Dinosauria Catalog - The Origins Museum Institute
Transcription
Dinosauria Catalog - The Origins Museum Institute
The Origins Museum Institute presents Dinosauria From The Genesis Exhibit THE LATE PALEOZOIC ERA from 345 to 250 million years ago Inundated by the inland invasion of the seas throughout the Mississippian (345 million years ago), the first division of the Carboniferous period, immense tropical forest regions were fossilized into deep carbon bands in the Earth’s crust. During the Pennsylvanian (310 million years ago), the remaining division of the period, reptiles diverged from their amphibious ancestors and began to proliferate on the land. By the Permian (280 million years ago), named for the discovery of fossils near the region of Perm in western Siberia, mammal-like reptiles (synapsids) had appeared, characterized by developments that would ultimately distinguish them from their reptile ancestors and lead to a new class of animals. All of the continental land masses of the Cambrian collided during the Permian, forming the supercontinent of Pangaea. A series of devastating ice ages were triggered by the encroachment of the supercontinent on both poles. The resulting widespread extinction that ended the Permian period and the Paleozoic Era was of such massive proportions that it has never been equaled, over 90 percent of all species vanishing. The decimation of the once dominant multitudes of mammal-like reptiles provided an ecological niche that favored the surviving reptile groups. 71. Class Reptilia, Subclass Synapsida Aerosaurus wellesi Early Permian, New Mexico 67. Subclass Labyrinthodontia, Order Anthracosauria Seymouria baylorensis Early Permian, Texas With a sturdy, terrestrial-type skeleton characteristic of reptiles and a skull exhibiting prominent ear notches characteristic of amphibians, Seymouria was long regarded as a primitive reptile and the closest link to their amphibian ancestry. Because related forms document the passage of a gilled larval stage, a characteristic thoroughly eliminating them from the reptilian class, Seymouria is now placed among the anthracosaurs (“Coal Reptiles”) by most authorities, thus including this distinctly terrestrial amphibian in the group of Permian labyrinthodont. Although Seymouria (named for the town of Seymour, Texas where it was discovered) differed very little from the Pennsylvanian ancestors of the reptiles, it left no known descendants. This exquisite skeleton is from the U.S. National Museum. The most primitive of the early synapsid reptile groups, the pelycosaurs may have led a somewhat amphibious existence, similar to that of alligators. Catching and killing its prey in jaws lined with an extraordinary number of sharp teeth, Aerosaurus (“Air Lizard”) was apparently an aggressive predator. Equipped with an unusually long and flat swimming tail, this young pelycosaur probably preyed on fish as well as on smaller, slower reptiles and amphibians. Although undoubtedly cold-blooded, because of their specialized bone structure the archaic synapsid reptiles are considered to be the ancestors of the therapsids, the later and more advanced reptile group that subsequently gave rise to warm-blooded mammals. Some scientists have also noted affinities between the Early Permian pelycosaurs and the later Permian freshwater mesosaurs, the possible forerunners of the air-breathing, marine ichthyosaurs. Collected by Charles Camp and Samuel Welles in 1928 and identified by Wann Langston and Robert Reisz in 1981, this cluster of Early Permian specimens also contains the dissociated bones of various other reptiles and amphibians including an Eryops and an Edaphosaurus. University of California at Berkeley. 70. Subclass Synapsida, Order Pelycosauria Dimetrodon limbatus Early Permian, Texas The aggressive, carnivorous Dimetrodon (“Dual Sized Tooth”), distinguished by a prominent dorsal fin along its back, prowled the upland regions of the swampy deltas of northern Texas along the edge of a shallow, drifting sea. The dominant predator of this area, Dimetrodon fed without competition on an abundant population of large, defenseless amphibians. Belonging to the archaic order of finback pelycosaurs, these early cold-blooded synapsids were ancestral to the mammal-like reptiles of later Permian times. Equipped with a ferocious arcade of palatine teeth (in addition to the sharp teeth that lined their jaws). Incapable of sustaining an extended chase, they probably hid in the lush vegetation, waiting for unsuspecting prey to stray too close to escape a quick, surprise attack. The creature’s impressive dorsal “sail” provided a large surface area for warming the blood when exposed to sunlight and cooling it when in the shade, although such spectacular features may evolve more rapidly for the purpose of mating displays than for thermal functions. By the Middle Permian, when the climate along the delta became too dry to sustain them anymore, the Dimetrodons disappeared quite suddenly from the fossil record. Brigham Young University. THE TRIASSIC PERIOD from 230 million to 195 million years ago There were no flowers or grasses at the beginning of the Triassic period (named for a tri-layered sequence of strata first discovered in southern Germany). Surrounded by a vast ocean, the supercontinent of Pangaea almost entirely circled the globe at the equator. Identical fossil remains found as far apart as Africa and North America suggest that the fauna of this period roamed freely across vast expanses o land that are now separated by the oceans. The Triassic marked the dawning of the Mesozoic (“Middle Life”) Era, or the Age of Dinosaurs. Because much of Pangaea lay along the equator, climates were generally tropical, and warm, ferny swamps teemed with primitive amphibians and crocodiles. In the arid desert regions, reptiles flourished with the extinction of mammal-like reptiles, producing the highly successful archosaur group which gave rise to crocodiles, pterosaurs, and a new kind of animal gradually appearing in the Late Triassic fossil record: the “Dinosauria” (“Terrible Lizards”), which continued appear in increasing varieties for the next 140 million years. The gradual breakup of Pangaea beginning at the end of the Triassic globally separated dinosaur populations along with the first true mammals, tiny burrowing descendants of the mammal-like reptiles. 76. Class Reptilia, Subclass Archosauria Euparkeria capensis Early Triassic, South Africa Crowding the archaic Permian protomammals out of their dominant position in the competition for food, the rapidly evolving Triassic reptiles known as thecodonts (“Socket Tooth”) were distinguished by the development of remarkably open skulls, hollow bones that were lightweight but very strong, and jaws lined with individually socketed teeth. This group of ancestral archosaurians (“Ruling Reptiles”) gave rise to the early saurischians and ornithischians of the Late Triassic (collectively known as the dinosaurs) as well as to crocodiles, birds, and the extinct flying reptiles. Exemplifying this important parent group, Euparkeria was a small creature that walked on 4 legs but, like the modem crocodile, was capable of bipedal sprinting due to its powerful hind leg muscles. A predator whose advantage over its prey was its swiftness, Euparkeria was extremely close to the immediate ancestry of the dinosaurs. This magnificent partial skeleton is from the famous Karroo Formation. South African Museum. 197. Class Reptilia, Subclass Archosauria Eoraptor lunensis Triassic, Argentina The earliest known dinosaur, the “dawn raptor” preyed on insects and small 4-legged animals. Although it was an efficient, bipedal predator, its primitive jaws lacked the “hinged” flexibility of its carnivorous successors. The Argentine Museum of Natural Sciences. 196. Class Reptilia, Subclass Archosauria Herrerasaurus Triassic, Argentina This early dinosaur was a primitive, bipedal carnivore with a flexible jaw capable of entrapping its struggling prey. Argentine Museum of Natural Sciences. 201. Class Reptilia, Subclass Archosauria Sharovipteryx mirabilis Triassic, Kyrgyzstan Formerly known as Podopteryx, this small gliding animal soared from tree to tree like a flying squirrel. Its name means “Sharov’s Wings.” It lived during the early Triassic period about 245 million years ago in Asia. With hind legs supporting a flight membrane, it appears to be an ancestor of the pterosaurs. From the Russian Paleontological Institute. 77. Class Amphibia, Order Temnospondyli Thoosuchus Triassic, Russia Still dependent on laying its eggs in water and undergoing a larval stage with gills for breathing (as its fishlike ancestors did when they first appeared during the Devonian Period), this small, bottom-dwelling amphibian, its skull deeply grooved with nervebearing canals, was only partially adapted to terrestrial life. Private collection. 78. Class Amphibia, Suborder Stereospondyla Aphaneramma Late Triassic, Arizona As a group, the Triassic stereospondyls, with their flat heads and upward gazing eyes, were primarily bottom-dwelling predators. Their weakly-limbed skeletons show an enigmatic regression in development compared to those of their more sturdily-limbed ancestors. Collected in the Moenkoepi Formation of Meteor Crater by Charles Camp, Samuel Welles, and Frank Peabody in 1938. University of California, Berkeley. 79. Class Amphibia, Suborder Stereospondyla Hadrokkosaurus bradyi Late Triassic, Arizona This large amphibian from the Moenkoepi Formation has the superficial appearance of a large frog. The pronounced indentation between the eyes, the pineal opening, sensitive to light and shade, was used as a “third eye” by early bottom-dwelling fish and amphibians. Derived from an ancient lineage of marine ancestors, the once photosensitive pineal body is still retained as a vestigial organ within the brains of all modern vertebrates. University of California, Berkeley. 80. Class Amphibia, Suborder Stereospondyla Cyclotosaurus Late Triassic, Arizona Resembling a giant salamander, this amphibian spent most of its life lying motionless on the murky bottoms of ancient lakes and rivers of southwestern North America, its tiny limbs useless for support on land. Often victims of drought, the fossilized remains of whole populations are sometimes found desperately crowded together in dried ponds. Derived from the labyrinthodonts of the Permian, they were extinct by the end of the Triassic. University of California, Berkeley. 81. Subclass Archosauria, Order Thecodontia Aetosaurus ferratus Late Triassic, Germany Resembling the armored dinosaurs of a later age, Aetosaurus (“Eagle Lizard”) actually belonged to the prolific thecodont group. Although some of these archosaurs were small, bipedal carnivores, others were broad, 4-legged herbivores with heavily armored bodies for protection against predators. Sporting an ominous battery of spikes and bony plates, the well-known Aetosaurus lived along the fringes of the Triassic marshes of Central Europe. Although closely related to the ferocious aquatic crocodilians and phytosaurs of its time, as evidenced by this mummified skeleton with a perfectly preserved hide, Aetosaurus was probably a passive animal, devoted mostly to uprooting vegetation with its blunt, upturned snout. Widespread throughout the northern hemisphere, aetosaurs are also found in Triassic formations of North America. Humboldt Museum. 82. Dinosaur trackway Grallator Late Triassic, Connecticut 195. Class Reptilia, Subclass Archosauria Pachypleurosaurus edwardsi Triassic, Switzerland This perfectly preserved in-situ skeleton belonged to a small nothosaur, a marine reptile originally descended from terrestrial stock, and direct ancestor of the giant plesiosaurs of the Jurassic. Private collection. Left in the mud by one of the earliest dinosaurs, this footprint of an ancient Triassic coelurosaur (“Hollow-Tailed Lizard”) gives no hint of the gigantic proportions of its descendants which were to follow. Ranging up to 10 feet in length, it ran on its hind legs counterbalanced by a long tail. This little theropod (“Beast Foot”), with its serpentine neck and lightweight, hollow bones, is believed to have been swift and agile, chasing prey and tearing it with sharp claws and teeth. Such behavior strongly suggests a warm-blooded metabolism, a theory popularized by the revolutionary paleontologist Robert Bakker. Among the first tracks to be found in North America, they were discovered in the Connecticut Valley during the early 19th Century by Edward Hitchock, who died convinced that he had found evidence of prehistoric birds. When later compared to verified Coelophysis tracks, they proved to be remarkably similar. Private collection. 83. Subclass Archosauria, Suborder Theropoda Coelophysis bauri Late Triassic, New Mexico Among the earliest of the dinosaurs to appear in the fossil record, Coelophysis (“Hollow Form”) was a long-legged, 3-fingered predator armed with long, slender jaws lined with sharp, serrated teeth. With grasping claws freed by its upright posture, Coelophysis was a formidable hunter, suspected of cannibalistic tendencies due to the presence of young skeletons found within the body cavities of some adult individuals that evidently fed on their own young. Inhabiting the Late Triassic coniferous forests, this graceful forerunner of the giant theropod dinosaurs of later ages (as well as the first birds) appears to have lived in wandering herds throughout northern New Mexico. Ranging from 3 to 10 feet in length and weighing up to 65 pounds, these sleek, swift creatures are known primarily from the remains of what appears to have been an entire herd that perished together, possibly in a sandstorm or during a flood. This spectacularly exposed in-situ skeleton, with the remains of a devoured juvenile, is from the famous Chinle Formation of Ghost Ranch. American Museum of Natural History. 84-85. Order Crossopterygii, Family Coelacanthidae Whitea Triassic, British Columbia For nearly 400 million years, these sluggish, passive creatures have survived virtually unchanged. Derived from the primordial group of freshwater fishes that produced the first terrestrial vertebrates and distinguished by its primitive limblike fins, this ancient carnivorous fish shows the preliminary bone structure that led to amphibious walking on land. Well-known from ancient freshwater and marine deposits, no fossil coelacanths (“Hollow Spine”) dating less than 60 million years have been found. Formerly believed to be long extinct, a rare extant species was discovered in 1938 off the Comoro Islands in the Indian Ocean. Inhabiting the ocean at depths exceeding 500 feet, these bright bluefish swim with a peculiar synchronized rotation of the pelvic and pectoral fins, which are capable of assuming nearly any position. Possessing an elastic notochord instead of a segmented backbone, the coelacanth, unlike most fish, bears its young live rather than by laying eggs. Although their deep habitat appears to be devoid of prey, dissected specimens reveal that they feed on fish and squid, perhaps surviving between feedings by slowing their metabolisms in the cold waters. Royal Tyrrell Museum of Palaeontology. 86-87. Order Therapsida, Infraorder Cynodontia Thrinaxodon liorhinus Early Triassic, South Africa The transition from synapsid reptiles to mammals was gradual rather than a quantum leap. Replacing gorgonopsids as the dominant predators of the Early Triassic, cynodonts were small, fox-like creatures with fewer reptilian characteristics than their ancestors. Although exhibiting tiny pits in their skulls which appear to have held whiskers and possessing a palate separating the nasal passages from the mouth, distinctly mammalian features, the primitive reptilian jaw structure of cynodonts retained certain prominent bones that became greatly reduced in mammals and incorporated into the inner ear. Extremely close to the ancestry of the mammals, this group included Thrinaxodon, an advanced therapsid of the forests of Antarctica and the South African Karroo. Equipped with prominent canine teeth as well as other dental features characteristic of early mammals, Thrinaxodon is widely regarded as the original stock from which they arose. In-situ skeleton and 2 skulls from the Karroo Formation. University of California at Berkeley. THE JURASSIC PERIOD from 195 million to 140 million years ago The Jurassic period, with its lush rainforests, derives its name from an abundantly fossiliferous sequence of chalky deposits discovered in the Jura Mountains bordering Switzerland and France. For the 70 million year duration of the Jurassic period, the supercontinent of Pangaea was gradually being wrenched into 2 separate masses, Gondwanaland (Africa, South America, Australia, India, Arabia, and Antarctica) and Laurasia (Europe, Asia, Greenland, and North America). Among the survivors of the Triassic extinction, tiny primitive mammals began to diversify during the Early Jurassic. By the dawning of the Late Jurassic (150 million years ago), the Atlantic Ocean had formed and the drifting continents had barely begun to resemble their present shapes. Gigantic dinosaurs that cared for their young had evolved from their smaller Late Triassic ancestors. Land bridges between the continents allowed the thriving herds of dinosaurs to migrate across great distances. Palmlike plants appeared and flourished throughout the warm, swampy landscapes of the period, nourishing the largest creatures that ever walked the Earth. 89. Suborder Theropoda, Infraorder Coelurosauria Compsognathus longipes Late Jurassic, Germany Of all the known fossils of adult dinosaurs, Compsognathus (“Elegant Jaw”) has the distinction of being the smallest. Running upright on its strong hind legs and stalking the underbrush for smaller Reptiles along the forested seashores of Jurassic Germany and France, Compsognathus was a swift and capable hunter, armed with sharp claws and teeth. This tiny coelurosaur was a relative of such giant creatures as the Tyrannosaurus rex of a later age. More closely related to Archaeopteryx (the bird-dinosaur), this skeleton shows that they shared a very recent common ancestor, suggesting that birds may have inherited their warm-blooded metabolisms from their dinosaur forebears. The smaller bones exposed within the rib cage of this well-known specimen have been identified as the undigested skeleton of Bavarisaurus, a tiny lizard that was devoured just before the predator died. With its neck and tail bent backwards, this Compsognathus (discovered in 1861) was long thought to have died in agony, although its posture is now attributed to the tightening of tissues as the carcass dried out prior to fossilization. From the famous lithographic limestone of Solnhofen. Bavarian State Institute for Paleontology and Historical Geography. These small, feathered dinosaurs inhabited the Late Jurassic forests of Central Europe 150 million years ago. Although adapted for gliding and presumably for limited flight, these primitive ancestral birds still retained efficient, grasping claws on each wing, as well as a long bony tail and jaws lined with sharp archosaurian teeth. perhaps the most famous fossil in the world, the spectacular “Berlin specimen” was found in 1877. Owing to the faint preservation of its flight feathers (overlooked for many years), the juvenile specimen, known as the “Eichstatt specimen,” was long misidentified as a Compsognathus. 90-92. Class Aves, Subclass Archaeornithes Archaeopteryx lithographica Late Jurassic, Germany The mounted skeleton, cast from the “London specimen” and featured in a PBS Nova program, was discovered in 1861. These rare specimens, the most complete ever found, are spectacular evidence of an intermediate stage between Reptiles and birds. Because its remains have only been found in deposits of shallow marine lagoons rich in fossilized fish and squid, some believe Archaeopteryx (“Ancient Wing”) was capable of diving for aquatic prey. Others envision it darting along the ground bipedally, using its wings as spoilers to increase its speed while chasing insects and small Reptiles or for soaring from tree to tree. From the famous lithographic limestone of Solnhofen. Berlin specimen from the Humboldt Museum. Mounted specimen from the British Museum. Juvenile specimen from the Jura Museum. 201. Class Aves, Subclass Archaeornithes Confusciusornis sanctus Jurassic, China 194. Class Aves, Subclass Archaeornithes Cathayornis yandica Jurassic, China Roughly contemporary with the European Archaeopteryx, this feathered theropod dinosaur from Asia more closely resembled modern birds. Beijing Institute of Vertebrate Paleontology. Possessing no tailbones, these early ancestors of modern birds lived in Asia . About the size of ravens, their discovery revolutionized long held concepts about the origins of birds. Beijing Institute of Vertebrate Paleontology. Capable of sustained flight as well as aerial acrobatics, Rhamphorynchus was probably furry and warm blooded. The leathery wing membrane, extraordinarily preserved in this specimen, was reinforced with a lacing of special tissue. From the lithographic limestone of Solnhofen. Humboldt Museum. 93. Superorder Archosauria, Suborder Pterosauria Rhamphorynchus gemmingi Late Jurassic, Germany The first vertebrate creatures capable of flight, the ancient pterosaurs (“Winged Reptiles”), were not on the evolutionary path that led to birds. Descended from early Triassic archosaurs and divided into 2 distinct groups by certain physical characteristics, the older genera were Distinguished by their long tails. Among the most primitive, Rhamphorynchus (“Beak Snout”) had a slender tail that ended in a kite- shaped membrane which presumably served as a rudder. Growing to lengths of up to 2 feet, its average wingspan was about 4 feet. With an elongated, flexible neck and extended jaws lined with sharp, forward- slanting teeth, this predator hunted along the same reefs and shorelines as Pterodactylus. Coiling its neck for diving and lunging at fish and squid, Rhamphorynchus speared its prey in the intermeshing barbs of its teeth, which allowed for no escape. hollow, lightweight bones enabled them to venture significant distances over the shallower waters of the ancient German seas, where a number of the remains have been recovered, exquisitely preserved along with impressions of their furry coats. From the lithographic limestone of Solnhofen. Humboldt Museum. 94. Superorder Archosauria, Suborder Pterosauria Pterodactylus elegans Late Jurassic, Germany Once presumed to have been limited to gliding from high perches, they are now believed to have been thoroughly agile in flight. They were quite vulnerable to marine predators, judging by the numerous pterosaur bones found in the stomachs of ichthyosaurs. The long, narrow snout of Pterodactylus may have been an adaptation to probing in the sand for burrowing worms. From the lithographic limestone of Solnhofen. Humboldt Museum. 94. Superorder Archosauria, Suborder Pterosauria Pterodactylus kochi Late Jurassic, Germany Throughout the Jurassic, leathery-winged pterodactyls (“Wing Finger”) would have been a common sight gliding on the tropical breezes of the European coastal regions, catching up insects, small fish, and squid in their long, sharply-toothed beaks. Pterosaurs belong to the archosaur order, which also includes dinosaurs and crocodiles. The first flying Reptiles to be discovered, the raven-sized Pterodactylus, like all pterosaurs, was distinguished by its elongated fourth finger which supported a powerful flight membrane. Their 95. Superclass Chelicerata, Subclass Xiphosura Mesolimulus walchi Late Jurassic, Germany Related to arachnids and the extinct eurypterids, the xiphosurans range from the Cambrian to the present. Derived from trilobites that had invaded estuaries and streams and lost their antennae (along with other specialized modifications), they are today represented by “living fossils” known as horseshoe crabs. Virtually unchanged since the Permian, these arthropods dig along the beaches for worms and other small prey, often burying themselves in the sand. Possessing a pair of compound eyes (as well as a pair of simple ones) and equipped with a spiny telson for correcting its position when overturned, horseshoe crabs undergo periodic molting. Reflecting their ancient ancestry, in fetal form they resemble trilobites. During the spring mating season, males cling to the telsons of the females as they crawl along the sandy shores. Dramatic evidence of this same behavior in ancient forms is provided by the well-documented Jurassic Mesolimulus, whose trails sometimes lead to the remains of the individuals that left them. From the lithographic limestone of Solnhofen. Private collection. 96. Order Ornithischia, Family Hypsilophodontidae Othnielia rex Late Jurassic, Utah Possibly the smallest of the horny-beaked hypsilophodonts (“High Crest Tooth”) of Late Jurassic Colorado and Utah, Othnielia, formerly known as “Nanosaurus rex” (“Dwarf Lizard King”) only grew to a length of 3 to 4 feet. Agile sprinters when threatened or chasing prey, the omnivorous hypsilophodonts possibly flourished for longer than any other dinosaurs, about 100 million years. After depositing their eggs in the sand, the females appear to have carefully manipulated them into spiral clutches of regular depths and even spacing. Although they probably remained with their herds throughout adulthood, these dinosaurs (owing to their welldeveloped limbs) apparently left their nests immediately upon hatching, probably feeding first on adult feces, rich in fermentative bacteria, and later progressing to plants and insects. As adults they fed on everything from plants to small Reptiles and mammals. “Nanosaurus” was renamed Othnielia in 1977, on the centennial of its discovery by the great pioneer paleontologist Othniel C. Marsh. Featured in the 1993 Universal Studios film Jurassic Park, this spectacularly exposed in-situ skeleton is from the Morrison formation of Emery County, Utah. Brigham Young University. 97. Subclass Ichthyopterygia, Order Ichthyosauria Stenopterygius quadricissus Jurassic, Germany 98. Class Reptilia, Subclass Ichthyopterygia Ichthyosaurus megacephalus Jurassic, Germany Flourishing throughout the Mesozoic, the ichthyosaurs (“Fish Reptiles”) appeared during the Early Triassic. Growing up to 15 feet in length, they apparently evolved from land-dwelling creatures, possibly the pelycosaurs. From the limbs of their terrestrial ancestors, the ichthyosaurs evolved their strong, broad paddles. Stenopterygius was Distinguished by its paddles, which contained more joints and fewer digits than those of other species. This magnificent specimen is preserved with an epidermal silhouette. Humboldt Museum. Typifying these classic marine Reptiles, Ichthyosaurus was Distinguished by the presence of extra digits with fewer joints in its paddle. From other fossils, these air-breathing, dolphin-like creatures are known to have fed on pterosaurs and fishes and, rather than laying eggs, gave birth to live young. This rare and extremely young Ichthyosaurus appears to have died at birth, although the displacement of its vertebra could easily have occurred after its death. Private collection. fossilized remains of bacterial parasites that inhabited the digestive tracts of the hosts millions of years ago. From the Jurassic of Chihuahua, Mexico. Private collection. 99. Coprolites Ichthyosaur Jurassic, Mexico Found in deposits rich in ichthyosaur remains, these coprolites (“Dung Stones”) are the fossilized excrement of large, dolphin-like marine predators. When examined microscopically, coprolites can reveal not only the fibers of the digested matter, but also the 100. Subclass Euryapsida, Order Sauropterygia Plesiosaurus macrocephalus Jurassic, Scotland Derived from small Triassic nothosaurs similar to Mesosaurus, these gigantic marine Reptiles, ranging from 6 to 40 feet in length, flourished throughout the Jurassic Period in the oceans of Europe and North America. Their small heads were equipped with a battery of needle-like teeth for catching fish, with elegantly long, serpentine necks capable of darting and striking. Adapted for “underwater flight”, these formidable predators glided through the water by flapping their winglike paddles up and down in the manner of penguins. Originally descended from terrestrial stock, plesiosaurs (“Ribbon Reptiles”) were air-breathing and had to haul their heavy bodies onto the beaches to lay their eggs. First appearing in the Late Triassic, plesiosaurs gradually tapered to extinction by the end of the Cretaceous. This rare, in-situ juvenile specimen is from the Jurassic of Lyme-Regis, England and was found by Mary Anning. Royal Scottish Museum. 101. Order Saurischia, Suborder Sauropodomorpha Apatosaurus ajax Late Jurassic, Utah The regal, long-necked sauropods were by far the largest land animals that ever lived. With a long, sloping skull and a single nostril situated on top of its skull, Apatosaurus (“Mystery Lizard”) was a member of the diplodocid family, shorter in length but much heavier than the closely related Diplodocus. Measuring 70 ft. in length and weighing 33 tons, these gigantic herbivores possessed long, serpentine necks and even longer whiplike tails. Because the first skeletons discovered were not found associated with any skulls, this dinosaur was originally described as “Brontosaurus” (“Thunder Lizard”), a name which persisted until the genus was finally correctly identified as Apatosaurus, previously known only from a skull that had been named before the discovery of the headless Brontosaurus skeletons, hence its name prevails. For a long time some museums continued to traditionally exhibit their Apatosaurus skeletons with the heads of Camarasaurus. Known only from North American specimens, they lived in vast herds that ranged throughout Utah, Wyoming, Colorado, and Oklahoma. This classic leg bone was recovered from the famous Morrison Formation. Brigham Young University. 102. Order Saurischia, Suborder Sauropodomorpha Camarasaurus lentus Late Jurassic, Utah Herds of Camarasaurus (“Chamber Lizard”), a heavy sauropod (“Lizard Foot”), with a shorter neck and tail and a blunter snout than Apatosaurus, thrived in the Late Jurassic Colorado, Oklahoma, Utah, and Wyoming. Unique among the sauropods, the large nasal cavities on top of its head, similar to those found in elephants and tapirs, suggest the possibility o a flexible trunk. They grew to 60 feet in length and weighed about 20 tons. Discovered ill 1877 by the pioneer palaeontologist Edward Drinker Cope, these giant animals were once presumed to have been semi-aquatic swamp dwellers, too heavy to walk about on dry land. Camarasaurs and other sauropods are now believed to have avoided such murky places in favor of the drier plains, possibly migrating across vast distances. Sauropod footprints indicate that these gentle herbivores cared for their young, guarding them on all sides as they wandered together, feeding on the highest tree branches. Found in the famous Cleveland Lloyd Quarry. Carnegie Museum. 104. Order Saurischia, Suborder Sauropodomorpha Hypselosaurus priscum Cretaceous, France This 40 foot long sauropod, characterized by its short, blunt head and small teeth, laid the largest eggs known, about 12 inches in length. The folded embryo within would have hatched to a size twice the length of the egg. Larger eggs probably would have required a shell too thick for hatching. Hypselosaurus (“High Crested Lizard”) is the only sauropod dinosaur that has ever been found in association with its eggs. Private collection. 105. Gastroliths Sauropod Late Jurassic, Utah Because their teeth were adapted for cropping but not for chewing the tons of plants they required daily, sauropods and other dinosaurs deliberately swallowed stones to help grind food to a digestible pulp in their gizzards. Smoothly polished gizzard stones have been found to originate from rock sites as much as 15 miles from their final resting place among the bones of a dead dinosaur, suggesting that these creatures, were somewhat particular about the types of rocks they ate. Private collection. Aiding in the precision of the movement of the enormous hindquarters and spiked tail, an enlargement of the spinal cord at the pelvis, much larger than the creature’s brain, was long believed to be its “second brain.” For its bulk, Stegosaurus had the smallest skull of any dinosaur, with a brain no larger than a walnut. Its front legs, only half the length of its back legs, indicate that it probably evolved from a 2-legged ancestor and could easily have reared on its strong hind limbs to feed on the higher tree branches. The deadly spikes attached to the tip of its powerful tail were used as a defense against predators in Late Jurassic Colorado, Oklahoma, Utah, and Wyoming. Found in the Cleveland Lloyd Quarry. U.S. National Museum. 106. Order Ornithischia, Suborder Stegosauria Stegosaurus stenops Late Jurassic, Utah Discovered in Colorado in 1877, Stegosaurus (“Plate Lizard”) grew to a length of up to 30 feet and weighed up to 2 tons. The bony armored plates guarding the neck, back, and tail of the huge Jurassic herbivore were attached to powerful skin muscles, and were probably capable of being raised and lowered to protect the spine or the flanks. Honeycombed with capillaries, they may also have served as a heat exchange for warming (or perhaps cooling) the animal’s bloodstream, although their actual placement and function remain an enigma. 107. Order Saurischia, Suborder Theropoda Allosaurus fragilis Late Jurassic, Utah Its massive, yet open, lightweight skull a marvel of engineering, the fearsome Allosaurus (“Different Lizard”) was the predominant carnivore in North America during the Late Jurassic, and probably fed on such herbivorous dinosaurs as Apatosaurus (Brontosaurus) and Stegosaurus, its flexible skull capable of expanding to accommodate oversized chunks of food. While some believe that they were too huge and clumsy to hunt, scavenging on carrion instead, others believe them to have been agile runners and swimmers, capable of hunting in packs for large prey. Trackways found in Texas indicate that such predators indeed chased herds of much larger dinosaurs, and Apatosaurus vertebrae, scarred by the teeth of an Allosaurus, have been unearthed. Sporting strong, 3-fingered claws at the ends of its short arms, this creature grew to 36 feet or more in length and weighed 1 to 2 tons, possibly reaching full maturity in as little as 5 years. Although they have been found in Africa, Australia, and perhaps even Asia, Allosaurus are most common in North America, some 40 individuals coming from a single quarry (possibly having perished together). Found in the Cleveland Lloyd Quarry. 193. Order Saurischia, Suborder Sauropodomorpha “Ultrasaurus” Late Jurassic, Utah This massive vertebra of a giant brachiosaur is from the upper neck of one of the largest sauropod dinosaurs ever found, a gentle herbivore. Brigham Young University. 108. Order Ornithischia, Family Camptosauridae Camptosaurus browni Late Jurassic, Utah A primitive ornithopod (“Bird Foot”), ranging in length from 12 to 23 feet and weighing up to 1000 pounds, Camptosaurus (“Bent Lizard”) was much larger and less agile than its hypsilophodontid ancestors. Short but sturdy, its 5-fingered forelimbs were adapted to bearing weight, although fossilized footprints indicate that it was also thoroughly capable of walking bipedally on its 4-toed hind legs. Found in the Cleveland Lloyd Quarry. 109. Order Ornithischia, Family Camptosauridae Camptosaurus browni Late Jurassic, Utah Cropping low plants with its beaked jaws, this peaceful grazer lived in western North America and Europe from the Late Jurassic to the Early Cretaceous, suggesting a former connection of the 2 continents. The ancestor of the famous Iguanodon, this important animal is believed to have given rise to the diffuse groups of duckbilled dinosaurs which spread throughout the world in Cretaceous times. Found in the Cleveland Lloyd Quarry. THE CRETACEOUS PERIOD from 140 million to 65 million years ago Marking the appearance of flowers, the fertile Cretaceous period was named for vast deposits of chalk (creta in Latin), rich in fossils and widely distributed throughout the world. The distribution of the continents was beginning to resemble that of the present, although a great shallow sea flowing from Canada to the Gulf of Mexico separated the supercontinent of Laurasia into Asiamerica (East Asia and Western North America) and Euramerica (Europe and Eastern North America). Marked by a rich emergence of advanced dinosaurs, the Late Cretaceous, 75 million years ago, was a time of tremendous volcanic upheaval as mountain ranges and new islands were being formed. The drifting of the continents toward the polar regions brought on significant changes in the seasonal weather. No longer able to roam between the continents, many of the later dinosaurs evolved exclusively in Asiamerica. The dominant creatures of the period, most species lasted no more than 5 million years before disappearing from the fossil record, often followed by more advanced descendants. This period lasted until 65 million years ago, ending with a catastrophic mass-extinction second only to that of the Permian. Nearly half of all the Cretaceous plants and animals, both marine and terrestrial, suddenly vanished. Among them were the dinosaurs. 110. Infraclass Teleostei, Order Leptolepiformes Cearana Early Cretaceous, Brazil The appearance in the fossil record of the heterogeneous group of leptolepids marks the arrival of the modern teleost, or bony, fishes, which now dominate the world’s aquatic environments. Armed with a mouthful of small, sharp teeth, these elongate, tapering fish became widespread throughout the Late Jurassic and Early Cretaceous. Spectacularly preserved in mud concretions, Cearana is a welldocumented variety known to have preyed on its own species, some individuals having died in the process b):: choking on prey too large to swallow. Unlike more primitive fish with primarily cartilaginous skeletons, such as sharks and lobe-fins, the early teleosts possessed fully ossified internal skeletons. From the famous Santana Formation of Brazil. Private collection. 111. Phylum Mollusca, Class Cephalopoda Texanites texanum Late Cretaceous, Texas Gregarious creatures, the ammonoids lived in large schools near the bottom of deep seas, avoiding areas occupied by seaweed, clams and snails. Preying on fishes and invertebrates (including other ammonoids) captured with their tentacles, they fed through a beak resembling that of a parrot. Related to octopus and squid, their chief defense against predation was to squirt ink in the faces of their predators, which included large crabs, fish, and marine reptiles. In the deeper seas that once extended westward from the Gulf of Mexico, ammonoids were plentiful, although gigantic fossil forms such as Texanites and even larger varieties are extremely rare. These long enduring creatures disappeared abruptly, along with the dinosaurs, at the end of the Cretaceous Period 65 million years ago. From the famous Austin Chalk of Texas. Private collection. 1914 by Barnum Brown, this rare and highly specialized group of theropods was not fully understood until the discovery by John Ostrom in 1964 of Deinonychus, an early dromaeosaur succeeded in the Late Cretaceous by Dromaeosaurus (the first such dinosaur to be discovered), and the Mongolian Velociraptor. Because Dromaeosaurs (“Running Reptiles”) display a striking affinity to birds, particularly the Jurassic Archaeopteryx, there is little doubt that they were warm-blooded. Its brain had unusually large cerebral hemispheres for a dinosaur, suggesting complex behavior more birdlike than reptilian. Of the few fossilized battles between dinosaurs that have ever been found, two involved Dromaeosaurs, providing spectacular evidence of their behavior. One find in Montana indicates that they hunted in packs, while a one-on-one battle to the death with a Protoceratops was discovered in the Gobi Desert. Their combination of intelligence, swiftness, and the sickle claw places these creatures among the most dangerous of the dinosaurs. Royal Tyrrell Museum of Palaeontology. 112. Infraorder Deinonychosauria, Family Dromaeosauridae Dromaeosaurus albertensis Late Cretaceous, Canada Popularly known as a “Raptor”, this ferocious creature was armed with one lethal, sickle-shaped claw on each foot, held in a retracted position when it walked and extended for slashing the bellies of its victims with multiple kicks of its powerful hind legs. Discovered in 193. Infraorder Deinonychosauria, Family Dromaeosauridae Velociraptor mongoliensis Late Cretaceous, Mongolia This small but extremely vicious Chinese dromaeosaur, armed with the lethal sickle-claw, has been found preserved in deathlock combat with the early ceratopsian dinosaur Protoceratops. From the Polish-Mongolian Expedition and From the Russian Paleontological Institute. 113. Order Ornithischia, Family Hadrosauridae Prosaurolophus maximus Late Cretaceous, Alberta 193. Infraorder Deinonychosauria, Family Dromaeosauridae Deinonychus antirrhopus Late Cretaceous, Mongolia These large, swift, dinosaurs slashed their prey with sharp, sicklelike toe claws. The small Mongolian Velociraptor was followed by the larger North American Deinonychus. From the Harvard Museum. This Late Cretaceous duckbilled dinosaur, with its broad, flat muzzle and its small nasal crest, is presumed to have been an ancestor of Saurolophus (“Crested Reptile”), the forerunner of the prominently crested Parasaurolophus (“Beside Saurolophus”) of North America. These skull crests, both large and small, might have served as resonators for bellowing and honking during mating competition. Living in herds, the 26 foot long Prosaurolophus (“Early Crested Reptile”) was herbivorous and may well have fed and looked after its young. Royal Ontario Museum. 114. Order Ornithischia, Family Hadrosauridae Parasaurolophus walkeri Late Cretaceous, Alberta Unlike the earlier hadrosaurs, whose skulls were distinguished by small crests of solid bone, this advanced duckbill sported a prominent hollow crest extending from the back of its head by as much as 6 feet. This tubular crest, which may have been connected to the creature’s back by a fleshy frill, is believed to have functioned as a resonating chamber for mating calls and warnings. With chambers connecting to the nasal passage, the crest was long regarded as a snorkel for the storage of air while the animal was submerged, however a better understanding of the habitats of Parasaurolophus, a forest-dwelling dinosaur, has dispelled its popular image as a swamp dweller. Although the function of its spectacular crest is not fully understood, It is unlikely to have been used for defense due to its hollow structure. Ranging up to 33 feet in length, Parasaurolophus was indigenous to New Mexico, Utah, and Alberta. Royal Ontario Museum. 192. Order Ornithischia, Family Hadrosauridae Edmontosaurus regalis Late Cretaceous, Alberta This classic hadrosaur, with its massive skull, was among the largest of the duckbilled dinosaurs. Royal Ontario Museum. 115. Order Ornithischia, Family Hadrosauridae Anatosaurus annectens Late Cretaceous, Montana There were numerous species of Anatosaurus (“Duck Reptile”), also known as Edmontosaurus and Trachodon, that flourished up until the very end of the Cretaceous Period. Rare mummified hides of some of these classic duckbilled dinosaurs have preserved parts of their skin in minute detail. An unprecedented discovery in Alaska yielded the frozen, unfossilized remains of a related variety long buried in the permafrost, providing possible DNA samples as well as evidence that the present tundra region was once a subtropical, swampy home to these gentle dinosaurs. This rare skin impression of the underbelly from the mummified remains of an Anatosaurus was found in the famous Hell Creek Formation. American Museum of Natural History. 116. Order Ornithischia, Family Hadrosauridae Anatosaurus annectens Late Cretaceous, Montana This fibular fragment with a splendidly preserved skin impression is from a partially mummified duckbilled dinosaur found in the Hell Creek Formation. Private collection. times than they are now. Known for protecting and feeding their young, these duckbills may have relied upon their prominent head crests for recognition during their mating season. Although they possessed no significant defense against predators, their keen senses of vision, hearing, and smell served to warn them of approaching danger. Females and immature individuals had smaller crests. Like other crested duckbills, when Corythosaurus called to each other the sound resonated through the long air passages of their nostrils (which extended into their hollow crests) producing a far-reaching, bugle-like honk. Lined with blood vessels, such extended nasal passages also may have evolved for the purpose of warming the arctic air before its arrival into the lungs. From the Royal Ontario Museum. 117. Order Ornithischia, Family Hadrosauridae Corythosaurus casuarius Late Cretaceous, Alberta The crested Corythosaurus (“Helmet Lizard”), with its short muzzle and tall, hollow crest, lived in herds throughout the forests of western Canada where it browsed on tree leaves and pine cones. Equipped with strong, bulging cheeks, it was capable of chewing a mouthful of food while storing another. Growing to lengths of 33 feet and weighing over 4 tons, these peaceful herbivores ranged north of the Arctic Circle in search of food and migrated south during the long winters, which were less cold in Late Cretaceous 119. Coprolites Hadrosaur Cretaceous, Canada These coprolites were discovered in association with the fossilized remains of duck-billed dinosaur nests in Saskatchewan. Private collection. 118. Order Ornithischia, Family Hadrosauridae Maiasaura peeblesorum Late Cretaceous, Montana These gentle duckbilled dinosaurs left in the fossil record a very detailed picture of their way of life. Herds of Maiasaura (“Mother Lizard”) appear to have nested together in vast colonies along the shores of Cretaceous rivers of Montana. Juveniles and hatchlings of varying ages have been found in these nests, exhibiting evidence of prolonged parental care. The eggs were carefully arranged in regular patterns in large, dug-out mounds, each an adult body length from the next. Adults, which grew to lengths of 30 feet, appear to have returned to these nesting sites year after year. Browsing on their hind legs among the tree branches or cropping the undergrowth on all fours, these peaceful herbivores were defenseless against the ferocious weapons of their predators. Migratory hadrosaurs such as Maiasaura appear to have fed and protected their young until they were old enough to join the herd. In situ juvenile skeleton. Private collection. Nestling discovered in 1978 by John R. Homer. Museum of the Rockies. 123. Order Saurischia, Family Ornithomimidae Struthiomimus altus Late Cretaceous, Canada Prowling along the Cretaceous riverbanks of Southern Alberta and New Jersey, herds of predacious Struthiomimus (“Ostrich Mimic”) may have hunted everything from fruit and insects to eggs and small animals. Its toothless beak resembling that of a bird, Struthiomimus had sharp 3-fingered claws that were capable of grasping as well as digging. Although large, it was slender and built for swift running on its powerful hind legs, balanced by a long tail. Commonly known as ostrich dinosaurs, the ornithomimids ranged from North America to Mongolia. Equipped with unusually large brains, these creatures were among the more intelligent of the dinosaurs. Their particularly large eyes imply that they were highly adapted to hunting at night, when the small rodentlike mammals emerged from their hiding and other dinosaurs were hampered by the darkness. The ostrich dinosaurs are exemplified by Struthiomimus, whose strong hands appear to have been the most powerful of this widespread group. Because of their formidable digging abilities they are largely regarded as nest-robbers, feeding on the buried eggs of other dinosaurs and occasionally on tender hatchlings. Royal Ontario Museum. 124. Order Saurischia, Suborder Theropoda Oviraptor philoceratops Late Cretaceous, Mongolia In Mongolia in 1922, an Expedition from the American Museum of Natural History led by Roy Chapman Andrews set out to find remnants of early humankind. Instead they discovered Museum. several nests of Oviraptor (“Egg Thief”) eggs, the first dinosaur eggs ever located, in what must have been a large nesting ground. Because skeletons of the small, frilled dinosaur Protoceratops were found in association with the nests, the eggs were long attributed to them. More recent expeditions have uncovered a fossil female Oviraptor (a small, beaked, and toothless theropod) actually sitting on a clutch of unhatched eggs, revealing that Protoceratops was the intruder. From the historic Roy Chapman Andrews Gobi Expedition. Alf Museum. 191. Order Ornithischia, Suborder Ceratopsia Psittacosaurus Late Cretaceous, Mongolia This small, swift plant eater had a narrow beak and short horn-like bumps on its cheeks. The name of this primitive, early ceratopsian meant “Parrot Lizard.” Ancestral to the giant horned dinosaurs, it lived in Mongolia during the early Cretaceous period between 120 and 98 million years ago. From a private collection. 191. Order Ornithischia, Suborder Ceratopsia Protoceratops andrewsi Late Cretaceous, Mongolia This in-situ egg nest was found in association with the small frilled dinosaur, a social, sharply-beaked herbivore ancestral to the giant horned dinosaurs. From the Polish-Mongolian Expedition. 190. Order Ornithischia, Suborder Ceratopsia Protoceratops andrewsi Late Cretaceous, Mongolia Regarded as the earliest known horned dinosaur (although it only possessed small crests where the great horns of its giant descendants would be), this adult skull and exquisite mounted skeleton of a rare nestling are from the Polish-Mongolian Expedition. 125. Order Ornithischia, Suborder Ceratopsia Chasmosaurus belli Late Cretaceous, Alberta Of the long-frilled ceratopsian dinosaurs of the Late Cretaceous, Chasmosaurus (“Ravine Reptile”) is the earliest known. Extending from the back of the creature’s head, a sturdy, spiked frill protected its neck and part of its back. Large openings in the frill reduced the weight of the bone mass. It had 2 small horns over the brows and one on its parrot-beaked snout. Living in large, migrating herds and measuring over 16 feet in length, these herbivorous grazers have been found in Alberta, New Mexico, and Texas. Though not as wellequipped as its descendant, Triceratops (“Three Horned Face”), Chasmosaurus was capable of warding off attacks by fierce tyrannosaurs. Females of the genus are distinguished by their diminutive horns. This impressive creature was one of the first dinosaurs to be found along with an impression of its skin, the patterns of its large mosaic scales suggesting contrasting color patterns. Despite their obvious abundance throughout the Late Cretaceous, the ceratopsians, along with the rest of the dinosaurs, disappeared entirely at the end of the Cretaceous Period, 65 million years ago. Colossal skull from the Royal Ontario Museum. Distinguished by thickly domed skulls rimmed with bony spikes, rival males are believed to have established dominance over their herds through fierce contests of head-butting. This supposition is further evidenced by their neck and backbones, which were specially reinforced with strong ligaments for absorbing and distributing severe shocks to the head. Ancestral to the larger and more advanced Pachycephalosaurus of North America, Stegoceras may have evolved from an ancestor of such horned dinosaurs as Triceratops and migrated from Mongolia before the continents divided. Royal Ontario Museum. 126. Order Ornithischia, Family Pachycephalosauridae Stegoceras validus Late Cretaceous, Alberta Among the rarest of dinosaur remains, the enigmatic boneheads are known mostly from fragments of skulls and skeletons, making their relationships to other ornithischian groups difficult to prove. Growing to lengths of over 6 feet and weighing up to 120 pounds, Stegoceras (“Horny Roof”) had keen vision and an acute sense of smell. Possessing sharp teeth, this slow-moving bipedal creature is believed to have fed on insects, fruits, and leaves. Roaming in herds that ranged throughout the hills of eastern Asia and western North America, the boneheads apparently had lifestyles similar to those of modern sheep and goats. 125. Order Ornithischia, Suborder Ceratopsia Triceratops horridus Late Cretaceous, Wyoming These horn cores belonged to a Triceratops, the last and the largest of the many frilled, horned dinosaurs. Private collection. 127. Suborder Theropoda, Family Tyrannosauridae Albertosaurus sarcophagus Late Cretaceous, Alberta Sometimes known as Gorgosaurus, Albertosaurus (“Alberta Lizard”) was a smaller, slender tyrannosaur (“Tyrant Lizard”) that only grew to a length of about 26 feet and a weight of 2 tons. Despite its size in comparison to its larger relatives, it was a ferocious predator that probably hunted the herbivorous horned and armored dinosaurs of the Late Cretaceous lowland forests of Alberta. Presumably descended from a small, Late Triassic theropod and named for the province in Canada where it was discovered, Albertosaurus was also common in Montana just before the end of the Mesozoic Era. Various names have been given to the dozens of Albertosaurus skeletons discovered during the past century, but only recently has their relation to each other been recognized. Like all tyrannosaurs, Albertosaurus only had 2 fingers on each claw, the vestigial third finger a useless remnant of its 3-fingered Jurassic ancestors. Spectacularly complete in-situ skeleton of an extremely rare juvenile individual. Royal Tyrrell Museum of Palaeontology. 128. Superorder Archosauria, Suborder Pterosauria Quetzalcoatlus northropi Late Cretaceous, Texas Named for the ancient Mexican feathered serpent-god Quetzalcoatl, the giant Quetzalcoatlus was estimated at twice the size of the largest pterosaurs known at the time, the Pteranodons of the Kansas Chalk. Only one wing was found, indicating a wingspan of 36 to 39 ft. with a weight of 190 to 220 lbs., making it the largest known flying creature. With a low, narrow crest and a long, sharp, toothless beaks, this giant is better known from smaller, more revealing specimens found in the same region, which were either younger individuals or a smaller species. Although they were dynamically suited for fishing over the sea, their remains are not found in marine deposits or even near any sizable freshwater lakes, but are instead found in the silts of the vast floodplain of an ancient inland system of stream channels over 250 miles from the nearest sea, suggesting the possibility of their having been scavengers of carrion instead. Similar discoveries in Canada and Israel indicate the possibility of a wide distribution. This colossal humerus, discovered in 1971 by Douglas A. Lawson and Wann Langston Jr., belonged to one of the last of the flying reptiles. Texas Memorial Museum. 189. Superorder Archosauria, Suborder Pterosauria Pteranodon sternbergii Late Cretaceous, Kansas Among the last of the pterosaurs, this small-crested female individual had a 12-foot wingspan. Gliding over the inland sea, these creatures were adapted for fishing. This mounted skeleton is from Lane County, Kansas. Private collection. 129-132. Suborder Theropoda, Family Tyrannosauridae Tyrannosaurus rex Late Cretaceous, Montana One of the largest known predators to have walked the Earth (the females distinctly larger than the males), this invincibly massive, ferocious carnosaur of Late Cretaceous western North America and Asia was discovered in 1902 by the adventurous collector Barnum Brown. Surpassed in size only by the South American carnosaur Giganotosaurus, Tyrannosaurus rex (“Tyrant Lizard King”) grew to lengths of as much as 50 feet, towering 20 feet high and weighing 5 to 7 tons. Their heads alone measured 4 to 5 feet, sporting jaws lined with long, saw-edged teeth, each one supported by ranks of replacements. Reaching maturity within 5 years, the life spans of these enormous creatures may have surpassed 100 years. Fossil evidence of their actual metabolism is magnificently preserved in the sequential rings of growth-spurts which scar the teeth. While strictly regarded by some as nothing more than scavengers, owing to their greatly reduced forelimbs which could not even reach their mouths, others consider these creatures the most fearsome hunters ever to have existed, balanced for swift running by their long tails. Presumed to have dealt mostly with the animal’s senses, the brain of Tyrannosaurus rex was not small for a dinosaur although compared to the size of the creature’s body it was remarkably diminutive. This cranial endocast preserves the size and shape of the brain of a Tyrannosaurus rex. This gigantic foot, discovered in the famous Hell Creek Formation, is from the Natural History Museum of Los Angeles County. This isolated maxilla, once the largest known, is from the University of California, Berkeley. This spectacular skull, from one of the largest skeletons ever found, is from the Museum of the Rockies. Tarbosaurus, but owing to the minimal differences between its features and those of Tyrannosaurus rex, its name has frequently been assigned to that genus instead. Because of the immense size of these creatures, paleontologists also disagree about whether these fierce dinosaurs were active predators or strictly scavengers. Defending her nest of eggs (exceedingly rare fossils discovered in association with the species) this spectacular mounted skeleton is from the Polish-Mongolian Expedition. 188. Suborder Theropoda, Family Tyrannosauridae Tarbosaurus bataar (Tyrannosaurus bataar) Late Cretaceous, Mongolia First appearing in Central Asia around 74 million years ago, the tyrannosaurid family subsequently spread to North America where they flourished until the end of the Mesozoic Era. Smaller and more ancient than Tyrannosaurus rex, the Asian species is often assigned its own genus Tarbosaurus (“Alarming Reptile”). Discovered in 1955 in the Bataar region of Mongolia, it was immediately named contain only impressions of the decomposed remains of insects, amber in fact contains the actual preserved remains of its hapless victims. In 1992 revolutionary researchers at the University of California, Berkeley and the American Museum of Natural History in New York successfully recovered and cloned DNA from 30million-year-old insects preserved in amber. Suspended in amber for over 20 million years, these termites and tiny flies are from the Miocene of Colombia and the Domenican Republic. Private collection. 152. Insects in Amber Early Miocene, Kenya, Colombia, Domenican Republic Trapped in resin that oozed from the trunks of ancient conifer trees before hardening into amber, fossil insects are found throughout the world, preserved to the tiniest bristles, including their last exhalations and excretions. Related to the petroleums, amber is usually clear and yellow in color. Lumps containing insect inclusions provide an important record of the prehistoric faunas of the locales and geological periods in which they are found. Formerly thought to Cretaceous deposits and those of the Tertiary period, apparently the global fallout from a devastating collision with a large extraterrestrial body. Heat trapped on the planet’s surface by the dense atmospheric debris may have raised global temperatures, killing off the majority of land and sea organisms in a single catastrophic event. The gigantic Xixulub crater in Yucatan, Mexico was blasted by such an impact around the time of the Cretaceous extinction. 133. Dinosaur footprint Carnosaur Late Cretaceous, Utah This foot impression of a Cretaceous carnivore was found in the ceiling of a Utah coal mine. By measuring the height of the leg and the length of the stride, paleontologists are able to calculate from trackways of such footprints the speed at which the creatures may have been traveling, often indicating high, warm-blooded, metabolisms. Private collection. 134. Meteorite Meteor Crater, Arizona Since 1978, the extinction of the dinosaurs has largely been attributed to cosmic intervention in the form of an enormous meteorite impact on the Earth 65 million years ago. Discovered by scientists Walter and Luis Alvarez, Frank Asaro, and Helen Michel, a thin layer of the exceedingly rare element iridium (commonly found in meteorites) forms a boundary worldwide between Most meteorites come from the vicinity of the asteroid belt, the orbiting fragments of metal and rock which are the remnants of a failed planet and of the actual creation of our solar system some 4.6 billion years ago. About 22,000 years ago this meteorite, composed of 95% pure iron, left a crater over 3,600 feet wide and 525 feet deep. A fragment of the original meteorite, this pristine relic of the formation of the solar system was found in the famous depression of Meteor Crater, Arizona. Private collection. The Genesis Exhibit The Great International Fossil Collection is proud to include casts of important fossils housed in the following venerable collections UNITED STATES National Museum of Natural History (Smithsonian Institution) American Museum of Natural History Carnegie Museum Natural History Museum of Los Angeles County University of California, Berkeley University of California, Los Angeles Brigham Young University University of Texas, Austin University of Nebraska State Museum Harvard University Peabody Museum, Yale University Monroe Community College Page Museum Alf Museum Museum of the Rockies CANADA Royal Ontario Museum Royal Tyrrell Museum of Paleontology National Museum of Natural Science EUROPE AND GREAT Britain Musee deL’Homme University of Bonn Humboldt Museum Bavarian State Institute of Geology and Paleontology Vienna Natural History Museum Swedish Museum of Natural History British Museum of Natural History Royal Scottish Museum AS1A~ Australia~ AND THE MIDDLE EAST Beijing Institute of Vertebrate Paleontology and Paleoanthropology University of Adelaide Geological Survey of Pakistan AFRICA Kenya National Museum South African Museum Transvaal Museum National Museum of Tanzania Cairo Geological Museum National Museum of Ethiopia PRAISE FOR The Genesis Exhibit “After 65 million years, they’re back. In a BIG way.” BOSTON HERALD “Now underway at the World Trade Center in Boston... the entire story of life on Earth as represented by the fossil record.” THE TAUTON GAZETTE “This fascinating presentation...has educated and enthralled all who have been fortunate enough to view it.” MASSACHUSETTS GOVERNOR WILLIAM F. WELD “The only exhibit of its kind in the world... The Genesis Exhibit makes no bones about theatrical approach... when it comes to drama, intrigue and excitement, it’s tough to beat the theater of life.” AUSTIN AMERICAN STATESMAN “A window with a view into the most remote prehistory of man and animals...the unique feature of The Genesis Exhibit is that the casts are displayed as sculpture...quite different from the paleontological exhibits one usually sees in museums...response from the public continues to be surprise and pleasure from lay people and scientists alike.” HILL COUNTRY LIVING “To see the originals you would have to travel allover the world... an unprecedented collection of dinosaur fossil casts!” DISTINCT MAGAZINE “A new breed of both art and science... it’s amazing to see...” WESTLAKE PICAYUNE “It’s thrilling to see children get an idea of how really big... some of these creatures were... wonderful creations, almost works of art, because that’s what they look like at first glance.” HILL COUNTRY NEWS “Without a doubt, this exhibit provides unmatched educational opportunities for its visitors. Children and adults alike have a unique chance to learn about the history and development of life on our planet, and to place themselves within the awesome scope of that development... a testament to what can be accomplished when public and private entities work together for the benefit of everyone.” TEXAS GOVERNOR ANN RICHARDS “This collection contains some very important finds which have added tremendously to our knowledge...from June until the Labor Day weekend. 200,000 people visited Atlanta’s Fernbank Museum of Natural History to view the highly acclaimed collection... such a remarkable presentation... so readily accessible.” GEORGIA GOVERNOR ZELL MILLER BIBLIOGRAPHY Alvarez, Walter , Frank Asaro, Helen V. Michel and Luis Alvarez, “Evidence for a Major Meteorite Impact on the Earth 34 Million Years Ago: Implications for Eocene Extinctions,” Science 216 (1982). Andrews, Roy Chapman, All About Dinosaurs (New York: Random House, 1953). Arduini, P. and G. Teruzzi, Prehistoric Atlas (London: Macdonald and Co. Publishers Ltd., 1982). Bakker, Robert T., The Dinosaur Heresies (New York: William Morrow and Company, Inc., 1986). Beerbower, James R., Search For the Past (New Jersey: PrenticeHall, Inc., 1960). Benton, Michael J., The Dinosaur Encyclopedia (New York: Simon and Schuster, 1984). On the Trail of the Dinosaur (New York: Crescent Books, 1989). Bird, Roldand T., Bones F or Barnum Brown: Adventures of a Dinosaur Hunter (Fort Worth: Texas Christian University Press, 1985). Breton, D.L., “The Arthropod Sidneyia inexpectans, Middle Cambrian, Burgess Shale, British Columbia,” Philosophical Transactions of the Royal Society of London. B295 (1981) pp. 619653. Case, Gerard R., A Pictorial Guide to Fossils (New York: Van Nostrand Reinhold, 1982). Cluver, M.A., Fossil Reptiles of the South African Karoo (Cape Town: The South African Museum, 1978). Coates, M.I. and J A. Clack, “Polydactyly in the Earliest Known Tetrapod Limbs,” Nature Vol. 347 (1990) pp. 66 -69. Colbert, Edwin H., The Great Dinosaur Hunters and Their Discoveries (New York: Dover Publications, 1968, 1984). Dinosaurs: An Illustrated History (New Jersey: Hammond Incorporated, 1983). Cook, Johnathan, “Proper Names For Early Fingers, 4 Nature Vol. 347 (1990) pp. 14 -15. Czerkas, Sylvia, ed., Dinosaurs Past and Present, J Volumes I and II, (Los Angeles County Museum Press, 1986). Czerkas, Sylvia J. and Stephen A., Dinosaurs: A Global View (New York: Mallard Press, 1991). Eldredge, Niles, Fossils: The Evolution and Extinction of Species (New York: Harry N. Abrams, Inc., 1991). Fenton, Carroll Lane and Mildred Adams Fenton, revised and expanded by Patricia Vickers Rich, Thomas Hewitt Rich, and Mildred Adams Fenton, The Fossil Book: A Record of Prehistoric Life (New York: Doubleday and Company, Inc., 1989). 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