Performance of Planted Native Hardwood Trees
Transcription
Performance of Planted Native Hardwood Trees
Technical Article No. 10.3 Performance of Planted Native Hardwood Trees INTRODUCTION SURVEYING PLANTED STANDS Growth and management information of selected native hardwood tree species is contained within the Tãne’s Tree Trust Indigenous Plantation Database based on stand assessments during a recent survey by Tãne’s Tree Trust (TTT) as well as during an earlier survey in the 1980s by the former Forest Research Institute (Pardy et al. 1992). Details on tree and stand assessments are given in Technical Article No. 10.1. Briefly, DBH (diameter at breast height – 1.4 m above ground level) of all planted trees within plots in native hardwood tree plantations of known age were recorded and heights measured for a minimum sample of 12 trees per plot. Site factors and stand histories were collated and growth datasets from plantation surveys analysed. 10.3 Performance of Planted Native Hardwood Trees number of native hardwood tree species including the beeches (Nothofagus spp.) have been planted over many decades throughout New Zealand. This article provides a summary of the height and diameter growth of a selection of planted native hardwood tree species including the development of preliminary growth models. Details on the survey and the Tãne’s Tree Trust database are given in Technical Article No. 10.1 in this handbook. Technical Handbook Section 10: Native Tree Plantations A GROWTH MODELS SPECIES AND NUMBERS PLANTED Stand selection and analysis Compared to the native conifers (refer to Technical Article No. 10.2 in this Handbook), far fewer hardwood trees have been planted and virtually none have been planted as a long-term timber resource. Of the over 10,000 trees in the Tãne’s Tree Trust Indigenous Plantation Database, less than 1800 are of selected native hardwood trees including the beeches have been measured to date in about 100 plots. Other than for the beeches, most hardwood trees were within small multi-species groves such as in urban parks or small plantings in established gardens. Where possible, data used for developing the growth models was restricted to stands that had been planted at regular spacing and which had minimal edge effects. Trees planted in the open or on edges of stands will have significantly faster diameter growth due to less competition compared to trees growing within plantations, so were excluded from the data used to develop growth models. Growth modelling was therefore based on a restricted number of plots and trees that represented stands. Sigmoidal growth curves of the Bertalanfy-Chapman form were fitted using nonlinear regression. Models were obtained for mean height and quadratic mean DBH. Separate slope or asymptote parameters were fitted for each species. For the height model, an intercept of 0.5 m representing height at planting was used. For the DBH model, a zero intercept at age 4 was used, this being the average age at which breast height is achieved by most native hardwoods. Height/age curves Plot mean heights along with fitted height/age regression curves for the eight hardwood tree species are given in Figure 1 for the individual species. This varied from 60-110 years of age for the different species depending on data available in the Indigenous Plantation Database. Up to twenty planted hardwood tree species have been measured but there are only eight species with sufficient numbers across a range of ages to warrant analysis of diameter and height growth. The most commonly planted beeches were red beech (Nothofagus fusca) and black beech (Nothofagus solandri) with a smaller number of silver beech (Nothofagus menziesii). Most of the beeches were established in stands in the south of the North Island and in the South Island. At 60 years after establishment where growth has been assessed across all eight species, red beech and black beech were the tallest with mean heights exceeding 20 m (Figure 1). This was followed by silver beech, rewarewa and puriri at less than 20 m mean height, taraire at 15 m and karaka and kohekohe at 10 m. For those species where plots have been established in older stands, red beech exceeds 30 m mean height 110 years after planting, followed by black beech at under 30 m and silver beech leveling off at 20 m height growth. Five other hardwood species across 3 or more sites, were karaka (Corynocarpus laevigatus), puriri (Vitex lucens), kohekohe (Dysoxylum spectabile), rewarewa (Knightia excelsa) and taraire (Beilschmedia tarairi). Most of these species were planted on North Island sites and often established as amenity groves in urban parks. There was a limited range of ages represented by the eight species with only three species – red beech, black beech and puriri – where stands over 100 years since planting were measured. All other species had limited number of stands across a range of ages. 2 Diameter/age curves Diameter/age curves exceeding 100 years are shown for the hardwood trees in Figure 2. At 60 years after planting, mean DBH ranges from 30 cm for the slowest growing species rewarewa and silver beech to 50 cm for taraire and red beech. The bulk of hardwood tree species have a mean diameter around 40 cm 60 years after planting. Figure 1: Height/age regression curves for the native hardwood tree species assessed from data in the TTT Indigenous Plantation Database that included the recent nationwide survey of native plantations. For those species with older stands, red beech is approaching 60 cm mean diameter growth in 100 years after establishment with black beech at 50 cm and puriri 45 cm for stands planted for over a century. Predicted height and diameter growth The predicted height and diameter of each of the eight hardwood tree species for selected ages up to 80 years since planting is given in Table 2. These predictions are largely restricted to within the limits of the data so predictions do not extend to 80 years for the limited stands assessed for rewarewa and taraire. Figure 4: Mean DBH/age regression curves to age 110 years for the native hardwood tree species based on growth data collated in the TTT Indigenous Plantation Database. Table 2: Predicted height and diameter for native hardwood tree species from the growth models developed for each species based on the assessment of native plantations in the nationwide surveys. Height (m) Age (years) Red beech Black beech Silver beech Puriri Karaka 10 20 40 80 5.9 10.9 18.9 28.4 5.4 9.9 17.0 25.6 4.3 7.8 13.4 20.2 5.6 9.7 15.0 19.5 Diameter (cm) 3.8 6.5 10.1 13.0 3.7 6.3 9.6 12.4 5.6 9.7 15.1 4.7 8.1 12.5 10 20 40 80 10.4 21.0 37.6 55.9 8.8 17.7 31.8 47.4 6.7 13.5 24.3 36.1 8.0 16.2 29.1 43.2 8.3 16.8 30.1 44.9 9.1 18.4 32.9 49.0 6.6 13.3 23.9 11.6 23.3 41.8 Of the selected hardwood trees assessed for height growth in planted stands, the fastest growing hardwoods are the beeches all of which are predicted to be between 20-30 m high 80 years after planting (Table 2). This is up to 35 cm annual height increment for red beech and 32 cm for black beech. Faster earlier growth rates of up to 50 cm height growth are predicted for several hardwood tree Kohekohe Rewarewa Taraire species at the 10 and 20 years periods including red beech, black beech, puriri and rewarewa. The tallest northern hardwood is puriri predicted to be nearly 20 m high 80 years after planting compared to kohekohe and karaka at only 12-13 m high over the same time period since planting. 3 The hardwood tree species with fastest predicted diameter growth include red beech (56 cm), black beech (47 cm) and kohekohe (49 cm) 80 years after planting (Table 2). Mean periodic annual diameter increment for these faster growing hardwood trees are 6-7 mm. Silver beech has the slowest diameter growth at only 36 cm 80 years after establishment. The beeches are generally the fastest growing native hardwood. Wardle (1984) indicates that red beech is the fastest growing and silver beech is slower than the other beeches. He has also found that the fastest growth period for managed beech stands tend to be at the young pole stages within the first few decades of establishment, and particularly where stands had been thinned. Rate of diameter growth is decreasing for many species from the earlier periods compared to the 80 year period. For instance, red beech is predicted to have a periodic diameter increment at 10 and 20 years of over 10mm but reducing to 9 mm at 40 years and 7 mm at 80 years after planting. Similarly, kohekohe had a periodic mean annual diameter increment of 9 mm at the 10 and 20 year period but growth rate is predicted to decline to 8 mm increment at 40 years and 6 mm at 80 years after planting. Variability between individual stands across all hardwood tree species is substantial due to the wide range of site and climatic factors that occur between stand locations. Many of the stands in the survey were not well managed after planting so that growth performance particularly within 5-10 years of planting is likely to have been seriously compromised by weed competition. As stands develop, there is inevitable increasing competition between trees and therefore a decrease in diameter growth rate as seen in the growth models developed for these selected native hardwood trees. Virtually no stands of native hardwood trees were thinned thus contributing to the decreasing diameter growth rates over time. Consequently, these average growth rates do not necessarily reflect the true potentials of each species. CONCLUSIONS Mean annual diameter growth rates of 10 mm and 50 cm for height for the faster growing native hardwood tree species within 20 years of planting is similar to those recorded previously (e.g. Bergin and Gea 2007; Pardy et al. 1992). However, growth rates have decreased over time across most species. It is inevitable that tree height growth rate will decrease with time as trees reach their natural heights, but diameter growth can continue for many decades but rate of growth will inevitably decrease with increasing within stand competition. References: Wardle, J. 1984: The New Zealand beeches: ecology, utilisation and management. New Zealand Forest Service, Caxton Press, Christchurch. 447p. Bergin, D.O.; Gea, L. 2007: Native trees – planting and early management for wood production. New Zealand Indigenous Tree Bulletin No. 3. New Zealand Forest Research Institute. Revised edition. 44p. Pardy, G. F.; Bergin, D. O.; Kimberley, M. O. 1992: Survey of native tree plantations. Forest Research Institute Bulletin No. 175. 24p. Authors: David Bergin and Mark Kimberley, Scion Contact: Tãne’s Tree Trust Website: www.tanestrees.org.nz The Ministry of Agriculture and Forestry does not necessarily endorse or support the content of the publication in any way. ISSN 2230-3014 October 2011 4 Printed by Scion Digital Print Centre, Rotorua