Eastern - Natuurtijdschriften
Transcription
Eastern - Natuurtijdschriften
CRANIUM 21,1-2 Mammal Collections Quaternary (2004) in the Museums of Yakutsk (Eastern Siberia, Yakutia, Russia) Gennady+G. Boeskorov and Dick Mol Summary The presence of permafrost of the Sakha territory collections of fossil Nature, the Russian Academy that belonging mammals parts of the to fossils are mammoths existed already dating kept housed in the in the the collection of the Mammoth Museum in more than 48,000 BP on the Regional Metals, right a and parts of spectacular of Yakutsk and museums: describe we but also of interesting most museums following paper parts to the Quaternary has been found in the Diamonds and Precious of Sciences and the Mammoth Museum. In this in of skeletal only of Eastern Siberia. The Geology of Museum of the Institute of Geological most Quaternary are not preservation permafrost occupies these collections mammoth, Mammuthus primigenius, woolly of the significant part (Yakutia) Republic reviewed here. The main conditions for unique creates of extinct animals. A carcasses are Siberian femur of Division, small a briefly Museum of Yakutsk, showing woolly that small the continentof Eurasia. Samenvatting Collecties De van Kwartaire Zoogdieren of bevroren bodem permafrost eeuwig de zachtere onderdelen van Vondsten uit de geslachtsdelen. hebben veel zoals de bijvoorbeeld Pleistoceen. in de inhoud Akademie hoofdstad van objecten van bijgedragen de en van vaste land van een dat het over de oosten o.a. van het uiterlijk wolharige maag van tot een de van jongste zijn over bijdrage wolharige Pleistocene Vroeg en de vinden in drie en wij Holocene meer het Zoölogisch het een gebruik zoogdieren tijdens on de het kort overzicht van van heeft de unieke de maar van op ook jaar geleden belangrijke de wolharige tentoongestelde mogelijkheid te ook om een zeer beschrijven. voorkwamen territory of Eastern Siberia is located which is by than 0° Celcius. and months) layers of relict of the Pleistocene a average annual air an Long short and the surface of temperature less winters severe keep summers the ground depth of hundreds of meters and in more than summer upper 2 one layers meters. of the In the melting during permafrost preserving in the kilometer. in Central ground north summer serves as a is not of is the melting a places in the of the than 1,5 Siberia - the shallower. So, the natural refrigerator remains of animals and ground. frozen to some deeper Eastern even to 8 upper example, For Yakutia (up the plants The wide distribution of for Fig 1 Territory of Sakha (Yakutia) Republic frozen Grondgebied perma- 19 van de het Laat Museum van de Eurazië. permafrost, caused zijn van in 1999) Introduction Most of the en Regionaal Natuurmusem, gespecialiseerd dan 48.000 van Jakutië, van gebitsorganen wolharige mammoet, Mammuthus primigenius, mammoeten ook Voedselresten die deze dieren Tikhonov, Een aantal we tevens van o.a. & musea: geven maar grondgebied steppenwisent. in Jakutsk dat zich In dit artikel maken skeletdelen, het leefomgeving te van zelfs organen als maag, darmen met name van (Vereshchagin verdeeld museum de en mammoet en holtes in wolharige Petersburg zijn van mammoet beter beeld in Sint volwassen kleine conservering Siberië en zijn eigen tentoonstellingsruimte. kleine, zeer unieke voor een deze Siberische vondsten Sinds 2002 heeft het toont aan van van de objecten worden in deze bijdrage afgebeeld. skeletelement zorg Jakutsk (Oost Siberië, Jakutië, Rusland). het Mammoet Museum. In deze en Mammoet Fauna, kleine rechter femur van huid, haar, kraakbeen, spieren De collecties daar Jakutië, Jakutsk. musea. als Wetenschappen van Museum in die mammoet organen Belangrijke collecties Russische Geologisch permafrost wolharige neushoorn, neushoorn hebben wolharige draagt dier zoals de informatie verschaft belangrijke aangetroffen een in de Musea Sakha (Jakutië) Republiek Dit op het Mammal Collections in the Museums of Yakutsk Quaternary frost in Siberia created for the preservation complete conditions really unique of bones of carcasses (Mammuthus primigenius) and parts The Museum. of bited and other mammals. the major part of such mammal Pleistocene Republic (Fig. 1). Investigations fauna and for expeditions the Pliocene extinct -10.000yBP) of the Mammoth is a nice been conducted for than 200 more world famous mammoth remains country and of Saint are kept in the Petersburg, e.g. skeleton with of parts Most years. Zoological the skin of Mammoth famous Berezovka from this (Equus Museum discovered of legs in the 1799), Mammoth found in rhinoceros wolf 1900; a from lenensis), of parts bison Steppe musk-ox (Alces spec.), ( Canis lupus) deer cave lion etc. the river so-called the basins girka and including (Bison In Museum of Nature 1891 the Museum Regional of Nature established in Yakutsk. From that time finds of Pleistocene placed were worth here mentioning reindeer (Cervus elaphus), Another collection of - one Early Olyrian originating from the Pleistocene Fauna belonging inhabited (which of the northern rivers Kolyma, (Coelodonta nasal bison, the following display: on horns, Yana) (Sher, 1971; Vangengeim, 1977) the broad-fronted moose (Cervalces was and the Vera and others. many It is mamma- woolly on River rhino- with both frontal antiquitatis) several (Equus vera) lati- beringi- mammoth a Tirekhtyakh (Indigirka (Bison priscus) horse Indi- mammals museum. in 1971, the cranium of the basin) ceros this horse (Panthera spelaea), ensis), steppe goat (Soergelia spec.) (Mammuthus primigenius), recovered the bank of the river and are Holocene in display on lian remains which skeleton and on woolly priscus), frons), Praeovibos Beringian musk-ox ( Regional woolly Lena ( Ovibos pallantis), red etc. end of the Pliocene rhinoceros woolly and there discoveries. There skeletal great importance is the Verkhoyansk (1877). The be a to the and the head of Vilyui (1771); of woolly a spectacular (Rangifer tarandus), so-called (the museum ( Coelodonta antiquitatis), Pleistocene part of the body including the head and parts of two can collections of ( Mammuthus primigenius), the moose Adam's stored in this of mammoth complete a Extensive in rhinoceros come are than 5000 more mammoth Yakutia Pleistocene entire museum. display the remains collecting have animals and the representatives of the Mammoth Fauna (120.000 (Yakutia) are of fossil remains from the in Sakha recovered were of territory of Yakutia during the end of found in this The collection largest mammals of the northeast of Russia which inha- mammoth woolly (Eastern Siberia, Yakutia, Russia) skulls of the and of the Lena horse steppe (Equus lenensis). The sub-fossil skeleton of the Greenland Balaena Whale, 1500 yBP, shore tochiya The the near radiocarbon mysticetus, found in 1973 on mouth of dated the East Siberian Sea the Bolshaya Kuropa- river is also of interest. Geological tute of Geology Precious Museum of the Instiof Diamonds and Metals, Siberian Division, Russian Academy of Sciences In the second half of the 20 th century the Institute of the Yakutian Branch of the Siberian Geology, Division started of to mammals, geological USSR up a especially originating institute the built from the and were Academy collection of the of of Sciences, Pleistocene Mammoth 2 Complete hind leg of the Berelekh mammoth permafrost. Step by step, paleontological Fig Fauna, the collections in that brought together in the Een Geological 20 het woolly mammoth from ’cemetery’ complete achterpoot moet van a van een wolharige zg. mammoet-’kerkhof’ van mam- Berelekh CRANIUM 21,1-2 (2004) Fig Fig 3 Part of the Abyi mammoth 5 Akana mammoth skeleton baby Mammoetskelet uit Akana Gedeelte van het Abyi mammoetkalf There are complete mammals in the including exhibits, unique some skeletons and of fossil Museum. These carcasses Geological include: o Complete hind Mammuthus cm. of leg Discovered in 1970 moth "cemetery" relekh, (Indigirka on the Berelekh on 1995). between 14.000 The frozen leg Basin). More than 8000 and cm length and o Fig nails Gedeelte van de mammoethuid gevonden Part of the so-called are Abyi first was some on a it x 25 (Ve- 1999). mammoth woolly was District, Russia and the third Yamal Peninsula, at baby (Fig. mammoth Effie, north of Fairbanks, Alaska, USA, the second 21 of which preserved This is the fourth find of baby (the in Kular separately & Tikhonov, 1990 and 4 Part of the Kular mammothskin 3). time- of 105 cm. The foot size is 24 four reshchagin a yBP (Mol, is covered with long leg red-brownish coloured hairs a in dating 12.000 found was the site. The skin of this reach 175 mam- the banks of the river Be- River bones have been found here span mammoth, woolly a primigenius (Fig. 2) measuring Russia). Dima, was The Magadan Masha Abyi on the mammoth Quaternary Mammal Fig Collections in the Museums of Yakutsk 6 Mammoth skeleton from the River Khroma baby discovered and excavated in 1990 was the bank of the stream the The Indigirka village Belaya following parts with milk dentition, trunk, 12 cm. long milk tusks, the left front ears, the o with leg was approximately According geological age of the sediments which included are mummy between 42.000 was - 27.000 yBP). in such sediments found. Absolute age a The in the In Large part of ments 170 moth x 130 (Fig. 4). operations skin cm.) It (maximum of the Kular the third woolly that 1977). was found in 1980 at the Kular stream the Omoloi find was preserved, mam- during mining are gold-mine (up of from Akana ted from River). Geological age: Late adult male (Fig. 5). the Chukochiya of a river River woolly The skeleton cm. bank near the of mammoth was the locality skeleton complete both as 2001). - the to an It is well as of the woolly rhi- (Boeskorov, In this wonderful frontal and horns, the nails Also the hind on yellow and the on fragments leg. nasal, are the hind and length The colour of the length of the of the hair is 15 gastro-intestinal tents have been preserved an Coelodonta belongs restricted to Eurasia, preserved wool is The Skeleton carcass front legs. The fragments of the skin with wool Pleistocene. o Late Pleistocene. rhinoceros in the world 2001a, Mol & de Vos, measure- woolly The bison of +/-1000 yBP (Flerov, the in near village Churapcha (Central Yakutia). specimen o Khroma adult female woolly rhinoceros and was found noceros is 29.500 River male a Found woolly rhinoceros, Churapcha antiquitatis (Fig. 7). dated in the Kar- ginian Interglacial of the Late Pleistocene (ap- mummy of (Fig. 6) (not mounted). the stream up skeleton complete mammoth Khromskaya Quba (Gulf). (1994) the to Lazarev the remains of this animal locality An almost 1988 o 1971 in that embe- was two months old at the time of death. proximately age the sediments in which the skeleton woolly limbs, ribs and of the unfused vertebrae. The mammoth parts of (Nizhne-Kolymsky District). Geological the head preserved: including uit de rivier Khroma ded: Late Pleistocene. (Abyi District). Gora are on River, 50 km down- skin and wool, parts of the other baby / Mammoetskelet (Eastern Siberia, Yakutia, Russia) (Lazarev et con- al., 1998). of the mounted skeleton, measured from the anterior end of the cranium to the first caudal excava- vertebra, Bolshaya withers is "Akana" Moscow on 22 160 this is cm. 260 cm., the Radiocarbon specimen height at dates in showed the animal CRANIUM 21,1-2 (2004) died 19.500 rhinoceros tion of a +/- 120 yBP (GIN-0594; 2001 Boeskorov, was "Ecomare" on Netherlands o Holotype The skull Delta Hill. for (Mol Equus (a was 2001). 33) (Fig. 8). found in the Lena River American-Khaya near withers at museum lenensis Russa- cranium N region height the & De Vos, Comparatively length crea- the island of Texel, the of the 1968 nov, Churapcha "life-size" model of the wool- rhinoceros ly The a). the base for the small horse with 125-140 cm. Basal of this cranium is 490 mm. Late Pleistocene. Fig 7 The Churapcha woolly o rhinoceros skeleton Equus Het skelet van een wolharige neushoorn gevonden in Churapcha Kular loi Fig 8 Skull (holotype) of Equus lenensis vonden in de delta van de Lena Russanov / De schedel vlakbij American-Khaya (holotype) heuvel 23 of Fragment van a lenensis foal of the Lena horse, (Fig. 9). gold-mine (up River). Equus Floof, lenensis Found at the stream the Omo- muscles and Russanov, 1968, skin ge- Quaternary Mammal Collections in the Museums of Yakutsk o Two well-preserved Panthera lions, the (Eastern Siberia, Yakutia, Russia) skulls of adult male spelaea (Fig. 11). and described 1987, in (Boyeskorov & Lazarev, at stream of the Ko- Duvanny outcrop (Lower lyma River) cave Discovered recently Baryshnikov 1997; & Boeskorov, 2000). The Mammoth Museum of the Institute of of the Applied Ecology North, Academy of Sciences of the Fig 9 Fragment of a Lena horse foal Sakha leg (Yakutia) Republic, (“The Mammoth Deel van een been van een Museum”). veulen uit de Lena In 1991 the Mammoth Museum specialises on mammoths Fauna. The aim of with hair are preserved. Geological age: this was and foundedand the study the Mammoth Fauna and its environment during the Pleistocene. Mammoth Late Pleistocene. The collection Museum consists of o Fragments vicola skulls found uisk of the Siberian (Fig. the on Vilyui City, right River lities near are 10). one sheep snow The left River one near Ovis niN on 1430 standing tion of These two locaarea during of than of Siberian snow was many not example, sheep / Schedels van study large to improve under- mammals at the end beginning Siberische 24 as a big as right female many woolly woolly people femur of a very woolly the main- mammoth assume. As small but mammoth, sneeuwschapen (Ovis nivicola) of the of the Holocene. parts) from land which indicates that the in the grown 10 Skulls than 1000 remains of of the Late Pleistocene and the extinc- mammoth (isolated skeletal present time. Fig the The collection contains much material of This circumstance the Late Pleistocene more of mammals of the Mammoth Fauna. This Pleistocene and the demonstrates the wider distribution of Ovis nivicola larger collection needs further the Lena situated far from the modern distribution of this species. the Verkhnevily- is N 4837 found Kachikatsy village. is Mammoth is to museum an full Mammuthus CRANIUM 21,1-2 (2004) Measurements Maximum 0 82 length femoris Caput width Maximum cm 11,9 cm shaft anteri, „ 6,8 cm or-posterior or-posterior Maximum width shaft ventral- n , Cm 93 cm 9,3 lateral width distal Maximum sis sis epiphyr v 1 .. Maximum width distal epiphyepiphy- ' mammoth woolly from three to Compared femora which Island Wrangel Mammoth Museum is cave lion N 6397 discovered at the 83,6 Duvanny outcrop Schedel de van werd in de (for cm a grottenleeuw Mammuthus Islands for length 84,2 are exilis off the these femur of Museum The most 12 Mammoth skin from the Bolshoy Lyakhovsky organs Bolshoy Lyakhovsky the studied by The results us. the are Mammuthus primigenius primigenius (BLUMENBACH, 1799), Mammoth Museum, a a carcass of Eurasia. exhibits in the woolly mammoth, steppe bison. Of great a partial mammoth, woolly than more part the small such carcasses parts of internal woolly rhinoceros, of an of the Lena horse and Early Holocene moose. a part of a fur mammoth, the There is also (and underfur) of woolly rhinoceros and bison. steppe Catalog of the museum are: o Number 7176 Locality the Kolyma Gender Female Individual ars) 50 50 AEY age River, Yakutia on caput femoris caput age, Groningen 14 C University, the fusion the Islands), 1994 cm., width 92 cm. part and the shaft an the the mammoth found Island (New Sibe- completely diaphysis fufu- indicaindica- old old individual 25 to an part of the skin is from the preserved. the shoulder ged femoris femoris is Caput ting including yBP (GrA 20382) with this 12). Max. length 219 (Fig. This of the shoulder, in underfur is Netherlands) sed woolly Bolshoy Lyakhovsky rian on 48.000 the the head, of the (AMS, the in Lower Stream of Ye(=African Elephant Ye- based exhibits interesting Part of the skin of the on Duvanny Yar, most Yakutsk. FeFe- dexter Remarks are woolly Some Table 1 Geological in shows in Table 1. presented mur cm). The right existed the skeletons of The Mammuthus exilis Yakutsk extensive collection of an was Channel USA. eiland carcass primigenius, the between 59 and 180 - and skin, horns of the part of Mammoethuid van for the remains of of legs as Island are already rhinoceros and woolly from significant osteological museum are Fig in The cm. total of 14 femora of BP on the continental years interest cm. Mammuthus primigenius (#7176) mammoths woolly 48.000 with unfused The shoulder- California, 14 femora estimated between 150 Mammoth a of (Shoulderheights cm a measu- follows: as be less than 180 collected coast are specimen might Arctic 7176 in the # smaller. The and 101 of # 7176 fits better in length ontsluiting bij Duvanny cm for # 7176 height reported Siberian (2003), even subadult epiphyses), 98,5 N 6397 die ontdekt Holocene Tikhonov et al. given by rements the (in the been of have Tikhonov et al. Ocean) by 11 Skull of the , Cm 15,3 cm sis ventrolateral Fig „ 18,3 cm ant.-post. eye-opening, some spots ear and a the fur and The thickness of the skin part is 22 mm. The skin belon- adult individual of medium size. This piece of (1996) after its skin was figured by Engesser discovery on et al. Bolshoy Lyakhovs- Mammal Collections in the Museums of Yakutsk Quaternary Fig (Eastern Siberia, Yakutia, Russia) 14 Part of the front leg of the Maksunuokha mammoth Gedeelte de van voorpoot van de Maksunuokha mammoet Fig 13 Fragment of of the left hind the mammoth skin with remains leg wer-arm Deel van de mammoethuid met resten de linker van is 53 achterpoot loured hair According longed ky Island. The 14C results age of 26.860 the skin on +/_ 290 yBP (Lazarev et of Fragment the skin of a woolly with the remains of the left hind recovered was from Bolshoy Island 1995. The size of this piece 140 cm. It is from the posterior Another mammoth o cm. cm.) The an long yellow has fallen off the (2001) this individual which died at 18-20 leg an co- leg. be- age of years. 1971 the mammoth Mammoth Lyakhovsky of skin is 200 Section of mummified gastro-intestinal organs of the famous Shandrin Mammoth It leg (Fig. 13). was carcass found near of (Fig. 16). the In Shandrin the Shandrin River, x part of the body. part of this find is represented by the tibia and fibula with muscles and skin et to (up to 54 al., 2001). o o The sole of the front foot is to Lazarev et al. approximately an gave cm. cracked, its width is 28 (Lazarev al, 2001). Part of the front leg (Fig. 14) of a woolly mam- moth, found in 1995 and excavated in 1997 on the bank of the Maksunuokha River, Ust-Yans- ky District. piece The front of the shoulder served. There leg, it is are leg with the carpus and region very well preon the front dark red hair and the thickness of the skin is 29 mm. Short hair was cranial and caudal side of o was remains of wool preserved the age: Late Pleistocene (Lazarev Part of a a front 15) found 1994. The ted by the on leg of woolly carpus the leg. Geological et al., 2001). mammoth Bolshoy Lyakhovsky preserved part of on Fig (Fig. Island the leg is represen- length hoy Lyakhovsky leg found on the Bols- Island in with skin and the lower-arm without muscles and skin. The 15 Part of the mammoth front Deel van een voorpoot Bolshoy Lyakhovsky of the lo- 26 van de eiland mammoet gevonden op CRANIUM 21,1-2 a of the tributary parts of the were cm. gastro-intestinal were carcass of the Indigirka well preserved. cut into several branches, ved in organs inside the The whole weight slices, sizes of 70 x 35 x 15 vegetation (grasses, bushes, mosses and seeds) were preser- good condition in the stomach and intes- tines. This exhibit is of the internal nists The frozen about 300 kg. The frozen organs is organs The remains of tree River. (2004) to one organs examen of six unique remains which enables the woolly paleobota- mammoth diet 1993). Radiocarbon dates sho- (Ukraintseva, wed the Shandrin Mammoth died 41.000 Fig yBP. 16 Section of mummified gastro-intestinal organs of the Shandrin mammoth The skeleton of the Shandrin Mammoth is display more the in details on museum in Novosibirsk. the Shandrin Mammoth on For Gedeelte we revan fer to o Vereshchagin & Tikhonov, (Fig. 17) was near the o skeleton contains two-third of the bones of the The part of the skin of 18) approximately same A recon- was outcrop in 1999. It animal. The at the shoulder of this specimen is 285 an cm. The woolly ween Fig Churapcha Mammoth was a male up interglacial years 17 Skeleton of the old (Lazarev Churapcha et al. van 27 de Churapcha measures to 19 1998). mammoth / Skelet woolly rhinoceros cm. 68 x mammoet (Fig. Duvanny Yar 60 cm It dates to the of the Late Pleistocene. mammoth which died somewhere bet- 30-40 a found at the famous site thickness is height gemummificeerd maagdarmkanaal mammoet Churapcha found in 1990 village Diring (Churapcha District). structed het (1999). The skeleton of the LatePleistocene Mammoth van de Shandrin and the Kargini- Mammal Collections in the Museums of Yakutsk Quaternary leton (Eastern Siberia, Yakutia, Russia) belongs to one individual young and the (cranium same relatively with both horns, part of the vertebral column, ribs and limb bo- nes). This partial skeleton was found Aldan River, Mamontova Gora and the o age built skeleton of the steppe bison, Bison The (Fig. 20). up outcrop, compilation expeditions lyma 18 Part of Gedeelte van woolly van een wolharige neushoorn o 38.500 Part of the Pleistocene rica diffe- in the lower stream of the Ko- yBP (GIN-11021). rhinoceros skin de huid steppe during River. The absolute radiocarbon date for the cranium is Fig skeleton is from several individuals of the bison which have been collected rent the 1976, is Late Pleistocene. Compilation priscus near (= borealis) skull (Fig. 21). the Lena River bank tata- saiga antelope Saiga near a It was found on mouth of Batamai River in the Late Pleistocene sediments. o Compilation Coelodonta skeleton of antoquitatis a woolly rhinoceros, 19). (Fig. third of the bones used for the Fig 19 Compilation Samengesteld skeleton of skelet van een de rivier Aldan in een a About compilation woolly ske- An extremely large woolly mammoth and (Fig. heavy 22) tusk of collected a male in the rhinoceros wolharige neushoorn; ontsluiting o one- een derde deel behoort tot een enkel individu, gevonden bij in Mamontova Gora in 1976. De ouderdom is Laat Pleistoceen 28 CRANIUM 21,1-2 (2004) 29.500 500 +/- yBP (MAG-943) (Boeskorov, 2001b). o of Part an Peninsula hind near leg, 120 Fig 20 Compilation skeleton of een ma. van een aantal individuen uit de De schedel steppewisent, benedenloop is (GIN-11021) with hair for this age gedateerd bestaande uit van de and pie- preser- specimen is 8080 +/- mammoth remains of on the New Siberian Islands KolyAs op 38.500 jaar obvious from these lists, is the major part is found in the Far North, above the Polar Circle, B.P where the best conditions for the soft remains Anabar District. outer curvature of this well o leg well are yBP (GIN-11727). Some notes Samengesteld skelet Bykov bison steppe a the part of the vertebrae, bones from the front ved. Absolute (Alces spec.) on the delta of the Lena River. The of skin covered ces moose found in 2001 of the cranium, fragment one Holocene Early (Fig. 24) carcass Part of a The maximum measures preserved mummified Lena horse, Equus 340 tusk is 80 of carcass lenensis length cm. on head lips) the shore of (including the an right (Fig. 23), of the torso soft and up to 8 on was found Indigirka Ri- The eye-opening and are pre- the neck and the anterior are cm. weight Dyokarskoye. ear, the adult female and the anterior part of the torso served. The hair part Lake on preservation of be found. One of the most promi- kg. in 1981 in the lower stream of the ver The can brown in coloured, length. dense, Absolute age is Fig Large tusk of 22 Grote slagtand sing localities male a wolharige van een to woolly search mammoth mammoetstier for of remains the Mammoth Fauna is the New Siberian Islands. On Bolshoy Island, Lyakhovsky New Siberian Islands larger parts which are been found. Museum to the in the mammoth the three carcasses of the Mammoth 1994 and 1995 have also 500 skeletal parts over to at least Mammoth Museum have Expeditions during museum to 9 further archipelago, of mummified now belonging brought belonging species of the Mammoth Fauna. The investigations for remains of mammoth started in century. The first important collections of the 19th Fig 21 Part of the Pleistocene saiga antelope mammoth skull Island Gedeelte van de schedel van een Pleistocene Bunge saiga-antilope were made of by and Baron E. 1886. A 29 remains an Bolshoy Toll in the large monograph Lyakhovsky expedition on years of Dr. A. 1885 and the results of these Quaternary Mammal Collections in the Museums of Yakutsk (Eastern Siberia, Yakutia, Russia) M. paleontologist (1906). An carcass was the entirely were complete specimen in up and Lena van het (Chersky, 1891). tion under the place published by leadership of Baron E. collection quently Fig Toll of studied 24 Part of an bones by Early was the Holocene an collected took onal exten- made and subsefamous moose / quantities 30 of on and Tikhonov not mention mammoth mammoth hunters who have least that we each tusks are by professilicences for exportation of mammoth ivory worldwide. The tusks of the woolly mammoth, years old, condition, and Gedeelte & the New Siberian Islands times 50.000 Russian carcass large year, expedi- in 1900-1903. On the New Siberian Islands, including Bolshoy Lyakhovsky Island, sive Chersky I. we For refer (1964) Last but to carcass history Garutt need Another Russian Polar only outside Russia. (1999). has been (Paris, the woolly Vereshchagin investigations is (Fig. 25) to paard National mammoth details of the de mummie the Museum France), van including feet. This beautiful History Gedeelte extremely skin of the head and ended / in condition, good 23 Part of the Lena horse mummy Lyakhovsky Many soft parts of the carcass Fig complete in 1906 on dug out Bolshoy Island. Pavlova van een Vroeg are, for are even preserved example, up to some- in excellent used in the Holoceen elandkarkas ivory CRANIUM 21,1-2 in South East industry Japan Asia because of the lack of and (Hong Kong) ivory from (by (2004) CITES regulations protected) present day elephants. Acknowledgements We wish to thank: Mr. Bernard France, for his support in a visit to the writing this new Peter Lazarev, samples the Yakutsk, for by Isotope of Petersburg, Russia) manuscript the Netherlands, for 25 The Complete after in 2001; with us complete the Nether- (Zoological Museum, for critical reading mammoth Parijs. Veel the Mayhew, Voorburg, improving the dug English. out on mammoet, in 1906 opgegraven Naturelle in gene- Radiocarbon Groningen, David were der Plicht at van Research, lands; Dr. Alexei Tikhonov and Dr. which Dr. Hans Laboratory, University Fig paper provided radio-carbondating carried out Centre Saint paper of remains of Pleistocene mammals of Yakutia for rously Mande, Mammoth Museum and the other mentioned museums in this Dr. Buigues Saint (CERPOLEX/Mammuthus), zachte delen Bolshoy Lyakhovsky Island; op waren now Bolshoy Lyakhovsky eiland, in een uitstekende 31 conditie, in MNHN nu at Paris in het Museum National d’Histoire waaronder de hoofdhuid en de voeten Quaternary Mammal Collections in the Museums of Yakutsk (Eastern Siberia, Yakutia, Russia) Literature G. & G. Baryshnikov, pleistocene cave from Felidae) 2001. The Boeskorov, Tikhonov, A., lion, Panthera spelaea (Carnivora, Russia. Cranium 18,1: 7-24. Yakutia, L. on Wrangel S. Agenbroad, Vartanyan, of the mammoth Comparative analysis 2003. populations Island and the Channel Islands. Deinsea 9: 415-420. G. Boeskorov, 2001a. G., Woolly rhino ( Coelodonta distributionin Northeast Asia. Deinsea antiquitatis) V. Ukraintseva, V., 1993. Hot G., 2001b. G. Boeskorov, history of the On taxonomic studies of the Late Pleistocene horses of Eastern matter. Conference to the Brauner: 76-85. Odessa: of Late Pleistocene Yakutia. Acta Nauka, Vereshchagin, lions, Panthera (Leo) spelaea Cracoviencia Zoologica Moscow: of the Description by mammal collection found 172 pp. basement of of Northern Asia. In Russian. N. K. & A. N. in Tikhonov, Yakutsk, 39 1990. pp. In Russian. 223-227. 40, 2: N. K. & A. N. Tikhonov, 1999. Exterior of the mammoth. Cranium 16,1: 4-44. post-Tertiary New Siberian 1885-1886. Notes of Russian expedition Paleontologica Exterior of the Mammoth. Vereshchagin, 1891. Chersky, I., in Siberia. In Russian. 1997. New finds Lazarev, E. A., 1977. 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