Pacific Willow
Transcription
Pacific Willow
Pacific Willow Salix lasiandra Benth. A short lived, non colonial, fast growing, native, pioneer willow is often mistaken for peach leaf willow or resembling cottonwood suckers (5, 10, 15). Pacific willow is used for windbreaks, inexpensive erosion control and energy plantations in other countries (12, 15, 16). Many willow species, especially pacific willow, are self-infertile and freely hybridizes with other willow species. Willow bark has aspirin like properties used by Native Americans to cure many common illnesses. Two varieties occur commonly in the Pacific Northwest, lasiandra and caudata (9). Also referred to as Shinning or whiplash willow and rarely red or yellow willow. SYN: S. lucida var. lasiandra and S. lucida ssp. lasiandra. Plant Characteristics glaucous, lanceolate shaped, finely toothed, 5-10 cm (2-3.9 in) long, mature leaves bore on glandular lobed petioles (1, 9). Inflorescence: Dioecious aments are solitary, thick, hairless, light reddish, 6-8 mm (0.24-0.31 in) long. Catkin buds are slightly larger than the duck shaped leaf buds (5). Floral bracts are elliptic to oblanceolate, often shallowly toothed, pale yellow, deciduous and hairy (6, 8). Staminate catkins are 18-65 mm (0.70-2.55 in) long (1). Characteristic of pacific willow are the 4 to 5 stamens and nectaries (1). Roots: Rather aggressive moderately deep, widespread fibrous root system. Roots will bind to Iron and other toxic metals to avoid poisoning. VAM, vascular arbuscular mycorrhizae, occurs on willow roots to aid in nutrient uptake. (12) GENERAL HABITAT CONSIDERATIONS Soil Types and Conditions: Inhabits a wide variety of deep, sandy, gravelly, fine textured or acidic to neutral soils typically of alluvial deposits. Tolerates a thick mollic epipedon occurring as far as 1.6 m (5.2 ft) from active water (4). Pacific willows avoid limestone deposits (14). Moisture/temperature requirements: Grows in stream or river banks, valleys, flood plains, alluvium bottomlands, fast moving creeks, woodlands, freshwater swamps, willow-alder thickets and roadside ditches. Occupies moderate elevations from sea level to 0-2,950 m (8,500 ft) in more flood prone areas than other willow species (7, 13, 15). As the elevation increases up to 1,981 m (6,500 ft), var. caudata becomes shrubbier. Management Considerations UTILIZATION VEGETATIVE CHARACTERISTICS Form and Stature: This multi-stemmed shrub or tree normally is 5-15 m (16-49 ft), but has been observed reaching 20 m (65 ft) tall (6, 8). The dark gray to dark brown, vertically ridged, scaly trunk can be up to 60 cm (23 in) thick bearing flexible highly brittle, glossy stems. Young twigs like young leaves are covered with fine spreading hairs and with age become smooth (6, 8). Height and crown diameter are influenced by site conditions (4). Leaves: Juvenile leaves are reddish to yellowish green and coarsely toothed with long silky hairs maturing into green or Cultivars ‘Nehalem’ released 1988 Corvallis, Oregon ‘Roland’ 1987 released Alaska PMC Forage and Palatability: Preferred by cattle when other palatable forages decrease (16). Wildlife Use: Deer, elk, moose and rabbits browse willow leaves and branches (3). Wildlife browsing throughout the year drastically reduces development. Aments are eaten by small game and song birds (7). In California, the endangered Least Bell’s vireo frequently nests in the Pacific willow (15). Heavy beaver use in the winter is common. Sawflys damage willow branches and leaves by mining, boring, defoliating and producing galls (12). Pacific willows provide cover for wildlife and cattle seeking shade. PLANTING Growth Season and Seedling Establishment: Blooms mid April to mid May, flowering as soon as leaves emerge. Establishment by 17.8-25cm (7-10 in) long, one-half-inch thick cuttings by seeds is possible. Cuttings must be collected, between November and March, and planted the same day. Roots will appear within 10 days of planting, so quick placement is important. (10) Prerooted cuttings will achieve highest establishment. Willows store carbohydrates in the early stages of active growth, suggesting prerooted cuttings have more reserves than non-rooted cuttings. Seed establishment should take place in late spring. The seedbed must be moist until the seeds germinate and establish. Fertilizing with wood ash or nitrogen enhances growth. Note that urea based fertilizer inhibits production. Seeds must be collected and planted promptly because the tiny seeds have no dormancy, germination will occur within 12-24 hours. DISTURBANCE Response to Competition: Naturally, this species is replaced by cottonwoods or poplars. Overgrazing can cause willows to be replaced by grasses, especially Kentucky bluegrass. Intolerant of shading, dense grasses can inhibit seedling establishment. (15) Response to Grazing: Pacific willow will resprout from the root crown after light to moderate grazing or browsing (11, 12). Regeneration may take more than 3 years if grazing is removed or the riparian corridor is fenced off. Proper grazing can stimulate enhanced density and growth (4). Late summer grazing won’t stimulate regrowth. Response to Fire: Plants with high vigor resprout from root crowns or stem bases following a light to moderate fire. Extreme fires in riparian areas are rare but those fires will remove organic matter damaging or killing the roots. (15) Response to Drought: Moderately tolerant to periodic droughts. Once pacific willow is established, roots will be extended searching for water and eventually form support for the stream banks. Sources: 1. Argus, George W. 2003. A guide to the identification of Salix (willows) in Alberta. June-July 2003. Ontario, Canada. 78p. 2. Argus, George W. 2003. A guide to the identification of willows in Alaska, the Yukon Territory and adjacent regions. Workshop on willow identification. June-July 2003. Ontario, Canada. 104p. 3. Brunsfeld, Steven and Fredrich Johnson. 1985. Field Guide To The Willows Of East-Central Idaho. Bulletin 39. Moscow, Idaho: University of Idaho, Forest, Wildlife, and Range Sciences. 95p. 4. Clary, Warren P. et al. 1996. Response of a depleted sagebrush steppe riparian system to grazing control and woody plantings. Intermountain research station: research paper INT-RP492. 32p. 5. Collet, Dominique M. 2004. Willows of Interior Alaska. US Fish and Wildlife Service. 111p. 6. Cook, Sarah. 1997. A Field Guide to the Common Wetland Plants of Western Washington and Northwestern Oregon. Seattle: Seattle Audubon Society. 417p. 7. Darris, Dale and Scott Lambert. 1993. Native willow varieties for the Pacific Northwest. U.S. Dept. of Agriculture and Soil conservation. Corvallis, Oregon 15p. 8. Davis, Ray J. 1952. Flora of Idaho. Dubuque, IO: WM.C. Brown Company. 836p. 9. Hitchcock, C. Leo and Arthur Cronquist. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. 10. Klein, Kevin. 2003. Plant Data Sheet: Salix lucida ssp. lasiandra (pacific willow). Washington State University. 2p. 11. Kindschy, Robert R. 1898. Regrowth of Willow following simulated Beaver cutting. Wildl. Soc. Bull. 17: 290-294. 12. Kovalchik, Bernard L. 1992. Growth and Yield of willows in Central Oregon compared to reports in world literature. ProceedingsSymposium on Ecology and mamagement of riparian shrub communities. Sun Valley, ID. 83-88p. 13. Manning, M.E. and W.G Padgett. 1992. Distribution of willows on forested of Nevada and Eastern CA. ProceedingsSymposium on Ecology and mamagement of riparian shrub communities. Sun Valley, ID. 83-88p. Pacific Willow Salix lasiandra Benth. 14. Plants for a future. 1997-2000. Salix exigua. [Accessed: April 12, 2005]. 15. Uchytil, Ronald J. 1989. Salix lasiandra. In: Fire Effects Information System, [Online]. U.S.D.A, Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [2005, April 19]. 16. USDA, NRCS. 2004. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA. Prepared by OSU Rangeland Ecology and Management, April 2005.
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