen
unmarked;
ventral
anteclypeus
dirty;
antefrons
are
yellow.
and
yellow
margin; occiput yellow spot extending
to
its hind
part.
Prothorax.
Front and
—
hind
lobe
and
yellow
all
bordered
with black
margin.
Pterothorax
with
side
a
broadly yellow
Abdomen.
—
and baso-lateral
spot
margin
of 6
backwardly
—
A
complete
separated by
of
mesothoracic collar
the antehumeral
Dorsal and
baso-lateral
(Figs 15, 16),
stripe:
and
yellow;
to
14).
(Fig.
median dorsal
a
black
stripes
stripe
7
a
on
stripe
converging
the
lateral spots of segment
confluent
above: lateral
postero-lateral
suture.
I, dorsal stripe, auricula
baso-lateral and lateral
of
segment 2,
spots
segment 3,
spots
progressively
smaller
9, yellow; ventrally segments 7
on
are
tubercle present
directed blunt
spines,
on
from
to
9
are
4
to
7, and each distal
provided
with
a process
the sternite which is covered with 7
segment 8 with
a
to
process about half way
10
(cf.
New
Figs
14-22.
Davidius
(15)
Abdominal
(17)
Anal
(20)
Fig. 15),
a
in
davidi shaanxiensis
segments
appendages,
Posterior
taxa
hamule,
7
dorsal
left
and
8,
view;
view;
—
and segment 9 with
ssp.
right
—
n.
view;
(18)
(Figs
—
left
14-21:
(16)
Same, left
(21) Penis,
a
433
Davidius from China
Fig.
Abdominal
view;
view; —
—
(19)
(22)
29:
2): (14) Pterothorax; —
segment
Anterior
Vulvar
9,
ventral
hamule,
scale,
ventral
margined irregular yellow swelling
and
view;
left view;
—
—
view.
ending
in
little tooth.
Anal
forked,
appendages
the
outer
(Figs
branches
17,
18).
strongly
—
Shorter than segment
divergent
and
10;
sharply pointed,
branches smaller and inclined downward and recurved
superiors
the inner
slightly; inferior shorter,
434
H.-q. Zhu,
rounded with
nipple-like
a
Z.-d.
robust and
are
in
equal
the
tapering;
19-21).
—
Anterior hamule
but the fore
length
Li
above.
process
Genitalia of second segment (Figs
both branches
Yan & S.-s.
penis small,
scale lobe
robuster;
one
deeply forked,
posterior
than
slightly longer
hamule
post-lobe and
both recurved backward.
female.
mm;
Total
—
coloration
extensive
length
similar
follows:
as
male, but
area
on
extending
smaller from 3
the vulvar scale
abdominal
mm,
the
yellow
-lateral linear spot which is
progressively
42
to
to
32
length
yellow
segment 2 confluent;
to
the
jugal
hind
mm,
suture
on
wing length
of abdomen
markings
longitudinal
a
3
to
29
more
baso-
7, the lateral spot
7; the baso-lateral spot of segment 8 very obscure;
(Fig. 22) strongly
notched and almost
as
long
as
the sternite of
segment 9.
—
Remark.
The
three unmarked
from each
new
distal
subspecies
is
easily distinguished from
abdominal segments
and
by
the sternal
D. davidi assamensis Laidlaw
processes
7 to 9. which
on
are
by
the
different
other.
REFERENCES
CHAO,
H.-f.,
Ada
1954.
enl.
sin.
FRASER. F.C..
Classification of Chinese
1934.
The
Taylor & Francis,
HAMADA,
K. &
dragonflies
of the
family Gomphidae (Odonata).
III.
4(3): 213-275.
K.
fauna of
British India,
including Ceylon
and
Burma, Odonata.
Vol. 2.
London.
INOUE,
1985.
The
dragonflies of Japan
in colour.
Vols
I
& 2.
Kodansha,
Tokyo.
NEEDHAM.
20
pis
J.G..
excl.
1930.
A
manual of the
dragonflies
of China.
Zool.
sin.
(A)
11(1).
344+11
pp„