THE RUSSIAN PLATFORM AS A KEY REGION FOR

Rivista Italiana di Prleontoloeia e Stratigrafia
volume 1l0
no.
l
,
pr IIl;' ,t'.;r; I
.a,prirzoo+
THE RUSSIAN PLATFORM AS A KEY REGION FOR VOLGIAN/TITHONIAN
CORRELATION: A REVIE\tr OF THE MEDITERRANEAN FAUNAL ELEMENTS
AND AMMONITE BIOSTRATIGRAPHY OF THE VOLGIAN STAGE
MIKHAIL A. ROGOV'
Receit-ted September 19, 2002; accepted October 3, 2003
Key uords. Jurassic, Volgian, Tithonian, Boreal-Teth1'an correla-
Introduction
tion, amnronites, biostratigraphy, Russian Platform.
Abstract.
A review of the ammonite faunas and biostratieraphy
of the uppern.rost Kimmeridgian - lowermosr Middle Volgian of Central
Russia is presented. The major role of the Sub-n-rediterranean ammonites
in the Volgian assemblages is described, providins additional detail to
the existine scheme of Tithonian-Volgian correlations. The base of the
Tithonian and the base of the Volgian are coincident, based on the association of Neoc/retocerds steraspis-Lingulaticeras soLenoides.The Klimovi
Zone corresponds to nrost of the Hybonotum Zone. However, the top
of the Hybonotum Zone, based on the distribution of Paralingulaticeras, Íalls within the Sokolovi Zone.The allocation of r Tenuicostata
Subzone (with two horizons: neoburgense and puschi) in the Pseudoscythica Zone is proven. The neoburgense horizon may be correlated
with the cíliata md penicillatum horizons of Neuburg and, probably,
with the Semiforme Zone. The findings oÍ Lingulatìceras blaschkei, Pseudolissoceras, rnd Sutneria in the Panderi Zone of the Middle Tìthonian
suggest that the base of the Middle Volgian may not be younger rhan
the Middle Tithonian Fallauxi Zone.
Riassunto. Viene fornita una revisione delle faune ad ammoniti
e della biostratigrafia dalla sommità del Kimmeridgiano alla parte più
bassa del Volgiano medio della Russia centrale. Viene descritto il ruolo
principale degli ammoniti submediterranei nelle associazioni Volgiane.
Ciò permette di specificare I'esistente schema di correlazione Titoniano-Volgiano. Fondandosi sull'associazion e N eochetoceras sterasp is - Lin gulaticeras solenoides, la base del Tironiano e la base del Volgiano coincidono. La Zona a Klimovi corrisponde alla maggior parte della Zona
ad Hybonotum. Tuttavia la parte sommitale della Zona ad Hybonotum,
basata sulla distribuzione di Paralingulaticeras, ricad,e nella Zona a Sokolovi. Nella Zona a Pseudoscythica viene dimostrata la ripartizione di
una Sottozona a Tenuicostata (con due orizzontí: neoburgense e puscbi).
lorizzonte a neoburgense può essere correlato con gli orizzonti a cjliata e penicillatzz di Neuburg e, probabilmente, con la Zona a Semiforme. I ritrovamenti nellaZona a Panderi dei medio-tìtonìani Lingulaticeras blaschkei, Pseudolissoceras e Sutneria suggeriscono che la base
del Volgiano medio può non essere più giovane della Zona a Fallauxi
del Titoniano medio.
Geological Institute of Russian Academy
of
Over the last few decades several papers devoted to the analysis of mixed Boreal-Tethyan associations
of Mollusca, and Tithonian-Volgian correlations (Zeiss
2001Kutek & Zeiss 1997;Hoedemaeker 1991; Sachs et
a\.1979: Mesezhnikov 1989; Sey tr Kalacheva1993,1.997;
Schweigert 1.993, 1.994, 2000; Scherzinger Er Schweigert
1999) have been published. Unfortunately, material from
the Russian Platform was either not studied during those
investigations, or only ammonites never described and illustrated before were mentioned.
Material and Methods
This paper is based on the author's field investigations and collections, and on data collected by PA. Gerasimoq M.S. Mesezhnikov and
VV Mitta. The ammonites illustrated are kept in the Geological Institute
of Russian Academy of Sciences (RAS) (author's collection, no. GIN
MIV MK), Paleontological Institute of RAS (PA. Gerasimov's collection,
no. PIN 4861) and in the State Vernadsky Museum (VV Mitta's collection, authors collection, no. SGM BX 17) in Moscow. Unfortunately, the
maiority of the collection of M.S. Mesezhnikov (S. Petersburg, VNIGRI)
remained inaccessible. Also, most of the Kimmeridgian-Middle Volgian
ammonites with Tethyan affinities, quoted by M.S. Mesezhnikov (Mesezhnìkov et a|.1977; Blom et al. 1984; Mesezhnikov 1989; Yakovleva
1993) have not yet been recovered from his collection.
Hitherto, only one outcrop of the Russian Platform (Gorodishchi, Fig. 1,3) yielding an Upper Kimmeridgian-Middle Volgian ammonite
succession composed of Sub-mediterranean ammonites has been discovered. Last year all the levels containing Tethyan ammonites, identified
in the Gorodishchi section, were traced through a number of localities
in the middle Volga area (Fig. t), since it is important to ascertain that
the sequences of Sub-mediterranean ammonites in different secrions
are coincident. As a result, the wide distribution of these ammonites
throughout the middle Russirn Sea became apparent.
Sciences, Pyzhevskii Lane,7,119017, Moscon, Russia. E-mail: [email protected]
l!. A. Rogn'
322
Llr[]q
IIlz TFf ro
.lllll :ffirr
+ ffi rz
x
1
ITTTilTI
LYSKO\'O
llilllllìlltìllT
s [-,] t:
lH 6 F_E r+
iostratigraphy,:
PN - Parrderi Zone
PU - pust'hilrorizotr
N - neoburgetT.rc horizon
B
- PseLrdoscythica SLrbzone
- Sokolovi Zone
L - efinroti horizon
P
S
Strr.:r
K
\\,\S1l
N - N.srera.rprs -L.solenoides Beds
Fl, F2 - Fallax Subzone
[illlfllT]lrllll,lllll
s
z
[l]llllllllllllllrillllllillll
t
:
lllllllTllllTllTllTfl
:s
llffl]l
lfffffffffl]
lflffllfltr
ll
ffi8
t\t
15
rss
n
20
E Tflfl
ifEffiT8r
ffiilr0r
168
Tttrl
aIT----------ffi
I
ffi
-r
"
PI
Pì
N
't.
>\." ||
\l \ì\tì
krn
L
,
S
7
L
N
Iol crnrost Miclc{lc Vrlgirn
Rr'.rpino,6 Murzicr'. Z- lsrdr:
r.,i.i). lì - Istaritcs.
l3 - Littgrlaticcr-.r.r, l-{ -
ParalingtÌaticertslfotttttntcsicllr, l5 - outcrops Ioc;rlin'.
Ammonites, zonal nomenclature and correlation
cr:rnrplc scc Kutek & Zciss 1997. p1.20. tir. l). M.rcroconchs h;rrc;r lol'er
point of rib irrrc;rtion thiìt nlrkcs thcnr ve'rv sinril,rr to IlotLaiskyt klinot'i
Since there ;ire different rìpprorches in conrprring
the brse of thc Tithonian rnd Volgi'rn stirscs (Schwcisert
199-{t H:rntzpclsue ct .il. l99S). ir is nece sr.rrr to beqin
(llov.riskr' IM]. As thc evolutionrrrv tcndcncies of rlicro-,rncl nrrrcroconchs
in thc.t.rrrrutirTrltìnttes Ilot;'aisÀ1,,r ìine.rge rrre virirrble (sc'c';rlso dat;r on
thc Klinrovj Zonc). rhe gcrrus 1/oi;.ri,,,(-l'.r c:rn bc'subclivicìccl furthcr into
nricro- rncl rttlcroconchilte suLrgcner,t, rìlso rrcccptecl ior cocr',rl pcrisphincticl rr:rnrorite s ol Southcrn Ccrrrrrrrr' (Schu'cigc'rt 1996). F-rccpt tor S.lr-
this re',,ierv with r description of thc ch:rrircteristics of
the uppenlos t Ki n-rr.nerid si;rn F:rl ;rx Subzone (Auti ssioI
dorcnsis Zone).
(Pl. l.
Upper Kimnrcridgian
Arrtissiodorcnsis Zonc Zicgler, l9 62
loels ilre lin()\\'n rrlso ixnr l)ollntl (Kuteli & Zciss
1997). but the .rrtnronite rrssocirtions ol tìrc lìussirrn plrrttornr.ìrc nlorc
divcrsc. Thc ìorr cl prrt ol thc I:rrllrr SuLrzonc c()ntiì;ns rrunrcrous .S.lrlllIr
î,run,rì horizons. IJoth
spltinctcsJìtlla.t (llovriskr') l\,l.nrl,,1r1,rcosta/,lttrtusstrbtrndorircll).rrlor),--1.
jasottoitlcs (l).rvlorr).J. cr gr..llll-s-,ior1or.cl-.1-s (Cortcru), nrorc r,rrclv Ncor'lrctoterts c[. sttbnttdttrttn 1l:ont.7. LingtÌttittrts solenoitlas (Qucnstctlt) rrncl
unîreque nt,'i.'7i116131il.i sp. Thc uppcr part oi thc subzonc contit;ns nl:rinlv
S,tnnatispltittttcs sp. nor: IM, nr]. trlicrocorrchs ol this sf.ccics cli[icr ironr
(lloveiskr')
lnrl
in its
lol
riLr
coeificicnt (usurlll rbour
ii*.
l51.,4spidoceras sp. intl., Gnrle.v,r sp. ind. tn(1 AulrcostelhLúils
thcil presc'ncr'. up to the brsr. oi tlrc Lol cr
\trlgirn. is cxtrcnrelv r.rrc. It is inrport:rnt to notc th,rt Ocy'rcloccr;r-,, rs we]l
ci. jasottoidcs (l)rrvlov), thor-rgh
Fallax Sr,rbzone Iloveiskv, 1941. This subzorrc ìs n idch'clisrribrrtcd
in thc nritl.llc Volgr lrc.r. lJ,rscrl ttn thc tlistributron of,rnr:ronites, rìris sutrzonc nl;ì\' bc sLrbdividetl inro tl'o ;rrrts. s hìch c,rn be iurthcr consicle,rc.tl .rs
.S./.r1lr.r
natisphínacs sp. nor:. this Ievel inclucles corìlnron Lingulatic'cras solenoidcs
(Quc'nstcdt) (lìoeov 2002b. pl.. iir. l) nttl Naot'l:ctocarts cx !t'. subnutt"tlrnr (lrrrrt.) (lìogov2002e.pl. t,iir. 1,2):Ocbetoccr,T.ic\gr.zro (Oppel)
2.51
iol
x
Attl"ttostcplutta-t do rrot cxtcntl into thc Tìthonien ;rnri c,rrr
,rs gcncra
Lre
consiclered
charlctcristic oi the'Kinrnreridsi:rn (Schs'eìecrt, pc'r'. conrnr. (for'
Otlntotcras)\. Anroug thc oppcliids it is possibìc to notc thc plc\cncc ()f
?T,n'ancllicct',ts tii.Jt',utt'iscttunt (l"ont;rnrrcs) (Corodishchi, lsrtlv sections;
lìogor l00lr. pl. l, iir. l; Pl. l. iig. 7) tttlNeotlrtoc'?rds ci.rcboilleti"tlutnt
(Bcrckhcnrcr & Hiilder) ([sach scction, Pì. l. lig.S; r'erv closc ro N. rcbou!cri.rtttiltt irr\elru.i1.rrJJ00.1'l.l.ri.l.i'rrc.rrrlre r{,1)(,irhr'lrllrrsrrh't.onc. Ncotlx,tot'er.ts ci. rt'bouletitnntn (not illustrete cl) s,rs rrlso cluotccì
ironr the Uppe r Kinrnrerirlgi.rn oi the North Subc.rspiln rcgion (Bogcllnov
193-{) pcrrritting thc corrclation ol thc uppcnrost lìalì;rr Subzone lith
rhc raboulatinttlr horizon oi thc Southern (lcnrrrnr' (Rogov 2002r).
Upper Jurrtssic of tbe \/olgt area
323
Fig.2 - Schenre of the
ammonite distribution. Abbrevi.rtions: U.Kirnrn. - Upuer Kinrnreridgien, Autiss. -
HORIZON
ORBEDSWTIII
z
î
Au
f
F-:
=
f
Z
Tithonian-
Volgian correl;.rtion based on
Subboreal Province (qmmorite succ€ssion
ofthe Russian Plate)
Submeditemaoean Province (after Geyssmt, 1997; Schweiger! 1999; Schweigert et al., 199Q. Middle Titho
t is s
io doren
s
is.
-l
J
v l-:
>trG
z
-)
tv
T
iF
!
I
o
=
L
F
1>
=
F
,.:
J
-)
4
+3
<=
z
z =i
fz ';>Z
Y2
E
q
Trì
>z
-J
Mocnrs-
frl
hcimcnsis
l
F
-J
zt.l
[ì icclcnsc
q
v
c0
, nsrftrsrri
ll
-
U lmcnsc
O
u
o
:Y
Ruepcì lianunr
T
(pail)
a
Lower Volgian
the Neocltetotu'Lts stcl'Ltlfris Lingul"tticarus solutoides Bcris (rt the brrse
of thc zone) tncl iltc lithograplicunt eJ'intot'i horizon.
. N. .tlcril-r?l.i L. solenoides Becls Roqori 2001. N. -\Ic'ir.'?l.i
(Oppel) (Pl.l, fir. l0-11) occurrcnccs irre nunrcroLrs, u'hilst 1/or',ri.sÀ'tz
(If or,;riskr.), Lingulctticer,ts solenoitles (Qr"re nsteclt), Sunerid if.
bracheri Bc'rckhcnrer ;ìl'r' nlol'e rale (Pl. l, fig. l).These berls nr,rv correspond to tha rueppelliLlttutlt tnLl rietllittgensis horizorrs lncl ln urrnrnrecl horizon (Schs eigc'rt Ei Zeiss 1998) bets'een rhc ruappcllianutt
tnd eigeltingense horizons of Southcrn Gr'nlliìn\' (Ssrbie). A nunrbcr
of horizons frorl Soutlrcrn Gernrrrnr, :rlso cont:rin Nc,oty';cfot,rir-r ex gr.
klintoti
(Oppel) tnd Litryul,tticetLts solatoidcs (Quenstc'clt) (Schwcigert,
pers. conrnr.),
l,lrile 2l',r/ilgrrlaticerts;rre iLrsent ol'\'crv l'irre (rs
In]] ch:ìrirctcriz-cd bv,r vcry high point of rib furcatìon. The first appearmce ol Subdicboto,nocerLls (Sphinctoceras) r'Llso occurs in this horizon.
Russirrrr hoirlizguLtticeras tnd Fontitnnesielln dilfer fronr the European
gcner,r in thc;rbsc'ncc of ventrolatcnl tubercles and in the small shell
size. Ns'ertheless, t1'pe of ribbing rnd spectrum of variability within Russi,rn and tvpical Mc'dìtc'rranc'an lmnronites rre the sxne. Thus,
Russirrr specìnrens nl;.rv be considered ls endernic species of the genera
P,tralingul,rticer"ts a,nd FoiltLtnilesie ll"t.The efintoxi horìz,on corresponds
appro\inrrtelv to the cf. e)'stettense-noentsheinensis horizons of Southe rn Gerrr.rrrr.rl, (Irir.2). The level rvith nunlerous Paralingulrtticeras is abscnt in Srvirbir (Schrvcigcrt, pers. cournr.) end Polrrnd. Probably these
rnrnrorrites penetrilted into the nriddle Volg;r rrer froÌl the Northern
CrLrcrsus, n,lrere dimorphs o'i Paralingulaticer"ts - Fontannesiella proI
ithogrdp hi
,rre representccl bv L
nt (Fonurn nes) occur.
Sokolovi Zone Ilovaisky, 1941
vct in thc Russirn Plrtiornr,
but rn rnr:ronite verv closc'to N. notlu/osunr Berclihenrer & H<tlder
(nricroconclri;rtc'counterp;ìl't of N. pruecursor) r';rs cliscovcrec] in tlre
Mr-rrzicv section (nor in situ) (Pl.l, fiu.9).
. eJinroz,i Horiz-on Rogorl 2002. Thc r';rri;rblc spccic's P,rr.rIiuguIctticents cy'rrot'i (Iìogor) (Pl.I, fir. Il-1-j) is conrnron, inclurling
nrorphs resenrbling 2 ex gr. ltrcfurleirti (Oppel) (Rogov 2002b, pl., fig.
4) and P perceruli (Fout;rnncs) (Rogov 2002b, pl., fi:l.Z) in thc r"rppcrnrost pirrt. pr'rssin{ into the Sokolovi Zone. NtocJtttorclr-. ci. rrcrìr.rP/i
(Oppel), Fottt,tttnasiclla sp. (Pl.l, [ì9.l,1), Haplocair., sp. (Pl.l, fiiì.6),
a,nd Gr,t"-esia (inclucling
cu
;n
rhc riedlingensi,s horiz-on). The sepiìrrìtc lo'c'l with N. pracurrsor Zciss
(eigeltingense horizorr) h,rs not Lrccn found
q
l
Klimovi Zone Michailov, 1964
In thc nriddle Vrlga art':r thc Klinrovi zonc'is subdivicled into
steraspis
J
-
al
)<
F
F
fO
o
J
G. cl. gr,tt'csiatta (Orb.)) ;Ìrc r;ìrc. Ilot;visktt
hlintot'i (llovriskv) flvf, nr] lncl 1/otlri-sÈ.lu sp. nor:
it will be possible to subdivide this zone rnto two
t sokoloti horiz-on below lncl t ptti:ida horizon rbove,
Evcntu;rlhi
frrnn,rl Ircrrizons,
;rs
it
is done
for tlre
z-one
in Pol;.rnd (Kutek 6i Zeiss 1997), but at present
insufficicnt. Accorcling to Mesezhnikov (Mesezhnikov et :rl.
lgZZ; Blorr c't al. l9tÌ-l), I. sokolot'i (lloveisky) occLrrs ne;ìr the base of
thc zonc, tosctlrcr with htralingulaticeras b"tebcrleini (Oppel), P perce'',r/r (Font;rnnes). In the ìipper p;rrt oI the Sokolovi Zone, Ilouaishya
p.rr.'rrlr (llovaiskv) lM, nrl is present together l'ith rare Borell amnronites (Subdiclntonocer,ts (Sphinctoceras)). The base of the Sokolovi
Zone (brsed on the presence ol htralingulaticerrs) lies rvithin the Hybonoturl Zone (Rouor'2002b), end the top is located rvithin the PalrrtinLrs Zone (Kutek El Zeiss 1997).
clrrtrr rrre
M. A. Rogoo
324
; i"{
a
z
aì
<N
ZP
RA\(,t
F
={:f:
5
EI I
'i r= E==ll
;-tE ll
It EE !i;itl
sttl
=
Zl
{i^
?3
I
I
>t>o
^c
=-H
I
hr
ú,.
luEllri'l
;;
_,e
iÈ;t
I
ìÉ.ii ,ii*
l :E
z
o
J
o
J
=iiìi
{ ;tr;Èi ll
a
Li
a
(llARI\
o
T
<V
ú,
7r.jú;J-= .=
I2
a
a''Èì=
À.: \'E È
(, Al\1r\r()\lll
cJ
?
I lr:
-=:Éiì^:5i-
.,
o
,
l**È:sììllPiF'
Z
S
O
q
I'lillì1il
:l
A
I
I
2
5
I
rizon; An - Anaspidocerds; F
- Fontannesielld; F2 - Fallax
I Ilotaisk)'a:L- Linguhticer,tslT lt et cerzr ; N.s.- L. s. N oc b et o cerrts
:N - N
Subzone, uppcr
cera
s
prrt;
e o c
o
e
steraspis- Lingulaticeras so/enoides Beds; P
P.e.
- efimovi
Litholosy:
clay, 6
a
1
-
Parz/ inguLtticeras;
horiz-on; P.p. - puschi horizon; Ps.
gatites; S - Stntt,ttisphinctcs: St.
silt:
neoburgense
Horizon Rogor', 2002. Anrtspidoceras neoburgense
"rsena
pl., fiia.10) iìre rArer.
Vith the exception of the anrrronites, other r.nol-
lusc,r frr:rnr the neoburgense horizon h;rve a Sub-rlediterrlne.rn origìn.
Belenrnites ,rre represented only bv Hil:olithes, lnd :r ver), divcrse bivelve
asscnrbl;rsc irrcludes con.ìmon M),ophorelLt, and l:rrge Liosn"ea. For rhe
îirst tirre since the Oxfordian Procerìtltium is found at this level. This
Neoburgense event is not defined in Poland and, probably, is connected
to rn increasing ìnfluence of the North-Caucrsian Basin.The neoburgelse horizon is s,ìdely distributcd jn Centr,rl Russìa. In edclition to the
Vof gr r.Lre;r, A. neoburgense (Oppcl) is knorvn rrlso ironr the Ural River
(llovrìskv & Florensky 1941, pI.23, fis.42,42a) and frorr the Moscow
;rrea (S:rsonov 1953; not illustrated). The prcsence of the,4. neoburgense
(Oppcl) lnd
the
S. dsenLl (Oppel) neoburgense horizon is correlated with
penicillatun md ciliata horizons of Neuburg (Fig.2). This horizon,
probably corresponds to the Semiforme Zone of the other Tethyen areas,
in r.hich the species,4. neoburgense (Oppel) a,nd S. .tserna (Oppel) also
xre very nun'rerous (Kutek & \Wierzbowski 1986; Fòzy et al. 1994).
. puschi Horiz-on Kutek and Zerss, 1974. The index species of
the Puschi Zonc oî Polend (Kutck & Zeiss 197.1) was later replaced by
the Tenuicost:rtunr (Zeiss 1977),6ut the first nanre is more preferable
in Rnssir, x P tenuicostatun (Michailov) trlso occurs ìnrhe neoburgense
horizon. It is probable that Mesezhnikov's (1982) opinton rh:;. P. tenu
icosttttun (Michailov) occurs throughout the Pseudoscythica Zone was
rvrong. In the Gorodishchi section P tenuicostatunt (Michailov) and P
puschi Kutek & Zeiss (Rouoi' 2002, pl., fir.l3) occur approxirnately in
ìdentic;rf anrounts, but in the Polevye-Bikshiki secrion P. tenuicost"ttunt
(Michailov) stronuly prevrils. Danubisphinctes sp. jur: (Pl.l, fig.1Z) also
occurs sporldic,rlly in the Polewe-Bikshikì sections and rare Pseudor:ìt'gatites a[f. seorsus (Oppel) are knorvn fronr the Gorodishchi section (Irie.1,
3). Based on the presence o'i Danubispbinctes spurius (Schneid) in the
coeval deposits of Poland, the upper part of the Tenuicostata Subzone
can be compared rvith the uppernrost Prlmatus Subzone of Neuburg.
Some researchers (e.g. Callomon
the uppermost Kimmeridgi;rn ro lorverr.r.rost Middle Voleian).
Abbre vi;rtions: A. - Aulacostephatrus; A.n. - neoburgense ho-
e IIi
.
(Oppel) fM, ml is very conrnron (P1.1, fig.l6). Ilo-oaisk1,a cf. pseudosc;,r/ricz (llovaìsky), Pseudot,irgttítes tenuicostdtrz (Michailov) (Pl. l,
fig.l8; this species is considered s Pseutloi:irgatites because of the los,
positìon of the point of rib furcation in the inner rvhorls), Sutneria
(Oppel) (Pl.l, fir.2,3) and ?Pseudolissocerds sp. (Rogov 2002b,
Panderi Zone Rosanov, 1906
- Arnrlonite rlnse charts in the Gorodishchi section (from
Titrrtrn
Zetss 1997) probablv occur in the sanre subzone.
Tenuicostxtum Subzone Zeiss, 1977
Middle Volgian
6
Fig.
Pseudoscythica Subzone Ilovaisky, 1941. In the nrajority of
sections, <>nly I. pseudoscl,t/:ica (llovaisky) [M, m I is known from this
level. I. schascbkozae (llor,aisk1,) ancl L ianscbini (Ilovaisky) (Kutek &
- drrk-grri' cley;
-
d:rrk-r.:r.ty;'{
phosphorite nodules.
er;r1,, b
-
Pseudovir-
- Sutilerid.
I - qr,rl crlc.rreous clay; 3 - nr.rrl; 5 - bituminous shal1,
Pseudoscythica Zone Ilovaisky, 1941
The Pseudoscythica Zone is subdivided ìnto trvo subzones
Pseudoscvthicrr and Tenuicosrrrunr.
tr Birkelund
1982;
Hantzpereue et al. 1998; Scherzinger & Schweigert 1999) suggest the
presence of a gap between the Lorver and Middle Volgian. However,
due to the lÌradual chanses in the Pseudovigratitinae-Virgatitinae linerge, thìs is considered ìmprobable. In the lorvest 0.5 nr of the Panderi
Zone in the Gorodishchi section, Dorsopllnitinae occur together with
amnronites, which are closer to Pseutlooirgatites, than to Zar"tiskites.
Therefore, we cannot be sure th:rt the base of the Panderi Zone tn
Russia ,rnd the base of the Scythicus Zone of Poland are coincident.
The allocirtion of the detached level rvith Zaraiskites contadictionis
(llovaisky) andZ. tliprosopa (Ilovaisky), as proposed by Zeiss (2OOl),
ìs questionable. These anrmonites rre found together wirh Zaraisbites
s cyt b i c u s (Vischniakoff ), D ors o p I an it e s p an deri (d' O rbigny), Padovia
paz:loti (Michrlski) (Mikhailov 1964) and correspond closely to the
r';rriabilitv of Z. scytbicus (Vischniakoff). Only the trmnronites named
by Ilovaisky (19a1, p.128, pl. 26, fig. 50) as Z. contradictionis var. B
differ appreciably front Z. settbicus (Vischniakoff). These .rnrntonites,
which resenrble Ilouaiskya, are found in the phosphate nodules horizon in VetÌianka. It is possible that they rvere redeposited from the
Lower Voleian.
The Pancleri Zone oÍ the Central Russia still cennot be subdivided into f;runal horizons. Hoq,ever, the presence of Zaraishites regu-
Upper J urassic of the Volga area
laris Kutek in the bituminous shales of the Gorodishchi section and Z.
quenstedti (Rouillier), Z. scythicus (Vischniakoff) in the lower part of
Panderi Zone suggests that in the future it will be possible to trace also
in Central Russia the horizons identified in Poland by Kutek (1994). Z.
regularìs Kutek occurs also in the northern areas of the Russian platfornr (Sachs 1976, pl. 22, fig. 2).
325
Dorsoplanites panderi
deriZone cannot be younger than the Middle Tithonian Fallauxi Zone
(Rogov 2002b). On the other hand, findings oí Zaraiskites regularis
Kutek in the Upper Tithonian of the Carpatho-Balkan zone allows
us to compxre the uppermost part of the Panderi Zone with the part
of Upper Tithonian Calpionella.Zone A (Kutek 1994; Kurek & Zeiss
1997). Ar present x more precise correlation of the Panderi-Virgatus
boundary with the Tethyan zonal succession is not possible,
is (Vischniakoff ), Pa.o loo ia p av looi (Michalskl),Acuticostites acuticostatus (Michalski),2{. bitrifurcatus Mitta (Mitta
I 993) are found in the Panderi Zone. Anrmonites with a Tethyan affinity are very rare. According ro Mesezhnikov (Mesezhnikov et al. 1977;
Blom et al. I 984; Yakovleva I 993), in the Gorodishchi section the Panderi Zone contains unfrequent Pseudolissoceras, Sutneria, Glochiceras.
Unfortunately, these ammonites have yet to be found in his collection stored in VNIGNI (S. Petersburg). R:r'e Lingulaticeras blaschkei
Cecca Er Enay (Pl.l, fig. l3,la) xre known from the Panderi Zone of
Chuvashiya and the Tver area (Rogov & Egorov 2002, fig. 1.2 a, b), as
well as in the Fallauxi Zone (Richteri Subzone) of Ardèche (Cecca Ec
Enay 1991). The presence of this species and anrmonites of the genera
Pseudolissoceras r'nd Sutneria, indicates that the lower oart of the Pan-
Achnowledgemenrs. I wish to thank Dr. G. Schweigert (Stuttgaft) for providing so much information concerning the ammonite succession of South Germany, including unpublished data. Dr. V V Mitta and Dr. O. A. Erlanger from Moscow kindly loaned specimens of
Lingulaticeras blaschkei from the Panderi Zone. Biostratigraphic studies of the Kimmeridgian-Volgian deposits and the presentation of the
results was nrade possible with the help of Dr. S. A. Kurenkov (GIN
RAS, Moscorv). Valuable suggestions were made by Dipl.-Ing. A. Scherzinger (Hattingen) and by reviewers, Drs. G. Schweigert (Stuttgart) and
J.C.V Cope (Cardiff). This publication has been supported by RFBR
grant no.03-05-64297.
In addition to Zaraishites, tlso
(d'Orbigny), D . tlors
op la n
PLATE I
All illustrated in natural size and collected by the author, except for marked specinrens.
l. Sutneria {f . bracheri Berckh. Klinrovi Zone, N. steraspis - L. solenoides Beds. Gorodishchi. (x2) GIN MIV 667/l;2. Sutneria cf . asema (Opp.).
Pseudoscythica Zone, Tenuicostatum Subzone,neoburgense horizon. Gorodishchi (x2). GIN MK 576; 3. Sutneria asema (Opp.). Pseudoscythica
Zone, Tenuicostatum Subzone, neoburgense horizon. Gorodishchi (xa). GIN MK 609; 4, 5. Lingulaticeras blaschhei (Cecca, & Enay). Panderi Zone.
BX 17/1. Collected by VV Mitta; 5 - Kimry (Tver area). PIN 4861/25. Collected by PA. Gerasim ov;6. Haploceras sp. ,uv.
Klimovi Zone, eJìmooi horizon. Gorodishchi. SGM 572-7 (x2); 7. ?Tarantelliceras franciscanum (Font.). Autissiodorensis Zone, Fallax Subzone.
Isedy. GIN MIV 285; L Neochetoceras cf. rebouletianun (Berckh. t{. Hólder). AutissiodorensisZone, Fallax Subzone. Isady. GIN MIV 284; 9. Neochetoceras nodulosum Berckh. Er Hólder. Klimovi Zone, not in situ (?N. ster.tspis-L.solenoid.es Beds). Murzicy. GIN MIV 630; 10. Neochetoceras
cf. steraspis (Opp.). Klimovi Zone, N. stertlspis - L .solenoides Beds. Polevye-Bikshiki. GIN MIV 655; 17. Neochetoceras cf. steraspis (Opp.). Klimovi Zone, N. steraspis - L. solenoides Beds. Gorodishchi. SGM 572-22: 12, 13. Paralingulaticer"ts efimooi Rogov Klimovi Zone, efimovi horizon.
Gorodishchi. l2 - SGM 572-21; 13 - Holotype SGM 572-l l: 14. Fontannesiella sp., Klimovi Zone, efimoai horizon. Gorodishchi. SGM 522-6;
15. Ocbetocer,ts tf zio (Opp.). Autissiodorensis Zone, Fallax Subzone. Gorodishchi. GIN MIV 692/l;16. Anaspidoceras neobu,Eense (Opp.) [m],
Pseudoscythica Zone, Tenuicostatum Subzone,neoburgense horizon. Polevye-Bikshiki. GIN MK 607;17. Danubìspbinctes sp. juv. Pseudoscythica
Zone, Tenuicostatum Subzone, puscbi horizon Polevve-Bikshiki. GIN MK 589; 18. Pseudooirgatites tenuicostittzrr (Michailov), Pseudoscythica
4 - Per-vonrayskoe. SGM
Zone, Tenuicostrtum Subzone, puschi horizon Polerrre-Bikshiki.
GIN MK
582.
326
M.
A.
Rogot,
',t"{
,l
|II
7?:
;€l
*$
I
l5
I
18
PI,ATI.-
I
Uppar J rtr,t'sit' oJ'the
\ltlg
arcd
327
REFEIIENCES
Blonr G.1., Kuznetz.rx';r K.l. & Mcscz.hnikov M.S. (1934) Jur:issic-Cret:rccous boundrrrl bccls in the Miclcllc Vilg.ì
;rncl Rjrs;rn ;rrciìs. txcLrrsion 0(r0. 27 Intern;rtionrrl Gcol.
Congr., Moscoui
198-1. Guicle booli of thc excursiorrs 059.
060, 0(r6: 33--19, Moscori'.
Boqci:rnov A.A. (1934) - Hvdrochloric clo:rcs oI the l-ou'cr Zavolz.hie . Ilrtll. Soc'. Natur. Alosc'o u. Scct. Góo1., l2: 315367, Moscos'. Iin Iìussiirn]
Crllorlon J.H. & Birkclund T. (1982) - Thc rrr.r.rnronirc z.rr.rcs
of tlre Borcrrl Vrl!Ìi,ìn (Uppe r JLrrassic) in E:rst Grce nland. In lrrrbrv A.li. E: Balkri,ill H.R. (Eds.): Arctic Gcologv ,rrrcl Gcopìrvsics. A'lcnt. Cdrtad. Soc. Pctntl. Gaol.,
8: 3-19-369, C.rl garr,'.
Cccc;r F. c\ Enav R. (199 l) - Les;tr.t.tr.nonitcs clcs z-r'rr.tc i Scnrifornrc ct ì Frrllauxi clu Tithonicltrc clc I'Arcle'chc (Sucl[:st dc la lìr:rncc): strrtiqr:rphic, peleror-rtolouic, prlcrobiogéogrephie . Palaunttogr,rpltittt. A., 2 I 9. I -3: I -tì 7,
Stuttg:rrt.
Ijijzv I. (1988) -
Upprcr Juressic f;rcics :rncl :Ìululonrtc sLrcccssion of thc Transclanubi:rr-r Ce rrtral lì.rrnr:c (Hunr:;rrr'). In
Roch;r R.B., So;rrcs A.P (tds.): 2ncl Int. Sr-nrpos. Juress.
Str:rtisr., 2: 5tì l-5tì3, Lisboa.
Gcvss:trtt I. (1997) - Titl-ronicn. In C:rriou L,.
"\ H:rntzpe rsuc P
(Eds.) - Biostr.rtier:rphie du Ju r:rssiquc Oues t-lìu nrpóclt
ct Móditcrr;rrrócn. Bull. Cctttrc
I
|J.ccb.
Elf Expktr. Pnil.,
Mcsczhnikor' lvt.S. (1939) - Tithonirrn, Volsirrrr rìnd Portlxndi;rn
st;lses (geologicrl trrrcl biologicrl cvcnts, corrclltion). In
Soliolov B.S. (Ed.): Scclinrentarv cover of thc L,arth ir-r
spi.rcc i.rnd timc. Strrtisrrrphl' rncl p;1le ontolos\': 100- 107.
Moscos'. Iin Russien]
Mcscz-hniliov N,l.S., D.rin l-.G., Kuznctzor';r K.I.
"\ Y;rkovlcv:-r
S.lì (1917) - Jur:rssic/Crctiìceolrs bouncl,rrv bcds in thc
Middlc Volg;r Arc;r (A Prospc'ctus to Gcological Excursion): -1-l pp., [-cningrrd.
Mikh.rilov N.P (196+) - Borc;rl l-:rtc Jur:rssic (l-orvcr Volgi:rn)
iìrìrnronites (Virg;rtosphinctin:rc). T1'dttsttct. Geologic,rI
Irtst., 107: Z-33. Moscou,. Iin Russien]
Mitt.r V.V. (1993) - Anrr.r.ror.ritcs ;rncl zon;rl str':rtiqrrphy of thc
Middlc Volgien clcposits oi tl.re Ccntrrrl lìussir. 133 pp.
Kiev. Iin Russien]
Roeov M.A. (2002r) - Ar-rtissioclorensis Zonc (Upper Kimnrcriclgirn) of thc Volgtr ;ì.rcr'r: iullnl()r1;tc rrssembhges,
biostr:rti gr:ìph)i corrcl;rtion. In. : Boqd:rnc>r' N.A., Vierz--
rl. (L,cls.): Modern qucstions oi seoloqv: 320-325. Moscot'. Iirr Russi;rn]
Rosov M.A. (2002b) - Str;rtiurrphv of tl.rc Lou,cr Volgi:rn deposits irr thc Russi;rn Pl,rtc errcl correl:rtion bctu.ecn Volqian
,rncl Titlror-rirrn St;recs. Srrarigrapbl, Geol. Correlatiott, 4:
bitsl<r' VL,., Vishncvsl<ava VS. ct
3
t lt
Hrrrtz-pcrsuc P, B:ruclin Ir., Mitt;r V., Olfcricv A. & Z:rkh:rrol
V. (1998) - The Upper Jur;rssic of thc VolrÌa Lr:rsirr: rrnrr.r.ronitc biostr,rtigrtrphv iÌnd occurre ncc of orqillic-c.ì|bon rich facics. Correl:rtions Lrctl,ccrr IJore:rl-Subborcal
;rnd Submcclitcrr;ìncrrr.ì Provitrccs. In Cr:-Lsquin-Solc:ru
S., Barrier E. (Eds.): Pcri-Tcthls Mcnroir -l: cpicr.rtonic
brsins oi Pcri-Tethr':rn pl;rtforurs. tllén. .1[us. nath. Hist.
79: 9-.1.1,
Sachs
l:
Mcsczhniliov M.S. & Shulginr N.l. (1979) - Stretisraphv of the Jur;rssic-Crct:rccous boundlrv beds in thc
Borc:rl Bclt. In Sechs VN. (Ed.): UppcrJurrssic and their
boundrrv n.ith Ce rt:rccous S1'sterl: 93- 102, Nor,osibirsk.
P,1f i5.
Jr"rrr-Lssic-Crct;rccoLrs bou ncl;rrr'. Ncii',s/.
Srllrigr;,
35.
37-6A, Bcrlirt & Stuttq:rrt.
Ilov;riskv D.I.
&
Irlorcnskv K.P (19+l)
-
Lcs ;rrt'rn.ror.ritcs
clu
Juressiquc sr,rpóricur clcs brrssins clcs riviòrcs Ourll e t Ilck.
Contribtftiorr d li cotnttlissttncc dc la géologic dc I'USSR..
Nour'. Ser., l(5):7-lL)5, Moscou'. Iin Russirnl
Kutck J. (1994) - Thc So'thicus Zonc (Midclle Volgien) in Pollnd: its ;ullnr()nitcs ;rncl biostrltigr;rphic snbclivisions.
Ac'ta gcol. Poltntiu., ++/l-2. l-13. \Versz:ur';r.
Kutck J. & Vicrzbos ski A. (1986) - A ne l' ;ìccount on thc Uppcr Jurassic stretigrirph\' ,Lncl ,rnrrlorritcs of the Ctl>rsztvn succe ssion, Picne rrv Klippcn Be lt, Polrnd.,4 cta Caol.
Polottic',t., 36/1: 29 | 3 15, W:rrszrtri';r.
KutcliJ. & Ze iss A. (197+) - Tithoni:rn-Vrlsi:tn anrnronitcs frorl
Brzost<is k:r nc:Ìr Torìr:ìsz(i\\' M;rzos'iccki. Ccntral I)ol.rncl.
Ac' tt C c o L P t t I n t i c',t., 2 1/ 3 : 5 O 5- 5 12, \\'rrrs zrrs'rr.
W;lrszil\\'i.t.
Mcse
zhnikoi' M.S. (l9tì2) - Titl.ronian (Vrluian) Stese . Zorre s of
the Jur:rssic Svstcrn in thc USSIì. Tr',ttts,tct. Itttcnlcpart.
Sn,ttígr. Cont., lA:' 120-l+6. [-cningrecl. Iir-r Russien]
Russì;rn]
Iin
S;ìson()\'N.T. (1953) - Stretirlrephv of theJurrssic ilnd Cretaceous
cìcposits of the Russi;rn pl:rtfomr. Bull. Soc. Ndtur. trloscou.
Scct. Géol., 28. 5: Zl-100, Moscos: [in Russi;u]
So'I.l. & K;rhcho'a E.D. (1993) - Biostr;rtigr;rphicrl criteria of
thc Jurrrssic-Crct:rce olrs bor-rndrrv for thc Russi:r. 60 pp.
S;rint-Petc'rsbure. Iin Russi:-rn]
Scv Ll. 3. K;rl;rcho'i-t E.D. (1999) - Lol'cr Bcrriasian of Southe rr-r
I)rinron'e (Frrr Lrrst Rtrssia) ,rncl tìrc problcrl of Borc;rlTet
I
h
r';rn co rrcLrt io
r.r. Pa l a a o gc
ogr., Pt
l tt a o c' l i n
r.,
Pa l tt e o
e
ui.,
50: 19-63, An'rstcrrl:rrl.
Schsciecrt G. (1999) - Die Anrnronrtcntc clcr Ncr-rburg-Ii)nll;ìtiol'ì (Oberjurri,
Sticllichc lir;rnkcnalb) :rncl ihrc Bcziclrungcn z-unr Volgirrrrr. J1rtt. llat,ar. Staatsslg. P,tl,iottt. bist. Gcol., 39: 3-
Scìrcrzinqcr A.
t
Kuteli J. & Zeiss A. (1992) - Thc hiehcst Kinrnrericlgi.rn encl
l-orvcr Volgien in Ccntr:rl Pol:rncl; thcir ;rnunonites ,rncl
bicrstrrrtiurlphv. Act,t Geol. Priottitit.. 17/3-1: l0l-198.
i c'cras :
Sachs V.N.,
Hocdcnr:rcker Ph.J. (199 1) - Tethvan-bore;rl correl:rtions;tncl
thc
& Egorov E.Yu. (2002) - Micldlc Tithonian G/o-
its distribution, r.ni gr;rtior-rs ;rnd s i gnificance for
thc Borcrrl-Tcthvrn corrclltion. In: Bogchnov N.A., Vicrzbitskv VL,., Vishnevsk.tv:r VS. et al. (L,ds.): Modcrn
(ìucstions of geologr': 325-329, Moscorv. [in Russian].
VN. (Ed.) (I976) -Str:rtigr:rph1' of thcJur,rssic Systcnr of
thc north of thc USSR. 136 pp. Moscorv. [irr Russian]
7: 97- 102, Prtris.
1t,tt., |
+8--16-+, Birnrir-rql-r;tnt.
Rosov M.A.
&
horiz-on
Iìr.rr
rr
ll,
Miirrcl'rcn.
en
igcrt G. (I99-1) - Subbore ,rlc Frlllncnclcnrcr.rtc' (Anrrnonoiclc.r) irl obcrcrr WeilÌjur:r (Oberliinrrlcrideiunr) dcr
Schu;ibiscl.rcn Alb. Prrl'i1,5: l -l l- 155, Stutts:u't.
Sclrs'eiscrt G. (199+) - Ubcr ciniec bcnrcrlicns\\/crtc Ar.t.r.nonitcn inr Obcrliinrrleridgiur'ìl clcr Schl':ibischen Alb
(Siìclrvestcletr tsch l.rnd ). S t t ttgtrtt'r Il e i t r. N n trktle., IJ.
l0.l: l- 15, Stuttgelt.
Schu
e
t
t
M.
328
A.
Schweigert G. (1996) - Die Hangende Bankkalk-Formation iirn
schwàbischen Oberjura.Jber Mitt. obenbein. geol. Ver.,
N.F., 78: 281-308, Stuttgan.
Schweigert G. (1998) - Die Ammonitenfauna des Nusplinger
Plattenkalks (Ober-Kimmeridgium, Beckeri-Zone, Ulmense-Subzone, Baden-\liirttemberg). Stuxgarter
B
ein
Naturkde., B. 267: 1-61, Stuttgart.
Schweigert G. (2000) - New biostratigraphic data from the
Kimmeridgian/Tithonian Boundary Beds of Srùl Germany In Hall R.L. 6c PL. Smith (Eds.): Advances in
Jurassic Research 2000. GeoResearch Forurn,, 6: 195-
202,Zuerich-Ueticon.
Schweigert G., Krishna J., Pandey B. 8c Pathak D.B. (199ó) -
Rogoo
A new approach to the correlation of the Upper Kimmeridgian Beckeri Zone across the Tethyan Sea. N. tb.
G e o l. Pakont., Abb., 202/ 3 t 3 4 5 -37 3, Stutt gart.
Yakovleva S.P (Ed.) (1993) - Unified stratigraphical scheme of
the Jurassic deposits of the Russian Platform (1993).28
sheets, 71 pp., Saint-Petersburg.
Zeiss A. (1977) - Some ammonites of the Kletnice Beds (Upper Tithonian) and remarks on correlation of the Uppermost Jurassic. Aca Geol. Polonica.,27/3: 369-386,
'Warszawa.
Zeiss
A.
(2001) - Die Ammonitenfauna der Tithonklippen von
Ernstbrunn, Niederósterreich. Denhschr. Naturhist. Mus.
Wien., 6: I 15 pp., rùlien.